Black flat bug Aradus niger, copyright Sam Houston.

Belongs within: Panheteroptera.

The Aradoidea are a group of flattened bugs, many species of which are found on or under bark of dead or dying trees (Grimaldi & Engel 2005). When wings are present, the sides of the abdomen extend outside the margins of the wings at rest (Carver et al. 1991).

Synapomorphies (from Shcherbakov & Popov 2002): Body strongly flattened dorsoventrally; extremely long stylets coiled within head; abdominal trichobothria absent.

    |--Kobdocoridae SP02
    |--Termitaradus [Termitaphididae] GE05
    |    `--T. australiensis CGW91
    `--Aradidae GE05
         |  i. s.: Clavaptera ornata Kirman 1985 L02
         |         Leuraptera yakasi Heiss 1990 L02
         |         Neocarventus uncus Kirman 1989 L02
         |         Archearadus burmensis GE05
         |         Odonia Bergroth 1889 B89
         |           `--*O. truncata [=Aradus truncatus] B89
         |         Artabanus B89
         |           |--A. atkinsoni Bergroth 1889 B89
         |           |--A. breviscutum Bergroth 1889 B89
         |           `--A. geniculatus B89
         |         Brachyrrhynchus B89
         |           |--B. feanus Bergroth 1889 B89
         |           |--B. membranaceus [incl. B. albipennis, B. orientalis] B89
         |           |--B. montanus Bergroth 1889 B89
         |           |--B. scabrosus B89
         |           |--B. setosus B89
         |           |--B. tagalicus Stål 1870 B89
         |           `--B. triangulus Bergroth 1889 B89
         |         Llaimacoris WEE74
         |         Emydocoris Usinger 1944 D56
         |           |--E. montanus Drake 1956 D56
         |           `--E. testidinatus D56
         |         Reeceicus Drake 1956 D56
         |           `--*R. saileri Drake 1956 D56
         |         Glyptocoris verus Drake 1956 D56
         |         Aglaocoris D56
         |           |--A. cubanus Drake 1956 D56
         |           |--A. comes Drake 1956 D56
         |           `--A. natalii D56
         |         Asterocoris (Peggicoris Drake 1956) D56
         |           `--A. (*P.) zeteki Drake 1956 D56
         |--Prosympiestus [Prosympiestinae] CGW91
         |--Isodermus [Isoderminae] CGW91
         |    `--I. planus CGW91
         |--Chinamyersiinae CGW91
         |    |--Kumaressa scutellata CGW91
         |    `--Tretocoris CGW91
         |--Calisius L02 [Calisiinae CGW91]
         |    |--C. interveniens K08
         |    |--C. pacificus Kirkaldy 1908 K08
         |    `--C. zealandicus Pendergrast 1968 L02
         |--Carventinae CGW91
         |    |--Carventus CGW91
         |    |--Glyptoaptera CGW91
         |    |--Probatoceps CGW91
         |    `--Acaraptera waipouensis Heiss 1990 CGW91, L02
         |--Aneurus L02 [Aneurinae CGW91]
         |    |  i. s.: A. prominens Pendergrast 1965 L02
         |    `--A. (Aneurodellus) L02
         |         |--A. (A.) brevipennis Heiss 1998 L02
         |         |--A. (A.) maoricus Heiss 1998 L02
         |         `--A. (A.) zealandensis Heiss 1998 L02
         |--Aradus [Aradinae] CGW91
         |    |  i. s.: A. abnormis Bergroth 1889 B89
         |    |         A. candidatus Bergroth 1889 B89
         |    |         A. flavicornis B89
         |    |         A. frigidus Kiritshenko 1913 TKM93
         |    |         A. lugubris B89
         |    |         A. varius B49
         |    `--A. (Quilnus) H01
         |         |--A. (Q.) brevirostris Horváth 1901 H01
         |         `--A. (Q.) niger H01
         `--Mezirinae CGW91
              |--Mezira thoracoceras [=Brachyrrhynhus thoracoceras] K08
              |--Neuroctenus par Bergr. 1887 CGW91, B89
              |--Aspisocoris termitophilus CGW91
              |--Chiastoplonia CGW91
              |--Glochocoris CGW91
              |--Clavicornia CGW91
              |--Drakiessa CGW91
              |--Chelanoderus CGW91
              |--Caecicoris CGW91
              `--Usingerida CGW91

*Type species of generic name indicated


[B49] Baker, E. W. 1949. A review of the mites of the family Cheyletidae in the United States National Museum. Proceedings of the United States National Museum 99 (3238): 267–320.

[B89] Bergroth, E. 1889. Viaggio di Leonardo Fea in Birmania e regioni vicine. XXII.—E. Bergrothii Commentarius de Aradidis in Burma et Tenasserim a L. Fea collectis. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 730–739.

[CGW91] Carver, M., G. F. Gross & T. E. Woodward. 1991. Hemiptera (bugs, leafhoppers, cicadas, aphids, scale insects etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1 pp. 429–509. Melbourne University Press: Carlton (Victoria).

[D56] Drake, C. J. 1956. New Neotropical genera and species of apterous aradids (Hemiptera). Journal of the Washington Academy of Sciences 46 (10): 322–327.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[H01] Horváth, G. 1901. Hemipterák [Hemiptera]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 245–274. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[K08] Kirkaldy, G. W. 1908. A catalogue of the Hemiptera of Fiji. Proceedings of the Linnean Society of New South Wales 33: 345–391, pl. 4.

[L02] Larivière, M.-C. 2002. Primary types of Heteroptera in the New Zealand Arthropod Collection: Pentatomomorpha described from New Zealand before the year 2000. New Zealand Entomologist 25: 79–85.

[SP02] Shcherbakov, D. E., & Yu. A. Popov. 2002. Superorder Cimicidea Laicharting, 1781: Order Hemiptera Linné, 1758. The bugs, cicadas, plantlice, scale insects, etc. [=Cimicida Laicharting, 1781, =Homoptera Leach, 1815+Heteroptera Latreille, 1810). In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 143–157. Kluwer Academic Publishers: Dordrecht.

[TKM93] Thaler, K., B. Knoflach & E. Meyer. 1993. Fragmenta faunistica Tirolensia—X. Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck 80: 311-325.

[WEE74] Woodward, T. E., V. F. Eastop & J. W. Evans. 1974. Hemiptera (bugs, leafhoppers, etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers. Supplement 1974 pp. 52–57. Melbourne University Press.

Last updated: 9 October 2017.

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