Podogennonta

The argestid Anoplosomella, from here.


Belongs within: Oligoarthra.
Contains: Leptastacus, Paraleptastacus, Psammastacus, Cylindropsyllinae, Psammopsyllinae, Leptocaris, Arenopontia, Parastenocaris, Forficatocaris, Remaneicaris, Phyllopodopsyllus, Cletodes, Diosaccinae, Stenhelia, Amphiascopsis, Amphiascus, Mesamphiascus, Amphiascoides, Robertsonia, Schizopera, Bryocamptus, Attheyella, Elaphoidella, Moraria, Nitocrella, Parapseudoleptomesochra, Nitocra, Stygonitocrella, Laophonte.

The Podogennonta is a major clade of harpacticoid copepods. Many taxa within the clade are interstitial in habitat.

Synapomorphies (from Seifried 2003): Females with endopod of mandible with 3 proximal lateral setae, 3 more distal lateral setae and 3 + 2 + 2 apical setae; endopod of maxillule with 4 setae; syncoxa of maxilla with (3 + 3), 3, 3 setae, the two proximal endites fused, distal praecoxal endite with 2 flagellate setae; both coxal endites with setae of characteristic shape: anteriorly 1 claw-like seta, posteriorly 2 strong, mostly unilaterally spinulose setae with subterminal flagellum; acessory armature of fused endopod segment: 1 seta on posterior surface, seta 9 displaced to proximal segment of endopod, seta 10 lacking; endopod 3-segmented with armature formula: 2, 1, 3; setae 4 and six lost. Maxilliped 4-segmented with praecoxa and coxa separated, basis, and 1-segmented endopod; endopodal setae 1 to 4 only represented by a group of 4 small setae on anterior surface of endopod, no setae geniculated. Leg 1 of characteristic shape: endopod-1 elongate, endopod-2 and endopod-3 short; endopod-3 with 1 innter seta, terminally with anterior claw, 1 middle geniculate seta and 1 posterior miniaturised seta; 1 inner seta lacking; exopod-3 terminally with 2 geniculated setae, 2 inner setae lacking. Formula of armature of leg 1: coxa 0-0, basis I-I, exopod I-0, I-1, III-2-0, endopod 0-1, 0-1, I-2-1. Leg 5 with endopodal lobe with six setae: 3 inner setae, 2 terminal setae, 1 outer seta; exopod foliated with 8 setae: 3 inner, 2 terminal, 3 outer setae.

Podogennonta
    |  i. s.: Antiboreodiosaccus Lang 1944 S03
    |         Dactylopina Brady 1910 S03
    |         Flavia Brady 1899 S03
    |         Mawsonella Brady 1918 S03
    |         Pyrocletodes Coull 1973 S03
    |         Tisemus Monard 1928 S03
    |         Metahuntemannia Smirnov 1946 S03
    |           |--M. spinipes Dahms & Pottek 1992 S03
    |           `--M. triarticulata Schriever 1984 S03
    |         Schizacron vervoorti (Fiers 1987) S03
    |--Argestidae S03
    |--Adenopleurellidae S03
    |--Cancrincolidae S03
    |--Cletopsyllus Willey 1935 N41b [Cletopsyllidae S03, Cletopsyllinae]
    |--Cristacoxidae S03
    |--Normanella Brady 1880 N41b [Normanellidae S03, Normanellinae]
    |--‘Stenhelia’ glacialis Brady 1918 N41b [Pseudotachidiidae S03, Pseudotachidiinae]
    |--Ismardis Leigh-Sharpe 1936 [Ismardiidae] S03
    |--Thompsonula N41b [Thompsonulidae S03]
    |    `--T. hyaenae (Thompson 1889) [incl. Robertsonia aculeifera Klie 1913] N41b
    |--Balaenophilus W81 [Balaenophilidae S03]
    |    `--B. unisetus W81
    |--Lourinia Wilson 1924 N41a (see below for synonymy)
    |    `--L. armata (Claus 1866) W67 (see below for synonymy)
    |--Laophontopsis Sars 1908 N41b [=Cleta Claus 1863 (preoc.) N41b; Laophontopsidae S03]
    |    |--*L. lamellifera (Claus 1863) [=*Cleta lamellifera] N41b
    |    `--L. secunda (Sewell 1924) N41b
    |--Parastenhelia Thompson & Scott 1903 N41b [incl. Microthalestris N41b; Parastenheliidae S03]
    |    |--‘Microthalestris’ forficula B26
    |    |--P. hornelli Thompson & Scott 1903 W67
    |    `--P. spinosa (Fischer 1860) W67
    |--Rhynchothalestris [Rhynchothalestridae] S03
    |    |--R. helgolandica (Claus 1863) S03
    |    |--R. rufocincta N44
    |    `--R. vanhoeffeni Brady 1910 N44
    |--Laophontodes Scott 1894 N41b [Ancorabolidae S03]
    |    |--L. armatus (Lang 1936) W67 [incl. Laophonte echinata Willey 1930 N41b]
    |    |--L. bicornis Scott 1896 N44
    |    |--L. brevis Nicholls 1944 N44
    |    `--L. typicus N44
    |--Huntemanniidae S03
    |    |--Huntemannia Poppe 1884 S03
    |    `--Nannopus S03
    |         |--N. abyssi Sars 1921 S03
    |         `--N. palustris Brady 1880 S03
    |--Dactylopusia N44 [Dactylopusiidae S03]
    |    |--D. micronyx N41b
    |    |--D. oculata Gurney 1927 N41b
    |    |--D. platysoma Thompson & Scott 1903 N41a
    |    |--D. tisboides N41b
    |    `--D. vulgaris N41b
    |--Delamarella Chappuis 1954 W86 [Latiremidae S03]
    |    |--D. arenicola Chappuis 1954 W86
    |    |--D. eximia (Božić 1969) W86
    |    |--D. galateae Cottarelli 1971 W86
    |    |--D. karamani Petkovski 1957 W86
    |    `--D. phyllosetosa Kunz 1984 W86
    |--Phyllognathopodidae S03
    |    |--Allophyllognathopus Kiefer 1967 R86
    |    |    `--A. brasiliensis Kiefer 1967 R86
    |    `--Phyllognathopus Mrazek 1893 R86
    |         |--P. bassoti Rouch 1972 R86
    |         `--P. viguieri (Maupas 1892) B01
    |--Leptastacidae [Leptastacinae] S03
    |    |--Leptastacus W86
    |    |--Paraleptastacus W86
    |    |--Psammastacus W86
    |    `--Arenocaris Nicholls 1935 W86
    |         `--A. bifida Nicholls 1935 W86
    |--Cylindropsyllidae S03
    |    |  i. s.: Ichnusella Cottarelli 1971 R86
    |    |           |--I. eione Cottarelli 1971 R86
    |    |           `--I. pasquinii (Cottarelli 1969) R86
    |    |--Cylindropsyllinae W86
    |    `--Psammopsyllinae W86
    |--Orthopsyllus Brady & Robertson 1873 N41a [Orthopsyllidae S03]
    |    |--O. linearis (Claus 1866) W67
    |    |--O. littoralis Nicholls 1942 N42b
    |    |--O. major Klie 1939 N41a
    |    |--O. propinquus Monard 1926 N42b
    |    |--O. rugosus Nicholls 1941 N42b
    |    |--O. similis Nicholls 1942 N42b
    |    `--O. wallini Lang 1934 N42b
    |--Metidae S03
    |    |--Laubieria W67
    |    |    |--*L. corallicola Soyer 1966 W67
    |    |    `--L. secunda Wells 1967 W67
    |    `--Metis Philippi 1843 N41a
    |         |--M. holothuriae (Edwards 1891) W67 [=Abacola holothuriae B26]
    |         |--M. ignea (see below for synonymy) B26
    |         `--M. jousseaumei (Richard 1892) [=Ilyopsyllus jousseaumei] N42b
    |--Darcythompsoniidae S03
    |    |--Kristensenia Por 1983 S03
    |    |--Leptocaris S03
    |    |--Darcythompsonia W86
    |    `--Horsiella N44
    |         |--H. brevicornis N44
    |         |--H. gurneyi Nicholls 1944 N44
    |         `--H. trisetosa Kunz 1935 N44
    |--Leptopontiidae [Leptopontiinae] S03
    |    |--Arenopontia W86
    |    |--Leptopontia Scott 1902 S03
    |    |    |--L. curvicauda Scott 1902 W86
    |    |    `--L. punctata Huys & Conroy-Dalton 1996 S03
    |    `--Notopontia Bodiou 1977 W86
    |         |--N. galapagoensis Mielke 1982 W86
    |         `--N. stephanieae Bodiou 1977 W86
    |--Parastenocarididae S03
    |    |--Parastenocaris CMA05
    |    |--Forficatocaris CMA05
    |    |--Remaneicaris CMA05
    |    |--Simplicaris Galassi & de Laurentiis 2004 CMA05
    |    |--Paraforficatocaris Jakobi 1972 R86
    |    |    `--P. paranaensis Jakobi 1972 R86
    |    |--Potamocaris Dussart 1970 CMA05
    |    |    `--P. bifida Dussart 1979 R86
    |    `--Murunducaris Reid 1994 CMA05
    |         `--M. juneae Reid 1994 CMA05
    |--Rhizothrix Brady & Robertson 1875 W86 [Rhizothricidae S03]
    |    |  i. s.: R. curvata Brady & Robertson 1880 W86
    |    |         R. gracilis (Scott 1903) W86
    |    |         R. minuta (Scott 1903) W86
    |    |         R. reducta Noodt 1952 W86
    |    |           |--R. r. reducta W86
    |    |           `--R. r. noodti Galhano 1970 W86
    |    |         R. scotti Lang 1936 W86
    |    |         R. spinosa Coull 1971 W86
    |    |         R. tenella (Wilson 1932) W86
    |    |         R. wilsoni Bodin 1977 W86
    |    `--R. (Rhizothrix) W67
    |         |--R. (R.) pubescens Por 1959 W67
    |         `--R. (R.) quadriseta Wells 1967 W67
    |--Tetragonicipitidae [Pteropsyllidae, Tetragonicepsidae] S03
    |    |--Phyllopodopsyllus W67
    |    |--Diagoniceps N44
    |    |--Laophontella Thompson & Scott 1903 W67, N41b
    |    |    `--L. armata (Willey 1935) [=Phyllopodopsyllus armatus] N44
    |    |--Tetragoniceps W67
    |    |    |--T. bergensis W67
    |    |    |--T. brownei Wells 1967 W67
    |    |    `--T. scotti W67
    |    `--Pteropsyllus W67
    |         |--P. consimilis (Scott 1894) W67
    |         `--P. plebius Monard 1935 W67
    |              |--P. p. plebius W67
    |              `--P. p. furcatus Kunz 1938 W67
    |--Harpacticidae S03
    |    |--Harpacticella SS03
    |    |--Perissocope litoralis Lang 1934 S03
    |    |--Zausopsis mirabilis Lang 1934 S03
    |    |--Tigriopus SS03
    |    |    |--T. brevicornis CCG01
    |    |    `--T. fulvus W67
    |    |--Zausodes S03
    |    |    |--Z. septimus Lang 1965 S03
    |    |    `--Z. sextus Lang 1965 S03
    |    |--Harpacticus S03
    |    |    |--H. chelifer (Müller 1776) W67
    |    |    |--H. compsonyx Monard 1926 W67
    |    |    |--H. gracilis Claus 1863 W67
    |    |    |--H. littoralis B26
    |    |    |--H. trisetosus Lang 1948 S03
    |    |    `--H. uniremis B26
    |    `--Zaus Goodsir 1845 S03, N42a
    |         |--Z. abbreviatus Sars 1911 N42a
    |         |--Z. aurelii Poppe 1884 [incl. Z. caeruleus Campbell 1929, Z. intermedius Nicholls 1939] N42a
    |         |--Z. contractus Thomson 1883 N42a
    |         |--Z. goodsiri Brady 1880 N42a
    |         |--Z. sarsi Nicholls 1942 N42a
    |         `--Z. spinatus Goodsir 1845 N42a
    |--Cletodidae S03
    |    |--Cletodes N41b
    |    |--Cletocamptus Schmankevitch 1875 R86
    |    |    |--C. deitersi (Richard 1897) S-MR03
    |    |    `--C. xenuus Por 1968 R86
    |    |--Parepactophanes Kunz 1935 W86
    |    |    |--P. dubia Noodt 1958 W86
    |    |    `--P. minuta Kunz 1935 W86
    |    |--Tryphoema Monard 1926 W86
    |    |    |--T. bocqueti (Božić 1954) W86
    |    |    |--T. lusitanica (Wells & Clark 1965) W86
    |    |    |--T. porca Monard 1926 W86
    |    |    |--T. ramabula (Pennak 1942) W86
    |    |    |--T. riedli Coull 1971 W86
    |    |    `--T. scilloniensis (Wells 1968) W86
    |    `--Enhydrosoma W86
    |         |--E. barnishi Wells 1967 W67
    |         |--E. buchholtzi (Boeck 1872) W67
    |         |--E. curticaudatum N41b
    |         |--E. curvirostre W67
    |         |--E. latipes (Scott 1909) W67
    |         |--E. littorale Wells 1967 W67
    |         |--E. longifurcatum Sars 1909 W67
    |         `--E. parapropinquum Gomez 2003 S03
    |--Thalestridae S03
    |    |  i. s.: Amenophia Boeck 1865 S03
    |    |         Paradactylopodia brevicornis (Claus 1866) W67
    |    |         Diarthrodes W67
    |    |           |--D. dissimilis W67
    |    |           |--D. glaber Wells 1967 W67
    |    |           |--D. minutus (Claus 1863) W67
    |    |           |--D. nanus (Scott 1914) W67
    |    |           `--D. sarsi (Scott 1909) W67
    |    |         Idomene [incl. Xouthous] W67
    |    |           |--I. aberrans Por 1964 W67
    |    |           |--I. aemula Thompson & Scott 1903 W67
    |    |           |--I. laticaudata (Thompson & Scott 1903) W67
    |    |           |--‘Xouthous’ maldivae Sewell 1940 W67
    |    |           `--I. simulans (Brady 1910) W67
    |    |         Dactylopodella flava N41b
    |    |--Dactylopodiinae N41a
    |    |    |--Dactylopodia tisboides (Claus 1863) W67
    |    |    `--Eudactylopus Scott 1909 N41a
    |    |         |--E. australis Nicholls 1941 N42b
    |    |         |--E. latipes (Scott 1894) N41a
    |    |         |--E. robustus (Claus 1863) W67
    |    |         `--E. spectabilis (Brian 1923) N41a
    |    `--Thalestrinae N41a
    |         |--Thalestris longimana BK77
    |         `--Phyllothalestris Sars 1905 N41a
    |              |--P. lata Nicholls 1942 N42b
    |              |--P. mysis (Claus 1863) N41a
    |              |    |--P. m. mysis N42b
    |              |    `--P. m. harringtoni Willey 1935 N42b
    |              |--P. paramysis Monard 1928 N42b
    |              |--P. royi (Monard 1928) N42b
    |              `--P. sarsi Sewell 1940 W67
    |--Diosaccidae [Miraciidae] S03
    |    |  i. s.: Pseudamphiascopsis ismaelensis (Monard 1936) W67
    |    |         Bulbamphiascus imus (Brady 1872) W67
    |    |         Haloschizopera junodi (Monard 1935) W67
    |    |         Paramphiascella robinsoni (Scott 1902) W67
    |    |         Melima W67
    |    |         Eoschizopera W86
    |    |         Helmutkunzia W86
    |    |         Typhlamphiascus Lang 1948 R86
    |    |           `--T. latifurcata Por 1968 R86
    |    |         Pseudostenhelia Wells 1967 W67
    |    |           `--*P. prima Wells 1967 W67
    |    |         Paramphiascoides Wells 1967 W67
    |    |           `--*P. mixtus Wells 1967 W67
    |    |         Actopsyllus Wells 1967 W67
    |    |           `--*A. longipes Wells 1967 W67
    |    |         Schizoperoides Por 1968 R86
    |    |           `--S. expeditionis Por 1968 R86
    |    |         Goffinella Wilson 1932 W86
    |    |           `--G. stylifer Wilson 1932 W86
    |    |         Balucopsylla Rao 1972 W86
    |    |           `--B. similis Rao 1972 W86
    |    |         Protopsammotopa Geddes 1968 W86
    |    |           |--P. norvegica Geddes 1968 W86
    |    |           `--P. wilsoni Wells 1977 W86
    |    |         Robertgurneya W67
    |    |           |--R. rostrata (Gurney 1927) W67
    |    |           |--R. similis (Scott 1896) W67
    |    |           `--R. spinulosa (Sars 1911) W67
    |    |         Psammotopa Pennak 1942 W86
    |    |           |--P. chappuisi Noodt 1955 W86
    |    |           |--P. phyllosetosa (Noodt 1952) W86
    |    |           |--P. polyphylla Noodt 1955 W86
    |    |           `--P. vulgaris Pennak 1942 W86
    |    |--Diosaccinae N41b
    |    |--Stenheliinae N41b
    |    |    |--Stenhelia N41b
    |    |    `--Pseudomesochra Scott 1902 [incl. Stenheliopsis Sars 1906] N41b
    |    `--Amphiascinae N41b
    |         |--Amphiascopsis N41b
    |         |--Amphiascus N41b
    |         |--Mesamphiascus N41b
    |         |--Amphiascoides N41b
    |         |--Robertsonia N41b
    |         `--Schizopera N41b
    |--Canthocamptidae S03
    |    |--Nannomesochra arupinensis (Brian 1925) W67
    |    |--Bryocamptus S-MR03
    |    |--Attheyella S-MR03
    |    |--Elaphoidella R86
    |    |--Moraria S-MR03
    |    |--Echinocamptus Chappuis 1928 R86
    |    |    `--E. georgevitchi (Chappuis 1924) B01
    |    |--Gulcamptus Miura 1969 R86
    |    |    `--G. uenoi Miura 1969 R86
    |    |--Antarctobiotus Chappuis 1930 R86
    |    |    `--A. bahamondei Ebert & Noodt 1975 R86
    |    |--Epactophanoides Borutzky 1966 R86
    |    |    `--E. udegeicus Borutzky 1966 R86
    |    |--Stygepactophanes Moeschler & Rouch 1984 R86
    |    |    `--*S. jurassicus Moeschler & Rouch 1984 R86
    |    |--Paracamptus Chappuis 1929 R86
    |    |    |--P. gasparoi Stoch 1998 B01
    |    |    `--P. nakamurai Chappuis 1958 R86
    |    |--Morariopsis Borutzky 1931 R86
    |    |    |--M. kieferi Petkovski 1959 R86
    |    |    `--M. scotenophila (Kiefer 1930) B01
    |    |--Canthocamptus Westwood 1836 R86
    |    |    |--C. insoletus Chappuis 1928 R86
    |    |    `--C. morimotoi Miura 1969 R86
    |    |--Spelaeocamptus Chappuis 1933 R86
    |    |    |--S. incertus Petkovski 1956 R86
    |    |    |--S. neotropicus Noodt 1963 R86
    |    |    `--S. spelaeus (Chappuis 1925) R86
    |    |--Mesochra Boeck 1865 R86, S03
    |    |    |--M. hiunumaensis Kikuchi 1972 R86
    |    |    |--M. lilljeborgi [incl. Laophonte hecate Brehm 1910] N41b
    |    |    |--M. pygmaea (Claus 1863) W67
    |    |    `--M. rostrata (Gurney 1927) W67
    |    |--Antrocamptus Chappuis 1956 R86
    |    |    |--A. catherinae Chappuis & Rouch 1960 R86
    |    |    |--A. chappuisi Rouch 1970 R86
    |    |    |--A. coiffaiti Chappius 1956 R86
    |    |    |--A. longifurcatus Rouch 1970 R86
    |    |    `--A. stygius Rouch 1970 R86
    |    |--Maraenobiotus Mrazek 1893 R86
    |    |    |--M. brucei R86
    |    |    |    |--M. b. brucei R86
    |    |    |    `--M. b. carpathicus Chappuis 1928 R86
    |    |    `--M. insignipes R86
    |    |         |--M. i. insignipes R86
    |    |         `--M. i. kyzylkumicus Borutzky 1972 R86
    |    `--Ceuthonectes Chappuis 1924 R86
    |         |--C. chappuisi Rouch 1980 R86
    |         |--C. gallicus Chappuis 1928 R86
    |         |--C. hungaricus Ponyi 1958 R86
    |         |--C. mirabilis Miura 1964 R86
    |         |--C. rouchi Petkovski 1984 R86
    |         |--C. serbicus Chappuis 1924 [incl. Morariodes colchidana Borutzky 1930, C. colchidanus] R86
    |         `--C. vievilleae Rouch 1980 R86
    |--Ameiridae S03
    |    |  i. s.: ‘Amphiascus’ mucronatus Brady 1910 N41b
    |    |         ‘Amphiascus’ proximus Scott 1914 N41b
    |    |--Psammoleptomesochra australis (Mielke 1994) [=Paramesochra australis] S03
    |    |--Pseudameira W67
    |    |--Proameira W67
    |    |--Nitocrella R86
    |    |--Leptomesochra W67
    |    |--Parapseudoleptomesochra S-MR03
    |    |--Nitocra S-MR03
    |    |--Stygonitocrella R86
    |    |--Sicamaeira W86
    |    |--Inermipes EHH05
    |    |--Pseudoleptomesochra Lang 1965 W86
    |    |    `--P. typica Lang 1965 W86
    |    |--Psyllocamptus Scott 1899 R86
    |    |    |--P. minutus Sars 1911 W67
    |    |    `--P. monachus Chappuis 1938 R86
    |    |--Paraleptomesochra Wells 1967 W86
    |    |    |--*P. minima Wells 1967 W67
    |    |    `--P. wellsi Rao 1972 W86
    |    |--Philoleptomesochra Wells 1967 W67
    |    |    |--*P. reductiforma Wells 1967 W67
    |    |    `--P. secunda Wells 1967 W67
    |    |--Karllangia Noodt 1964 W86
    |    |    |--*K. arenicola Noodt 1964 W67, W86
    |    |    |--K. psammophila Wells 1967 W86
    |    |    `--K. tertia Kunz 1975 W86
    |    |--Sarsameira Wilson 1924 [incl. Pseudosarsameira] W67
    |    |    |--S. exilis (Scott & Scott 1894) W67
    |    |    |--S. minor Wells 1967 W67
    |    |    `--S. pendula (Shen & Bai 1956) W67
    |    |--Psammameira Noodt 1952 W67
    |    |    |--*P. hyalina Noodt 1952 W67
    |    |    |--P. grandis (Nicholls 1939) [=Ameira grandis] W67
    |    |    `--P. reducta Wells 1967 W67
    |    |--Nitocrellopsis Petkovski 1976 R86
    |    |    |--N. elegans (Chappuis & Rouch 1959) R86
    |    |    |--N. intermedia (Chappuis 1937) R86
    |    |    `--N. ioneli (Dumont & Decraemer 1974) R86
    |    |--Praeleptomesochra Lang 1965 W86
    |    |    |--P. africana (Kunz 1951) W86
    |    |    |--P. phreatica Pesce 1981 R86
    |    |    |--P. pygmaea (Vervoort 1964) W86
    |    |    `--P. similis Lang 1965 W86
    |    |--Ameira W86
    |    |    |--A. exigua W67
    |    |    |--A. leptoderma (Klie 1950) W67
    |    |    |--A. longipes Boeck 1864 W67
    |    |    |--A. parvula (Claus 1866) [=Amphiascus parvulus] N41b
    |    |    `--A. simulans W67
    |    |--Pseudoleptomesochrella Lang 1965 W86
    |    |    |--P. bisetosa Lindgren 1975 W86
    |    |    |--P. halophila (Noodt 1952) W86
    |    |    |--P. incerta (Chappuis & Delamare Deboutteville 1956) W86
    |    |    |--P. marina (Chappuis & Rouch 1960) W86
    |    |    `--P. pontica Apostolov 1969 W86
    |    |--Parevansula Guille & Soyer 1966 W86
    |    |    |--P. elegans (Marinov 1974) W86
    |    |    |--P. mediterranea Guille & Soyer 1966 W86
    |    |    |--P. reductiforma (Wells 1967) W86
    |    |    |--P. secunda (Wells 1967) W86
    |    |    |--P. vermiformis Moore 1976 W86
    |    |    `--P. wellsi (Marinov 1973) W86
    |    `--Interleptomesochra Lang 1965 W86
    |         |--I. attenuata (Scott 1896) W86
    |         |--I. boguensis Lindgren 1975 W86
    |         |--I. elongata (Božić 1955) W86
    |         |--I. eulitoralis (Noodt 1952) W86
    |         |--I. noodti Galhano 1968 W86
    |         |--I. reducta Lang 1965 W86
    |         `--I. tenuicornis (Sars 1911) W86
    `--Laophontidae S03 [Namakosiramiidae MD01]
         |--Heterolaophonte quinquespinosa (Sewell 1924) W67
         |--Onychocamptus mohammed (Blanchard & Richard 1891) S-MR03 (see below for synonymy)
         |--Klieonychocamptus W67
         |--Pseudocletopsyllus Vervoort 1964 W67
         |--Mictyricola Nicholls 1957 W67
         |--Pholenota Vervoort 1964 W67
         |--Novolaophonte viatorum Cottarelli, Saporito & Puccetti 1983 W86
         |--Laophonte N41b
         |--Platychelipus Brady 1880 N41b
         |--Sarsocletodes Wilson 1924 [=Pseudocletodes Sars 1921 (preoc.), Pseudoplatychelipus Lang 1936] N41b
         |--Donsiella Stephenson 1935 N41b
         |--Psammolaophonte Wells 1967 W86
         |    `--*P. spinicauda Wells 1967 W67
         |--Apolethon Wells 1967 W67
         |    `--*A. fumator Wells 1967 W67
         |--Esola Edwards 1891 N41b
         |    `--E. spelea (Chappuis 1938) R86
         |--Cubanocleta Petkovski 1977 R86
         |    `--C. noodti Petkovski 1977 R86
         |--Mexicolaophonte Cottarelli 1977 W86
         |    `--M. arganoi Cottarelli 1977 W86
         |--Arenolaophonte Lang 1965 W86
         |    `--A. stygia Lang 1965 W86
         |--Stygolaophonte Lang 1965 W86
         |    `--S. arenophila Lang 1965 W86
         |--Pseudolaophonte Scott 1896 N41b
         |    `--P. spinosa (Thompson 1893) [incl. P. aculeata Scott 1896] N41b
         |--Harrietella Scott 1906 N41b
         |    `--H. simulans (Scott 1894) N41b
         |--Hemilaophonte Jakubisiak 1932 N41b
         |    `--H. janinae Jakubisiak 1932 N41b
         |--Lobitella Monard 1934 N41b
         |    `--L. apoda Monard 1934 N41b
         |--Tapholeon Wells 1967 W67
         |    |--*T. ornatus Wells 1967 W67
         |    `--T. uniarticulatus Wells 1967 W67
         |--Paralaophonte W67
         |    |--P. brevirostris (Claus 1863) W67
         |    |--P. congenera (Sars 1908) W67
         |    |--P. quinquespinosa P71
         |    `--P. tenera (Sars 1920) W67
         |--Asellopsis Brady & Robertson 1873 N41b
         |    |--A. duboscqui Monard 1926 N41b
         |    |--A. hispida Brady & Robertson 1873 N41b
         |    |--A. intermedia (Scott 1895) N41b
         |    `--A. littoralis Nicholls 1939 N41b
         |--Klieonychocamptoides Noodt 1958 W86
         |    |--K. arenicola (Chappuis & Delamare Deboutteville 1956) W86
         |    |--K. arganoi Cottarelli & Mura 1980 W86
         |    |--K. itoi Mielke 1981 W86
         |    `--K. remanei Noodt 1958 W86
         |--Echinolaophonte Nicholls 1941 N41b
         |    |--*E. horrida (Norman 1876) [=Laophonte horrida] N41b
         |    |--E. armiger (Gurney 1927) (see below for synonymy) N41b
         |    |--E. brevispinosa (Sars 1908) [=Laophonte brevispinosa] N41b
         |    `--E. mirabilis (Gurney 1927) [=Laophonte mirabilis] N41b
         |--Laophontina Norman & Scott 1905 N41b
         |    |--L. acantha Noodt 1955 W86
         |    |--L. distincta Wells 1967 W86
         |    |--L. dubia Norman & Scott 1905 N41b
         |    |--L. noodti Kunz 1983 W86
         |    |--L. reducta Coull & Zo 1980 W86
         |    |--L. triarticulata Coull & Zo 1980 W86
         |    `--L. variabilis Coull & Zo 1980 W86
         `--Aphrolaophonte Chappuis 1960 W86
              |--A. aequatorialis Cottarelli & Mura 1982 W86
              |--A. brevipes (Chappuis 1954) W86
              |--A. leonis Cottarelli & Mura 1982 W86
              |--A. michaelae Cottarelli & Mura 1982 W86
              |--A. monodi Chappuis 1960 W86
              |--A. pori Masry 1970 W86
              |--A. renaudi (Chappuis & Delamare Deboutteville 1956) W86
              `--A. schmidti Mielke 1981 W86

Echinolaophonte armiger (Gurney 1927) [=Laophonte armiger; incl. L. hystrix Brian 1928, L. steueri van Douwe 1929] N41b

Lourinia Wilson 1924 N41a [=Jurinia Claus 1866 (preoc.) N41a; incl. Ceylonia Thompson & Scott 1903 N41a, Ceyloniella Wilson 1924 N41a; Ceyloniellidae, Louriniidae S03]

Lourinia armata (Claus 1866) W67 [=Jurina armata N41a, Ceylonia armata N41a, Ceyloniella armata N41a; incl. Ceylonia aculeata Thompson & Scott 1903 N41a, Ceyloniella aculeata N41a, Ceylonia aculeata var. adriatica Brian 1923 N41a, Ceyloniella aculeata var. adriatica N41a]

Metis ignea [incl. Ilyopsyllus coriaceus Brady & Robertston 1873, I. natans Williams 1907, I. sarsi Sharpe 1911] B26

Onychocamptus mohammed (Blanchard & Richard 1891) S-MR03 [=Laophonte mohammed S-MR03; incl. L. humilis Brian 1929 N41b]

*Type species of generic name indicated

REFERENCES

[BK77] Barel, C. D. N., & P. G. N. Kramers. 1977. A survey of the echinoderm associates of the north-east Atlantic area. Zoologische Verhandelingen 156: 1-159.

[B26] Bigelow, H. B. 1926. Plankton of the offshore waters of the Gulf of Maine. Bulletin of the Bureau of Fisheries 40 (2): 1-509.

[B01] Brancelj, A. 2001. Male of Moraria radovnae Brancelj, 1988 (Copepoda: Crustacea), and notes on endemic and rare copepod species from Slovenia and neighbouring countries. Hydrobiologia 453/454: 513-524.

[CCG01] Canivet, V., P. Chambon & J. Gibert. 2001. Toxicity and bioaccumulation of arsenic and chromium in epigean and hypogean freshwater macroinvertebrates. Archives of Environmental Contamination and Toxicology 40: 345-354.

[CMA05] Corgosinho, P. H. C., & P. Martínez Arbizu. 2005. Two new interstitial species of Remaneicaris Jakobi 1972 from the Ribeirão do Ouro River, Minas Gerais, Brazil, with a redefinition of the genus (Crustacea, Copepoda, Harpacticoida, Parastenocarididae). Senckenbergiana Biologica 85 (2): 147-162.

[EHH05] Eberhard, S. M., S. A. Halse & W. F. Humphreys. 2005. Stygofauna in the Pilbara region, north-west Western Australia: a review. Journal of the Royal Society of Western Australia 88 (4): 167-176.

[MD01] Martin, J. W., & G. E. Davis. 2001. An updated classification of the Recent Crustacea. Natural History Museum Los Angeles County, Science Series 39: 1-124.

[N41a] Nicholls, A. G. 1941a. Littoral Copepoda from South Australia. (I) Harpacticoida. Records of the South Australian Museum 6 (4): 381-427.

[N41b] Nicholls, A. G. 1941b. A revision of the families Diosaccidae Sars, 1906 and Laophontidae T. Scott, 1905 (Copepoda, Harpacticoida). Records of the South Australian Museum 7 (1): 65-110.

[N42a] Nicholls, A. G. 1942a. A review of the genus Zaus Goodsir, and a description of two species of Laophonte Philippi (Copepoda, Harpacticoida). Annals and Magazine of Natural History, series 11, 9: 119-127.

[N42b] Nicholls, A. G. 1942b. Marine Copepoda from Western Australia. I. Littoral harpacticoids from Rottnest Island. Journal of the Royal Society of Western Australia 27: 135-141.

[N44] Nicholls, A. G. 1944. Littoral Copepoda from the Red Sea. Annals and Magazine of Natural History, series 11, 11: 487-503.

[P71] Por, F. D. 1971. One hundred years of Suez Canal—a century of Lessepsian migration: retrospect and viewpoints. Systematic Zoology 20 (2): 138-159.

[R86] Rouch, R. 1986. Copepoda: Les harpacticoïdes souterrains des eaux douces continentales. In Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) (L. Botosaneanu, ed.) pp. 321-355. E. J. Brill / Dr. W. Backhuys: Leiden.

[S03] Seifried, S. 2003. Phylogeny of Harpacticoida (Copepoda): Revision of “Maxillipedasphalea” and Exanechentera. Cuvillier Verlag: Göttingen.

[SS03] Seifried, S., & H. K. Schminke. 2003. Phylogenetic relationships at the base of Oligoarthra (Copepoda, Harpacticoida) with a new species as the cornerstone. Organisms Diversity & Evolution 3: 13-37.

[S-MR03] Suárez-Morales, E., & J. W. Reid. 2003. An updated checklist of the continental copepod fauna of the Yucatan Peninsula, Mexico, with notes on its regional associations. Crustaceana 76 (8): 977-991.

[W81] Watson, L. 1981. Sea Guide to Whales of the World. Hutchinson: London.

[W67] Wells, J. B. J. 1967. The littoral Copepoda (Crustacea) of Inhaca Island, Mozambique. Transactions of the Royal Society of Edinburgh 67: 189-358.

[W86] Wells, J. B. J. 1986. Copepoda: Marine-interstitial Harpacticoida. In Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) (L. Botosaneanu, ed.) pp. 356-381. E. J. Brill / Dr. W. Backhuys: Leiden.

2 comments:

  1. Dear Chris, that is an Anoplosomella. It is not considered Ameiridae anymore, being transfered to Argestidae.

    ReplyDelete
    Replies
    1. Thank you, Paulo, I've edited the picture caption accordingly.

      Delete

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