Oestroidea

Dexopollenia flava, copyright @sabikikori5.


Belongs within: Muscoidea.
Contains: Miltogramminae, Paramacronychiinae, Sarcophaginae, Rhiniidae, Calliphoridae, Tachinidae, Rhinophoridae, Pollenia, Sepimentum.

The Oestroidea are a group of calyptrate flies united by features including a row of bristles on the meron and wings with vein M1 distinctly bent forward distally.

Synapomorphies (from Kutty et al. 2010): Meron bearing vertical row of bristles; anatergite bearing patch of setulae ventral to proximal rim of lower calypter; distal part of wing vein M deflected forwards to join C before wing apex; phallus bearing cuticular denticles on ventral surface of distal section.

<==Oestroidea [Calliphoroidea, Tachinoidea]
    |  i. s.: Ulurumyia Michelsen & Pape 2017 [Ulurumyiidae] MP17
    |           `--*U. macalpinei Michelsen & Pape 2017 MP17
    |--+--Mystacinobia Holloway 1976 KP10, PG-M04 [Mystacinobiidae]
    |  |    `--*M. zelandica Holloway 1976 PG-M04
    |  `--Sarcophagidae [Sarcophagoidea] KP10
    |       |  i. s.: Athyrsia L59
    |       |         *Mulsantia campestris L59
    |       |         Anica campestris M90
    |       |         Sphecapata conica BM76
    |       |         Opsidea BM76
    |       |         Amobiopsis BM76
    |       |--Miltogramminae KP10
    |       `--+--Paramacronychiinae KP10
    |          `--Sarcophaginae KP10
    `--+--Rhiniidae KP10
       `--+--Calliphoridae KP10
          `--+--+--Tachinidae KP10
             |  `--+--Rhinophoridae KP10
             |     `--Helicoboscinae [Helicoboscini] KP0
             |          |--Gulmargia Rognes 1993 PG-M04
             |          |    `--*G. angustisquama Rognes 1993 PG-M04
             |          `--Eurychaeta Brauer & Bergenstamm 1891 (see below for synonymy) PG-M04
             |               |--*E. muscaria (Meigen 1826) [=Sarcophaga muscaria] PG-M04
             |               `--E. palpalis (Robineau-Desvoidy 1830) KP10, PG-M04 (see below for synonymy)
             `--+--Stevenia KP10
                |    |  i. s.: S. subpellucida F92
                |    |--S. hertingi Kugler 1978 KP10
                |    `--+--S. atramentaria (Meigen 1824) KP10
                |       `--Paykullia maculata KP10
                `--Polleniinae [Polleniini] KP10
                     |--Pollenia PG-M04
                     |--Sepimentum PG-M04
                     |--Wilhelmina Schmitz & Villeneuve 1932 PG-M04
                     |    `--*W. nepenthicola Schmitz & Villeneuve 1932 PG-M04
                     |--Nepenthomyia Kurahashi & Beaver 1979 PG-M04
                     |    `--*N. malayana Kurahashi & Beaver 1979 PG-M04
                     |--Nesodexia Villeneuve 1911 PG-M04
                     |    `--*N. corsicana Villeneuve 1911 PG-M04
                     |--Anthracomyza Malloch 1927 [=Anthracomyia Malloch 1927 non Rondani 1856] PG-M04
                     |    `--*A. atratula (Malloch 1927) [=*Anthracomyia atratula] PG-M04
                     |--Melanodexia Williston 1893 [incl. Melanodexiopsis Hall 1948] PG-M04
                     |    |--*M. tristis Williston 1893 PG-M04
                     |    |--M. grandis S87
                     |    `--*Melanodexiopsis’ tristina Hall 1948 PG-M04
                     |--Morinia Robineau-Desvoidy 1830 (see below for synonymy) PG-M04
                     |    |--M. melanoptera (Fallén 1817) W87, PG-M04 (see below for synonymy)
                     |    |--M. nana F92
                     |    |--M. piliforaea PG-M04
                     |    `--M. slenfini PG-M04
                     |--Xanthotrixus Aldrich 1930 PG-M04
                     |    |--*X. mongol Aldrich 1930 PG-M04
                     |    |--X. bazini PG-M04
                     |    |--X. draco Aldrich 1930 F92
                     |    |--X. melanurus Fan 1992 F92
                     |    `--X. uniapicalis Fan 1992 F92
                     `--Dexopollenia Townsend 1917 [incl. Lispoparea Aldrich 1930] PG-M04
                          |--*D. testacea Townsend 1917 PG-M04
                          |--D. flava (Aldrich 1930) F92 [=*Lispoparea flava PG-M04]
                          |--D. geniculata Malloch 1935 F92
                          |--D. maculata Villeneuve 1933 F92
                          |--D. nigriscens Fan 1992 F92
                          `--D. uniseta Fan 1992 F92

Eurychaeta Brauer & Bergenstamm 1891 [incl. Helicobosca Bezzi 1906, Theria Robineau-Desvoidy 1830 (preoc.)] PG-M04

Eurychaeta palpalis (Robineau-Desvoidy 1830) KP10, PG-M04 [=*Theria palpalis PG-M04, *Helicobosca palpalis PG-M04]

Morinia Robineau-Desvoidy 1830 [incl. Anthracomyia Rondani 1856, Calobatemyia Macquart 1835, Disticheria Enderlein 1934] PG-M04

Morinia melanoptera (Fallén 1817) W87, PG-M04 [=Musca melanoptera PG-M04, Anthracomyia melanoptera W87, *Disticheria melanoptera PG-M04; incl. *Anthracomyia genei Rondani 1862 PG-M04, A. geneji PG-M04, *Calobatemyia nigra Macquart 1835 PG-M04, *Morinia velox Robineau-Desvoidy 1830 PG-M04]

*Type species of generic name indicated

REFERENCES

[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[F92] Fan Z. 1992. Key to the Common Flies of China 2nd ed. Science Press: Beijing.

[KP10] Kutty, S. N., T. Pape, B. M. Wiegmann & R. Meier. 2010. Molecular phylogeny of the Calyptratae (Diptera: Cyclorrhapha) with an emphasis on the superfamily Oestroidea and the position of Mystacinobiidae and McAlpine's fly. Systematic Entomology 35: 614–635.

[L59] Lopes, H. de S. 1959. A revision of Australian Sarcophagidae (Diptera). Studia Ent. 2 (1-4): 33–67.

[M90] McAlpine, J. F. 1990. Insecta: Diptera adults. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1211–1252. John Wiley & Sones: New York.

[MP17] Michelsen, V., & T. Pape. 2017. Ulurumyiidae—a new family of calyptrate flies (Diptera). Systematic Entomology 42: 826–836.

[PG-M04] Peris, S. V., & D. González-Mora. 2004. Clave de identificación para los géneros de Calliphoridae del mundo. Subfamilias con vena remigium desnuda y creación de una nueva subfamilia. Bol. R. Soc. Esp. Hist. Nat. (Sec. Biol.) 99 (1–4): 115–144.

[S87] Shewell, G. E. 1987. Calliphoridae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 1133–1145. Research Branch, Agriculture Canada.

[W87] Wood, D. M. 1987. Rhinophoridae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 1187–1191. Research Branch, Agriculture Canada.

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