Hafenrefferia sp., photographed by Pfiegler Walter.

Belongs within: Circumdehiscentiae.

The Tenuialidae are a family of oribatid mites characterised by the presence of a long humeral process. They are most abundant in forest litter in temperate regions of the Holarctic. Depressions in the podosoma allow members of this family to retract their legs (Norton & Behan-Pelletier 2009).

Characters (from Norton & Behan-Pelletier 2009): Length 800–1200 µm. Subcapitulum diarthric; chelicerae chelate-dentate. Prodorsum and notogaster smooth, shiny. Prodorsum with true lamellae. Notogaster with longhumeral processes projecting anterolaterally to bothridium; octotaxic system absent; dorsophragmata and pleurophragmata present. Podosomal region with depressions for retracted legs; epimeral border IV with minitectum.

    |--Ceratotenuiala Aoki & Maruyama 1983 S04
    |    `--*C. echigoensis Aoki & Maruyama 1983 S04
    |--Tenuialoides Woolley & Higgins 1966 S04
    |    |--*T. medialis Woolley & Higgins 1966 S04
    |    `--T. fusiformis Aoki 1969 [incl. Hafenferrefia translamellata Aoki & Fujikawa 1979] S04
    |--Hafenrefferia Oudemans 1906 S04
    |    |--*H. gilvipes (Koch 1839) (see below for synonymy) S04
    |    `--H. acuta Aoki 1966 S04
    |--Peltenuiala Norton 1983 S04
    |    |--*P. pacifica Norton 1983 S04
    |    `--P. orbiculata (Aoki & Ohnishi 1974) [=Hafenferrefia orbiculata] S04
    |--Tenuiala Ewing 1913 S04
    |    |--*T. nuda Ewing 1913 S04
    |    |--T. crenulata Woolley & Higgins 1966 S04
    |    `--T. kurti Woolley & Higgins 1955 S04
    `--Hafenferrefia Jacot 1939 [incl. Hafenrefferiella Sellnick 1952] S04
         |--*H. nitidula (Banks 1906) [=Galumna nitidula] S04
         |--H. hyrcanica (Krivolutsky 1967) [=Hafenrefferiella hyrcanica] S04
         |--H. imperialis (Berlese 1910) (n. d.) [=Achipteria imperialis] S04
         |--H. lata Golosova 1980 S04
         `--H. nevesi (Sellnick 1952) [=Hafenrefferiella nevesi] S04

*Haffenrefferia gilvipes (Koch 1839) [=Oribata gilvipes; incl. Liacarus auritus Nordenskiöld 1901, L. pterotus Coggi 1900] S04

*Type species of generic name indicated


Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology, 3rd ed., pp. 430-564. Texas Tech University Press.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

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