Wings and genitalia of Sabulopsocus tractuosus, from Schmidt & New (2004).

Belongs within: Psocomorpha.
Contains: Elipsocidae, Lachesilloidea, Pseudocaecilioidea, Peripsocoidea.

The Homilopsocidea are a clade of bark-lice supported by two synapomorphies: the presence of an egg-guide exteneding from the dorsodistal margin of the female subgenital plate, and dorsal swelling of the dorsal valve of the gonapophyses (Yoshizawa 2002).

Characters (from Yoshizawa 2002): Head usually with rounded vertex except sharply pointed in Calopsocidae; postclypeus usually well bulged; epistomal suture with or without well-developed internal ridge; anteclypeus variable; male eyes usually much larger than female’s; ocelli complete, usually clustered on weakly developed tubercle; mandible short, with rounded outer margin except Calopsocidae with somewhat angled outer margin; lacinial tip usually bicuspid; galea usually flattened; labial palpus rounded. Prothorax less bulged dorsally; pterothorax strongly bulged dorsally; mesothorax with broad precoxal bridge and narrow trochantin; metaepisternum with broad membranous region. Forewing with various coloration and markings; posteroproximal corner smoothly rounded; venation generally of Caecilius-type. Hindwing with rounded posteroproximal corner; Sc faint. Male genitalia variable; phallosome complete. Female genitalia: subgenital plate usually with distal projection extended from dorsal margin. Gonapophyses usually complete; dorsal valve swollen dorsally, usually with subapical ventral process.

<==Homilopsocidea [Calopsocoidea, Elipsocoidea]
    |--+--Elipsocidae Y02
    |  `--Drymopsocus Smithers 1963 SN04
    |       `--*D. brunneus Smithers 1963 SN04
    `--+--Lachesilloidea Y02
       `--+--Pseudocaecilioidea Y02
          |--Peripsocoidea Y02
          `--Sabulopsocidae SN04
               |--Sabulopsocus Smithers 1969 SN04
               |    `--*S. tractuosus Smithers 1969 SN04
               `--Moapsocus Schmidt & New 2004 SN04
                    `--*M. angelicus Schmidt & New 2004 SN04

Homilopsocidea incertae sedis:
  Lesneiidae [Lesneiinae] SN04
    |--Hemicaecilius Enderlein 1903 SN04
    `--Lesneia Badonnel 1931 SN04
         |--*L. capensis Badonnel 1931 SN04
         |--L. niger Broadhead & Richards 1982 SN04
         |--L. pulchra Broadhead & Richards 1982 SN04
         `--L. stuvkenbergi Badonnel 1963 SN04
  Polypsocus Hagen 1866 M93 [incl. Ptilopsocus Enderlein 1900 M93; Polypsocidae L02]
    |--*P. corruptus (Hagen 1861) (see below for synonymy) M93
    `--P. fasciatus Banks 1908 M74

*Polypsocus corruptus (Hagen 1861) [=Psocus corruptus; incl. Psocus abruptus Hagen 1861, Polypsocus abruptus, Ptilopsocus annulicornis Banks 1903, Po. corruptus var. omissus Banks 1938, Po. corruptus var. pictilis Banks 1938] M93

*Type species of generic name indicated


[L02] Li F. 2002. Psocoptera of China vol. 1. Science Press: Beijing.

[M74] Mockford, E. L. 1974. Records and descriptions of Cuban Psocoptera. Entomologica Americana 48 (2): 103-215.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[SN04] Schmidt, E. R., & T. R. New. 2004. A systematic and phylogenetic revision of the family Elipsocidae (Insecta: Psocoptera), with the erection of two new families: Lesneiidae and Sabulopsocidae. Invertebrate Systematics 18: 157-213.

[Y02] Yoshizawa, K. 2002. Phylogeny and higher classification of suborder Psocomorpha (Insecta: Psocodea: ‘Psocoptera’). Zoological Journal of the Linnean Society 136: 371-400.

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