Aradus lugubris, photographed by Boris Loboda.

Belongs within: Panheteroptera.

The Aradoidea are a group of flattened bugs, many species of which are found on or under bark of dead or dying trees (Grimaldi & Engel 2005).

Synapomorphies (from Shcherbakov & Popov 2002): Body strongly flattened dorsoventrally; extremely long stylets coiled within head; abdominal trichobothria absent.

    |--Kobdocoridae SP02
    |--Termitaradus [Termitaphididae] GE05
    `--Aradidae GE05
         |--Acaraptera waipouensis Heiss 1990 L02
         |--Calisius zealandicus Pendergrast 1968 L02
         |--Clavaptera ornata Kirman 1985 L02
         |--Leuraptera yakasi Heiss 1990 L02
         |--Neocarventus uncus Kirman 1989 L02
         |--Neuroctenus par Bergr. 1887 B89
         |--Archearadus burmensis GE05
         |--Odonia Bergroth 1889 B89
         |    `--*O. truncata [=Aradus truncatus] B89
         |--Artabanus B89
         |    |--A. atkinsoni Bergroth 1889 B89
         |    |--A. breviscutum Bergroth 1889 B89
         |    `--A. geniculatus B89
         |--Aradus B89
         |    |--A. abnormis Bergroth 1889 B89
         |    |--A. candidatus Bergroth 1889 B89
         |    |--A. flavicornis B89
         |    |--A. lugubris B89
         |    `--A. varius B49
         |--Aneurus L02
         |    |  i. s.: A. prominens Pendergrast 1965 L02
         |    `--A. (Aneurodellus) L02
         |         |--A. (A.) brevipennis Heiss 1998 L02
         |         |--A. (A.) maoricus Heiss 1998 L02
         |         `--A. (A.) zealandensis Heiss 1998 L02
         `--Brachyrrhynchus B89
              |--B. feanus Bergroth 1889 B89
              |--B. membranaceus [incl. B. albipennis, B. orientalis] B89
              |--B. montanus Bergroth 1889 B89
              |--B. scabrosus B89
              |--B. setosus B89
              |--B. tagalicus Stål 1870 B89
              `--B. triangulus Bergroth 1889 B89

*Type species of generic name indicated


[B49] Baker, E. W. 1949. A review of the mites of the family Cheyletidae in the United States National Museum. Proceedings of the United States National Museum 99 (3238): 267-320.

[B89] Bergroth, E. 1889. Viaggio di Leonardo Fea in Birmania e regioni vicine. XXII.—E. Bergrothii Commentarius de Aradidis in Burma et Tenasserim a L. Fea collectis. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a 7: 730-739.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[L02] Larivière, M.-C. 2002. Primary types of Heteroptera in the New Zealand Arthropod Collection: Pentatomomorpha described from New Zealand before the year 2000. New Zealand Entomologist 25: 79-85.

[SP02] Shcherbakov, D. E., & Yu. A. Popov. 2002. Superorder Cimicidea Laicharting, 1781: Order Hemiptera Linné, 1758. The bugs, cicadas, plantlice, scale insects, etc. [=Cimicida Laicharting, 1781, =Homoptera Leach, 1815 + Heteroptera Latreille, 1810). In History of Insects (A. P. Rasnitsyn & D. L. J. Quicke, eds) pp. 143-157. Kluwer Academic Publishers: Dordrecht.

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