Black flat bug Aradus niger, copyright Sam Houston.

Belongs within: Panheteroptera.

The Aradoidea are a group of flattened bugs, many species of which are found on or under bark of dead or dying trees (Grimaldi & Engel 2005). When wings are present, the sides of the abdomen extend outside the margins of the wings at rest. Most living species are included in the family Aradidae with the exception of the Termitaphididae, a circumtropical group of blind, wingless species that live in termite nests (Carver et al. 1991).

Synapomorphies (from Shcherbakov & Popov 2002): Body strongly flattened dorsoventrally; extremely long stylets coiled within head; abdominal trichobothria absent.

    |--Kobdocoris RJ93 [Kobdocoridae SP02]
    |    `--K. aradinus Popov 1986 RJ93
    |--Termitaradus [Termitaphididae] GE05
    |    `--T. australiensis CGW91
    `--Aradidae GE05
         |  i. s.: Clavaptera ornata Kirman 1985 L02
         |         Leuraptera yakasi Heiss 1990 L02
         |         Neocarventus uncus Kirman 1989 L02
         |         Archearadus burmensis GE05
         |         Odonia Bergroth 1889 B89
         |           `--*O. truncata [=Aradus truncatus] B89
         |         Artabanus B89
         |           |--A. atkinsoni Bergroth 1889 B89
         |           |--A. breviscutum Bergroth 1889 B89
         |           `--A. geniculatus B89
         |         Brachyrrhynchus B89
         |           |--B. feanus Bergroth 1889 B89
         |           |--B. membranaceus [incl. B. albipennis, B. orientalis] B89
         |           |--B. montanus Bergroth 1889 B89
         |           |--B. scabrosus B89
         |           |--B. setosus B89
         |           |--B. tagalicus Stål 1870 B89
         |           `--B. triangulus Bergroth 1889 B89
         |         Llaimacoris WEE74
         |         Emydocoris Usinger 1944 D56
         |           |--E. montanus Drake 1956 D56
         |           `--E. testidinatus D56
         |         Reeceicus Drake 1956 D56
         |           `--*R. saileri Drake 1956 D56
         |         Glyptocoris verus Drake 1956 D56
         |         Aglaocoris D56
         |           |--A. cubanus Drake 1956 D56
         |           |--A. comes Drake 1956 D56
         |           `--A. natalii D56
         |         Asterocoris (Peggicoris Drake 1956) D56
         |           `--A. (*P.) zeteki Drake 1956 D56
         |--Prosympiestus [Prosympiestinae] CGW91
         |--Isodermus [Isoderminae] CGW91
         |    `--I. planus CGW91
         |--Chinamyersiinae CGW91
         |    |--Kumaressa scutellata CGW91
         |    `--Tretocoris CGW91
         |--Calisius L02 [Calisiinae CGW91]
         |    |--C. interveniens K08
         |    |--C. pacificus Kirkaldy 1908 K08
         |    `--C. zealandicus Pendergrast 1968 L02
         |--Carventinae CGW91
         |    |--Carventus CGW91
         |    |--Glyptoaptera CGW91
         |    |--Probatoceps CGW91
         |    `--Acaraptera waipouensis Heiss 1990 CGW91, L02
         |--Aneurus L02 [Aneurinae CGW91]
         |    |  i. s.: A. prominens Pendergrast 1965 L02
         |    `--A. (Aneurodellus) L02
         |         |--A. (A.) brevipennis Heiss 1998 L02
         |         |--A. (A.) maoricus Heiss 1998 L02
         |         `--A. (A.) zealandensis Heiss 1998 L02
         |--Aradus TKM93 [Aradinae CGW91]
         |    |  i. s.: A. abnormis Bergroth 1889 B89
         |    |         A. betulae G20
         |    |         A. candidatus Bergroth 1889 B89
         |    |         A. flavicornis B89
         |    |         A. frigidus Kiritshenko 1913 TKM93
         |    |         A. lugubris Boisduval 1835 B35
         |    |         A. nicholasi Popov 1989 RJ93
         |    |         A. varius B49
         |    `--A. (Quilnus) H01
         |         |--A. (Q.) brevirostris Horváth 1901 H01
         |         `--A. (Q.) niger H01
         `--Mezirinae [Dysodiinae] CGW91
              |--Neuroctenus par Bergr. 1887 CGW91, B89
              |--Aspisocoris termitophilus CGW91
              |--Chiastoplonia CGW91
              |--Glochocoris CGW91
              |--Clavicornia CGW91
              |--Drakiessa CGW91
              |--Chelanoderus CGW91
              |--Caecicoris CGW91
              |--Usingerida CGW91
              `--Mezira D-VTM96
                   |--M. froeschneri Davidová-Vilímová, Taylor & McPherson 1996 D-VTM96
                   |--M. granulata D-VTM96
                   |--M. novella D-VTM96
                   |--M. sayi Kormilev 1982 D-VTM96
                   |--M. smithi Kormilev 1982 D-VTM96
                   `--M. thoracoceras [=Brachyrrhynhus thoracoceras] K08

*Type species of generic name indicated


[B49] Baker, E. W. 1949. A review of the mites of the family Cheyletidae in the United States National Museum. Proceedings of the United States National Museum 99 (3238): 267–320.

[B89] Bergroth, E. 1889. Viaggio di Leonardo Fea in Birmania e regioni vicine. XXII.—E. Bergrothii Commentarius de Aradidis in Burma et Tenasserim a L. Fea collectis. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 730–739.

[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d'Urville. Faune entomologique de l'océan Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.

[CGW91] Carver, M., G. F. Gross & T. E. Woodward. 1991. Hemiptera (bugs, leafhoppers, cicadas, aphids, scale insects etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1 pp. 429–509. Melbourne University Press: Carlton (Victoria).

[D-VTM96] Davidová-Vilímová, J., S. J. Taylor & J. E. McPherson. 1996. A new species of Mezira Amyot and Serville (Heteroptera: Aradidae) from Florida, with a key to the small Mezira species of America north of Mexico. Proceedings of the Entomological Society of Washington 98 (4): 630–639.

[D56] Drake, C. J. 1956. New Neotropical genera and species of apterous aradids (Hemiptera). Journal of the Washington Academy of Sciences 46 (10): 322–327.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[H01] Horváth, G. 1901. Hemipterák [Hemiptera]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 245–274. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[K08] Kirkaldy, G. W. 1908. A catalogue of the Hemiptera of Fiji. Proceedings of the Linnean Society of New South Wales 33: 345–391, pl. 4.

[L02] Larivière, M.-C. 2002. Primary types of Heteroptera in the New Zealand Arthropod Collection: Pentatomomorpha described from New Zealand before the year 2000. New Zealand Entomologist 25: 79–85.

[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.

[SP02] Shcherbakov, D. E., & Yu. A. Popov. 2002. Superorder Cimicidea Laicharting, 1781: Order Hemiptera Linné, 1758. The bugs, cicadas, plantlice, scale insects, etc. [=Cimicida Laicharting, 1781, =Homoptera Leach, 1815+Heteroptera Latreille, 1810). In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 143–157. Kluwer Academic Publishers: Dordrecht.

[TKM93] Thaler, K., B. Knoflach & E. Meyer. 1993. Fragmenta faunistica Tirolensia—X. Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck 80: 311–325.

[WEE74] Woodward, T. E., V. F. Eastop & J. W. Evans. 1974. Hemiptera (bugs, leafhoppers, etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers. Supplement 1974 pp. 52–57. Melbourne University Press.

Last updated: 13 May 2022.

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