Liacaridae

Dorycranosus acutidens, photographed by David Porco.

Belongs within: Circumdehiscentiae.
Contains: Liacarus (Liacarus).

The Liacaridae are a family of oribatid mites with converging lamellae, and two pairs of humeral setae c1 and c2 placed near to each other. The lamellae remain well-separated in Planoristes acuspidatus but are fused apically in the remaining genera. Birsteinius has monodactyle legs whereas the legs of Opsioristes, Liacarus and Dorycranosus are tridactyle. Opsioristes eurhostus has five pairs of genital setae vs six in Liacarus and Dorycranosus (Balogh & Balogh 1992).

<==Liacaridae
    |--Opsioristes Woolley 1967 S04
    |    `--*O. eurhostus Woolley 1967 S04
    |--Planoristes Iturrondobeitia & Subías 1978 S04
    |    `--*P. acuspidatus Iturrondobeitia & Subías 1978 S04
    |--Birsteinius Krivolutsky 1965 S04
    |    |--*B. clavatus Krivolutsky 1965 S04
    |    |--B. krivolutskyi Rjabinin 1979 S04
    |    |--B. microchaetus Krivolutsky 1967 S04
    |    |--B. mongolicus (Mahunka 1964) [=Liacarus mongolicus] S04
    |    |--B. monticolus Wang & Shen 2001 S04
    |    |--B. perlongoides Hammer 1977 S04
    |    |--B. perlongus Krivolutsky 1965 S04
    |    `--B. trimucronatus Yamamoto & Aoki 2000 S04
    |--Liacarus Michael 1898 S04
    |    |  i. s.: L. globosus S22
    |    |--L. (Liacarus) S04
    |    |--L. (Procorynetes Woolley 1969) S04
    |    |    |--L. (*P.) nigerrimus Berlese 1916 S04
    |    |    |--L. (P.) andinus (Balogh 1984) [=Procorynetes andinus] S04
    |    |    |--L. (P.) breviclavatus Aoki 1970 S04
    |    |    |--L. (P.) clavatus Fujikawa & Aoki 1970 S04
    |    |    |--L. (P.) espeletiae (Balogh 1984) [=Procorynetes espeletiae] S04
    |    |    |--L. (P.) globifer (Kramer 1897) [=Leiosoma globiferum] S04
    |    |    `--L. (P.) tsendsureni (Bayartogtokh 1998) [=Procorynetes tsendsureni] S04
    |    `--L. (Rhaphidosus Woolley 1969 S04
    |         |--L. (*R.) carolinensis Banks 1906 S04
    |         |--L. (R.) acuminatus (Woolley 1969) [=Rhaphidosus acuminatus] S04
    |         |--L. (R.) alticola (Balogh 1984) [=Rhaphidosus alticola] S04
    |         |--L. (R.) bacillatus Fujikawa & Aoki 1970 S04
    |         `--L. (R.) carbonarius (Banks 1906) [=Notaspis carbonaria] S04
    `--Dorycranosus Woolley 1969 NB-P09, S04
         |--*D. abdominalis (Banks 1906) [=Liacarus abdominalis] S04
         |--D. acutidens (Aoki 1965) [=Liacarus (Dorycranosus) acutidens] S04
         |--D. acutus (Pschorn-Walcher 1951) (see below for synonymy) S04
         |--D. aequidentatus (Ewing 1918) [=Liacarus (Dorycranosus) aequidentatus] S04
         |--D. altaicus Krivolutsky 1974 [=Liacarus (Dorycranosus) altaicus] S04
         |--D. angustatus Weigmann 1976 [=Liacarus (Dorycranosus) angustatus] S04
         |--D. arcticus (Banks 1899) (n. d.) [=Oppia arctica, Liacarus (Dorycranosus) arcticus] S04
         |--D. badghysi (Krivolutsky 1966) [=Liacarus (Dorycranosus) badghysi] S04
         |--D. columbianus (Berlese 1908) [=Liacarus (Dorycranosus) columbianus] S04
         |--D. coronatus Golosova & Ljashchev 1984 [=Liacarus (Dorycranosus) coronatus] S04
         |--D. dickersoni Moraza 1990 [=Liacarus (Dorycranosus) dickersoni] S04
         |--D. floridensis (Berlese 1908) [=Liacarus (Dorycranosus) floridensis] S04
         |--D. frontinalis (Banks 1906) [=Liacarus (Dorycranosus) frontinalis] S04
         |--D. fusiformis (Ewing 1917) [=Liacarus (Dorycranosus) fusiformis] S04
         |--D. indentatus Aoki 1973 [=Liacarus (Dorycranosus) indentatus] S04
         |--D. keretinus (Nordenskiöld 1901) [=Liacarus (Dorycranosus) keretinus] S04
         |--D. latiusculus (Wen 1991) [=Liacarus (Dorycranosus) latiusculus] S04
         |--D. lingulatus Golosova & Ljashchev 1984 [=Liacarus (Dorycranosus) lingulatus] S04
         |--D. neonominatus (Subías 2004) (see below for synonymy) S04
         |--D. nitidus (Banks 1895) (see below for synonymy) S04
         |--D. paoliensis Woolley 1969 [=Liacarus (Dorycranosus) paoliensis] S04
         |--D. parallelus (Hammer 1967) [=Liacarus (Dorycranosus) parallelus] S04
         |--D. polychothomus (Wen 1991) [=Liacarus (Dorycranosus) polychothomus] S04
         |--D. pulcher Sergienko & Djaparidze 1981 [=Liacarus (Dorycranosus) pulcher] S04
         |--D. splendens (Coggi 1898) (see below for synonymy) S04
         |--D. tenuilamellatus (Hirauchi 1998) [=Liacarus (Dorycranosus) tenuilamellatus] S04
         |--D. triapicini Djaparidze 1983 [=Liacarus (Dorycranosus) triapicini] S04
         |--D. yezoensis (Fujikawa & Aoki 1970) [=Liacarus (Dorycranosus) yezoensis] S04
         `--D. zachvatkini (Kulijev 1962) [=Liacarus (Dorycranosus) zachvatkini; incl. D. ibericus Djaparidze 1973] S04

Dorycranosus acutus (Pschorn-Walcher 1951) [=Liacarus (Dorycranosus) acutus; incl. L. claviger Mihelčič 1956, Cultroribula grandis Mihelčič 1956, L. infissus Gunhold 1953] S04

Dorycranosus neonominatus (Subías 2004) [=Liacarus (Dorycranosus) neonominatus, Dorycranosus arcticus Grishina 1984 non Oppia arctica Banks 1899] S04

Dorycranosus nitidus (Banks 1895) [=Cepheus nitidus, Liacarus (Dorycranosus) nitidus; incl. L. nitidus fissuratus Jacot 1937, L. glaber Ewing 1909] S04

Dorycranosus splendens (Coggi 1898) [=Cepheus splendens, Liacarus (Dorycranosus) splendens; incl. L. alatus Berlese 1904, L. alatus curtipilis Willmann 1935, L. diversus Mihelčič 1956, L. ghilarovi Bashkirova 1958, L. moraviacus Willmann 1954, Dorycranosus ovatus Djaparidze 1973, L. punctulatus Mihelčič 1956] S04

*Type species of generic name indicated

REFERENCES

[BB92] Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[NB-P09] Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 430–564. Texas Tech University Press.

[S22] Schweizer, J. 1922. Beitrag zur Kenntnis der terrestrischen Milbenfauna der Schweiz. Verhandl. Naturf. Ges. Basel 33: 23–112, 4 pls.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.