Psocomorpha

Archipsocus floridanus, photographed by Ben Van Allen.


Belongs within: Psocoptera.
Contains: Psocetae, Homilopsocidea, Caeciliusetae, Epipsocetae.

The Psocomorpha is a major subgroup of the barklice, characterised by the presence of non-annulated antennae, and in winged forms of a sclerotised pterostigma.

Characters (from Yoshizawa 2002): Head usually with rounded vertex; antennae 13-segmented; flagellomeres never annulated; maxilla without stipito-galeal muscle; hypopharyngeal filaments fused for most of their length; labial palpus 1-segmented. Forewing with nodus and thickened pterostigma; nodulus hook-shaped, formed by truncated spines fused at their base; CuP ending together with A1 at wing margin. Apex of first axillary sclerite usually with minute proximal process. Subgenital plate lacking median sclerite.

<==Psocomorpha [Eupsocida, Psociformia]
    |--Archipsocidae [Archipsocetae, Archipsocoidea] Y02
    |    |--Pararchipsocus pacificus Y02
    |    |--Pseudarchipsocus Mockford 1974 M74
    |    |    `--*P. guajiro Mockford 1974 M74
    |    |--Archipsocopsis Badonnel 1948 M93
    |    |    |--*A. mendax M93
    |    |    |--A. fernandi L02
    |    |    |--A. frater (Mockford 1957) [=Archipsocus frater] M93
    |    |    |--A. inornata M93
    |    |    `--A. parvulus (Mockford 1953) L02, M93 [=Archipsocus (Archipsocopsis) parvulus M93]
    |    `--Archipsocus Hagen 1882 M93
    |         |--*A. puber Hagen 1882 L02
    |         |--A. albofasciatus Badonnel 1949 B55
    |         |--A. aneura Badonnel 1948 B55
    |         |--A. badonneli B86
    |         |--A. bicolor B55
    |         |--A. castrii B86
    |         |--A. floridanus Mockford 1953 M93
    |         |--A. fuscopalpus Badonnel 1955 B55
    |         |--A. ghesquierei Badonnel 1946 (see below for synonymy) B55
    |         |    |--A. g. ghesquierei B55
    |         |    `--A. g. longicornis Badonnel 1955 B55
    |         |--A. gurneyi Mockford 1953 M93
    |         |--A. machadoi Badonnel 1955 B55
    |         |--A. mendax Badonnel 1948 B55
    |         |--A. minutus Badonnel 1955 B55
    |         |--A. modestus B86
    |         |--A. nomas Gurney 1939 M93
    |         |--A. panama Gurney 1939 M74
    |         |--A. polytrichus Li 2002 L02
    |         |--A. recens Enderlein 1903 L02
    |         `--A. sanurensis Thornton 1984 L02
    `--+--+--Psocetae Y02
       |  `--+--Homilopsocidea Y02
       |     `--+--Caeciliusetae Y02
       |        `--Epipsocetae Y02
       `--Hemipsocidae [Hemipsocetae, Hemipsocoidea] Y02
            |--Anopistoscena Y02
            |--Cyclohemipsocus Li 2002 L02
            |    `--*C. chinensis Li 2002 L02
            |--Hemipsocus Selys-Longchamps 1872 M93
            |    |--*H. chloroticus (Hagen 1858) [=Psocus chloroticus; incl. H. hyalinus Enderlein 1906] M93
            |    |--H. africanus Enderlein 1907 E07
            |    |--H. funebris Badonnel 1969 L02
            |    |--H. luridus Enderlein 1903 (see below for synonymy) S95
            |    |--H. maritiensis Turner 1976 L02
            |    |--H. massulatus Li 2002 L02
            |    |--H. parallelicus Li 1996 L02
            |    `--H. pretiosus Banks 1930 M93
            `--Metahemipsocus Li 1995 L02
                 |--*M. longicornis Li 1995 L02
                 |--M. bellatulus Li 2002 L02
                 |--M. bicuspidatus Li 2002 L02
                 |--M. bimaculatus Li 2002 L02
                 |--M. cuneatus Li 1996 L02
                 |--M. flabellatus Li 2002 L02
                 |--M. guangxiensis Li 2002 L02
                 |--M. interaus Li 1995 L02
                 |--M. octofarius Li 1999 L02
                 |--M. recurvicornis (Li 1992) [=Hemipsocus recurvicornis] L02
                 |--M. scitulus Li 2002 L02
                 |--M. spilopterus Li 1996 L02
                 |--M. tenuatus Li 1997 L02
                 |--M. triangularis Li 2002 L02
                 |--M. trimerus Li 2002 L02
                 |--M. vitellinus Li 2002 L02
                 `--M. yunnanicus Li 2002 L02

Archipsocus ghesquierei Badonnel 1946 [incl. A. ghesquierei var. albomaculatus Badonnel 1955, A. neens f. macropterus Badonnel 1946] B55

Hemipsocus luridus Enderlein 1903 [=H. chloroticus var. luridus, H. selysianus luridus; incl. H. selysianus Enderlein 1919, H. luridus selysianus] S95

*Type species of generic name indicated

REFERENCES

[B55] Badonnel, A. 1955. Psocoptères de l'Angola. Diamang Publicações Culturais 26: 11-267.

[B86] Badonnel, A. 1986. Psocoptères de Colombie (Insecta, Psocoptera). Spixiana 9 (2): 179-223.

[E07] Enderlein, G. 1907. Wissenschaftliche Ergebnisse der Schwedischen Zoologischen Expedition nach dem Kilimandjaro, dem Meru und den Umgebenden Massaisteppen Deutsch-Ostafrikas 1905–1906, vol. 15. Corrodentia, pt 2. Copeognatha. Almqvist & Wiksells Boktryckeri-A. B.: Uppsala.

[L02] Li F. 2002. Psocoptera of China vol. 1. Science Press: Beijing.

[M74] Mockford, E. L. 1974. Records and descriptions of Cuban Psocoptera. Entomologica Americana 48 (2): 103-215.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[S95] Smithers, C. N. 1995. Psocoptera (Insecta) of Christmas Island. Invertebrate Taxonomy 9: 529-561.

[Y02] Yoshizawa, K. 2002. Phylogeny and higher classification of suborder Psocomorpha (Insecta: Psocodea: ‘Psocoptera’). Zoological Journal of the Linnean Society 136: 371-400.

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