Anthrax maculatus, photographed by Michael Jefferies.

Belongs within: Anthracinae.

The Anthracini are a cosmopolitan group of beeflies, characterised by the presence of an apical circlet of hairs around the compacted, often onion-shaped antenna. They include the cosmopolitan genus Anthrax, many species of which have the wings infuscated to varying degrees.

Characters (from Lambkin et al. 2003): Antennal basal stylomere bearing apical stylomere and circlet of hairs; R2+3 arising from R with distinct basal 90° bend at i-r crossvein, squama fringed with scales; pulvilli normal, large flattened membranous structures, almost as long as tarsal claws; laterotergite and mediotergites bare.

    |--Brachyanax ater LYG03
    |--Thraxan LYG03
    |--Xenox GEEL09
    |    |--X. habrosus GEEL09
    |    |--X. nigritus GEEL09
    |    `--X. tigrinus KME08
    `--Anthrax LYG03
         |--A. albofasciatus KME08
         |--A. analis A71
         |--A. argentiforns Macquart 1849 E12
         |--A. argyropygus KME08
         |--A. blandus K01
         |--A. cathetodiathmos GEEL09
         |--A. cintalapa GEEL09
         |--A. fasciventris Macquart 1849 E12
         |--A. flavus K01
         |--A. georgicus KME08
         |--A. gideon GEEL09
         |--A. irroratus KME08
         |--A. ixion K01
         |--A. leucopyga BM76
         |--A. maculatus LYG03
         |--A. maurus K01
         |--A. midas GEEL09
         |--A. minuta Macquart 1849 E12
         |--A. morio L02
         |--A. oedipus GEEL09
         |--A. pauper KME08
         |--A. picea KME08
         |--A. pluto KME08
         |--A. quinquefasciatus K01
         |--A. sinuata R35
         |--A. stellans KME08
         `--A. varipennis Macquart 1849 E12

*Type species of generic name indicated


[A71] Askew, R. R. 1971. Parasitic Insects. Heinemann Educational Books: London.

[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[GEEL09] Greathead, D. J., N. L. Evenhuis & C. J. Einicker Lamas. 2009. Bombyliidae (bee flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 565–576. Ottawa, NRC Research Press.

[K01] Kertész, K. 1901. Legyek [Dipteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 179–201. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[KME08] Kits, J. H., S. A. Marshall & N. L. Evenhuis. 2008. The bee flies (Diptera: Bombyliidae) of Ontario, with a key to the species of eastern Canada. Canadian Journal of Arthropod Identification 6: 1–52.

[LYG03] Lambkin, C. L., D. K. Yeates & D. J. Greathead. 2003. An evolutionary radiation of beeflies in semi-arid Australia: systematics of the Exoprosopini (Diptera: Bombyliidae). Invertebrate Systematics 17: 735–891.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[R35] Rayment, T. 1935. A Cluster of Bees: Sixty essays on the life-histories of Australian bees, with specific descriptions of over 100 new species. Endeavour Press: Sydney.

Last updated: 11 June 2017.

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