Cepheidae

Cepheus corae, from here.

Belongs within: Brachypylina.

The Cepheidae are a family of often relatively large oribatid mites, usually with a sculptured integument. Species of Cepheus can be found in moss, litter and rotting wood. Cepheus latus is a borer in birch twigs (Norton & Behan-Pelletier 2009). The genera Ommatocepheus, Eupterotegaeus, Conoppia, Sphodrocepheus and Tritegeus have tridactylous legs; the remaining genera are monodactyle (Balogh & Balogh 1992).

Characters (from Norton & Behan-Pelletier 2009): Length 500–1200 µm. Integument usually sculptured. Subcapitulum diarthritic; chelicerae chelate-dentate; subcapitular mentum without tectum. Prodorsum with true lamellae. Scalps absent. Lenticulus absent. Notogaster without setae centrodorsally, all notogastral setae in marginal or posteromarginal position. Three pairs of adanal setae, positioned adanally and postanally. Tarsus I with two solenidia.

Cepheidae S04
    |--Protocepheus Jacot 1928 S04
    |    `--*P. hericius (Michael 1887) S04 [=Tegeocranus hericius S04, Cepheus hericius BN82]
    |--Tikizetes Hammer 1967 S04
    |    `--*T. spinipes Hammer 1967 S04
    |--Pilocepheus Pérez-Íñigo 1992 S04
    |    `--*P. azoricus Pérez-Íñigo 1992 S04
    |--Conoppia Berlese 1908 [incl. Balzania Jacot 1929 non Spegazzini 1898 (ICBN), Phyllotegeus Berlese 1913] S04
    |    |--C. palmicincta (Michael 1884) S04 (see below for synonymy)
    |    `--C. setiformis Golosova & Karppinen 1985 S04
    |--Oribatodes Banks 1895 S04
    |    |--*O. mirabilis Banks 1895 S04 (see below for synonymy)
    |    `--O. crenulatus Csiszár 1962 S04
    |--Tereticepheus Bernini & Bernini 1990 S04
    |    |--*T. annamariae Bernini & Bernini 1990 S04
    |    `--T. undulatus (Willmann 1939) [=Cepheus undulatus] S04
    |--Hypocepheus Krivolutsky 1971 S04
    |    |--*H. mirabilis Krivolusky 1971 S04
    |    |--H. helveticus Mahunka & Mahunka-Papp 2002 S04
    |    `--H. krivolutskyi Calugar & Vasiliu 1976 S04
    |--Reticulocepheus Vasiliu & Calugar 1977 [incl. Cubacepheus Balogh & Mahunka 1979, Geocepheus Sitnikova 1979] S04
    |    |--*R. decoui Vasiliu & Calugar 1977 (see below for synonymy) S04
    |    |--R. areolatus (Sitnikova 1979) [=Geocepheus areolatus] S04
    |    |--R. grandis (Sitnikova 1979) [=Geocepheus grandis] S04
    |    `--R. reticulatus (Sitnikova 1979) [=Geocepheus reticulatus] S04
    |--Tritegeus Berlese 1913 S04
    |    |--T. bisulcatus Grandjean 1953 [incl. Cepheus bifidatus Michael 1880 non Nicolet 1855, *T. bifidatus] S04
    |    |--T. brevisetus Sitnikova 1975 S04
    |    |--T. major Golosova & Karppinen 1984 S04
    |    `--T. sculptus Bernini & Bernini 1990 S04
    |--Sphodrocepheus Woolley & Higgins 1963 S04
    |    |--*S. tridactylus Woolley & Higgins 1963 S04
    |    |--S. anthelionus Woolley & Higgins 1968 S04
    |    |--S. dentatus Fujikawa 1972 S04
    |    |--S. mitratus Aoki 1967 S04
    |    `--S. tuberculatus Mahunka 1988 S04
    |--Compactozetes Hammer 1966 [Compactozetidae] S04
    |    |--*C. rotoruensis Hammer 1966 S04
    |    |--C. hastatus Hammer 1973 S04
    |    |--C. niger Hammer 1966 S04
    |    |--C. serratus Balogh 1970 S04
    |    `--C. zeugus Luxton 1988 S04
    |--Ommatocepheus Berlese 1913 S04
    |    |--*O. ocellatus (Michael 1882) [=Cepheus ocellatus; incl. Scutovertex pulcherrimus Berlese 1913] S04
    |    |--O. clavatus Woolley & Higgins 1964 S04
    |    |    |--O. c. clavatus S04
    |    |    `--O. c. japonicus Aoki 1974 S04
    |    |--O. crassisetosus Pérez-Íñigo & Peña 1996 S04
    |    |--O. parvilamellatus Pérez-Íñigo & Pérez-Íñigo 1996 S04
    |    `--O. reticulatus Pérez-Íñigo & Peña 1997 S04
    |--Eupterotegaeus Berlese 1916 PL17, S04 [incl. Diodontocepheus Mihelčič 1958 S04]
    |    |--*E. ornatissimus (Berlese 1908) (see below for synonymy) S04
    |    |--E. armatus Aoki 1969 S04
    |    |--E. dentatus Sitnikova 1979 S04
    |    |--E. flavus (Ewing 1918) [=Cepheus flavus] S04
    |    |--E. nasalis Sitnikova 1979 S04
    |    |--E. pseudosculptus (Coggi 1900) [=Cepheus pseudosculptus] S04
    |    |--E. rhamphosus Higgins & Woolley 1968 S04
    |    |--E. rostratus Higgins & Woolley 1963 S04
    |    |--E. spinatus Higgins & Woolley 1963 S04
    |    `--E. xizangensis Wang & Solhoy 2001 S04
    |--Sadocepheus Aoki 1965 [incl. Hamotegeus Balogh & Mahunka 1969] S04
    |    |--*S. undulatus Aoki 1965 S04
    |    |    |--S. u. undulatus S04
    |    |    `--S. u. setiger Fujita & Fujikawa 1986 S04
    |    |--S. breviseta (Balogh 1986) [=Hamotegeus breviseta] S04
    |    |--S. dubius Hammer 1979 S04
    |    |--S. elevatus Mahunka 1987 S04
    |    |--S. foveolatus Luxton 1988 S04
    |    |--S. franzi (Balogh 1986) [=Hamotegeus franzi] S04
    |    |--S. granulatus (Balogh & Mahunka 1969) S04 [=*Hamotegeus granulatus BB92a]
    |    |--S. longiseta (Balogh 1986) [=Hamotegeus longiseta] S04
    |    |--S. makarcevae Sitnikova 1975 S04
    |    `--S. subniger (Ewing 1917) [=Tegeocranus subniger] S04
    `--Cepheus Koch 1835 [incl. Tegeocranus Nicolet 1855] S04
         |--*C. latus Koch 1835 S04 [=Tegeocranus latus BN82]
         |--C. brachiatus Sitnikova 1975 S04
         |--C. caucasicus Sitnikova 1975 S04
         |--C. cepheiformis (Nicolet 1855) [=Tegeocranus cepheiformis; incl. C. michaeli Jacot 1928] S04
         |--C. corae Jacot 1928 S04
         |--C. dentatus (Michael 1888) [=Tegeocranus dentatus; incl. C. grandis Sitnikova 1975] S04
         |--C. feideri Suciu & Panu 1972 S04
         |--C. gracilipes (Canestrini 1897) (n. d.) [=Tegeocranus gracilipes] S04
         |--C. granulatus Mihelčič 1958 S04
         |--C. heterosetosus (Sitnikova 1975) [=Oribatodes heterosetosus] S04
         |--C. hokkaiensis Fujikawa 1992 S04
         |--C. incisus Mihelčič 1958 S04
         |--C. jindingensis Wen & Sun 1990 S04
         |--C. kurosawai Aoki 1986 S04
         |--C. lobatus Mihelčič 1953 (n. d.) S04
         |--C. mirabiloides Jacot 1938 S04
         |--C. ovalis CF77
         |--C. pegazzanoae Bernini & Nannelli 1982 S04
         |--C. pertusus Haupt 1882 (n. d.) S04
         |--C. pustulatus (Pearce 1910) [=Tegeocranus pustulatus] S04
         |--C. takasago Aoki 1991 S04
         |--C. transversalis Balogh 1958 (n. d.) S04
         |--C. transylvanicus Cooreman 1951 S04
         |--C. tuberculosus Strenzke 1951 S04
         `--C. verrucosus Bernini 1971 S04

Conoppia palmicincta (Michael 1884) S04 [=Leiosoma palmicinctum S04, *Phyllotegeus palmicinctus G36; incl. Oppia microptera Berlese 1885 S04, *Conoppia microptera S04, Oribata microptera Canestrini 1896 non Oppia microptera Berlese 1885 S04]

*Eupterotegaeus ornatissimus (Berlese 1908) [=Tegeocranus ornatissimus; incl. Diodontocepheus steinboecki Mihelčič 1958] S04

*Oribatodes mirabilis Banks 1895 S04 [=Cepheus mirabilis BN82; incl. C. lamellatus Banks 1906 S04, Tegeocranus longisetus Berlese 1910 S04]

*Reticulocepheus decoui Vasiliu & Calugar 1977 [incl. Cepheus (Cubacepheus) lobatus Balogh & Mahunka 1979 non C. lobatus Mihelčič 1953] S04

*Type species of generic name indicated

REFERENCES

Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[BN82] Bernini, F., & R. Nannelli. 1982. Notulae oribatologicae XXVI. Contribution to the knowledge of the genus Cepheus (Oribatida, Acarida) in Italy. Redia 45: 155–188.

[CF77] Canestrini, G., & F. Fanzago. 1877. Intorno agli Acari Italiani. Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti, Serie 5, 4: 69–208, pls 2–7.

[G36] Grandjean, F. 1936. Observations sur les oribates (10^e série). Bulletin du Muséum, 2e série, 8 (3): 246–253.

Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 430–564. Texas Tech University Press.

[PL17] Pachl, P., A. C. Lindl, A. Krause, S. Scheu, I. Schaefer & M. Maraun. 2017. The tropics as an ancient cradle of oribatid mite diversity. Acarologia 57 (2): 309–322.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Last updated: 30 June 2017.