Belongs within: Oestroidea.
The Paramacronychiinae are a group of sarcophagid flies combining a non-plumose arista with strong hairs on the meron. They are diverse in habits, including necrophagous species, predators of snails and parasites of other insects.
Characters (from Pape & Dahlem 2010): Head profile with convex occipital region; antennal arista not plumose; eyes not or only slightly enlarged; postocular setae alternating in size; notopleuron without subprimary bristles; metasternal area bare; hind coxa bare posteriorly; coxopleural streak present; male midfemur with apical posteroventral ctenidium of stout, flattened bristles; male abdominal sternites 2–4 partly hidden by overlapping margins of corresponding tergites, terminalia usually protruding, with tergite 6 distinctly fused to syntergosternite 7+8 at more or less right angle.
<==Paramacronychiinae PD10
|--Chrysogramma [Chrysogrammatini] F92
| `--C. variabile F92
`--+--Sarcotachiini F92
`--+--Goniophyto Townsend 1927 [incl. Sinagria Rohdendorf in Fan 1965; Goniophytoini] F92
| |--*G. formosensis Townsend 1927 F92
| `--G. honshuensis Rohdendorf 1962 F92
`--Paramacronychiini F92
|--+--+--Nyctiinina F92
| | `--+--Dexagriina F92
| | `--Blaesoxiphellina F92
| `--+--Paramacronychia [Paramacronychiina] F92
| | `--P. flavipalpis F92
| `--+--Galapagomyiina F92
| `--Agria Robineau-Desvoidy 1830 [Agriina] F92
| |--*A. punctata Robineau-Desvoidy 1830 F92
| |--A. housei S87
| `--A. xiangchengensis Chao & Zhang 1988 F92
`--Wohlfahrtia Brauer & Bergenstamm 1889 [Wohlfahrtiina] F92
|--*W. magnifica (Schiner 1862) [=Sarcophila magnifica; incl. S. wohlfahrti Portschinsky 1875] F92
|--W. atra Aldrich 1926 F92
|--W. balassogloi (Portschinsky 1882) [=Sarcophila balassogloi] F92
|--W. bella (Macquart 1838) [=Agria bella] F92
|--W. cheni Rohdendorf 1956 F92
|--W. fedtschenkoi Rohdendorf 1956 F92
|--W. intermedia (Portschinsky 1887) [=Sarcophila intermedia] F92
|--W. meigeni (Schiner 1862) [=Sarcophila meigeni] F92
|--W. nuba (Wiedemann 1830) KP10
|--W. pavlovskyi Rohdendorf 1956 F92
|--W. stackelbergi Rohdendorf 1956 F92
|--W. trina (Wiedemann 1830) KP10
`--W. vigil S87
Paramacronychiini incertae sedis:
Cattasoma PD10
Brachicoma Rondani 1856 F92
|--B. devia (Fallén 1820) (see below for synonymy) F92
|--B. asiatica Rohdendorf & Verves 1979 F92
|--B. nigra Chao & Zhang 1988 F92
`--B. setosa PD10
Erythandra picipes PD10, S87
Toxonagria montanensis S87
Sarcofahrtia ravinia S87
Nyctia lugubris (Macquart 1843) KP10
Sarcophila Rondani 1856 KP10
|--*S. latifrons (Fallén 1817) [=Musca latifrons] F92
|--S. japonica (Rohdendorf 1962) I92
|--S. meridionalis Verves 1982 KP10
|--S. mongolica Chao & Zhang 1988 F92
`--S. rasnitzyni Rohdendorf & Verves 1985 F92
Angiometopa Brauer & Bergenstamm 1889 F92
|--*A. ruralis (Fallén 1817) [=Musca ruralis] F92
`--A. hikosana F92
Wohlfahrtiodes Villeneuve 1910 F92
|--*W. nudus Villeneuve 1910 F92
`--W. mongolicus Chao & Zhang 1988 F92
Brachicoma devia (Fallén 1820) [=Tachina devia; incl. T. nitidula Rondani 1856 non Meigen 1824, *Brachicoma nitidula] F92
*Type species of generic name indicated
REFERENCES
[F92] Fan Z. 1992. Key to the Common Flies of China 2nd ed. Science Press: Beijing.
[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.
[KP10] Kutty, S. N., T. Pape, B. M. Wiegmann & R. Meier. 2010. Molecular phylogeny of the Calyptratae (Diptera: Cyclorrhapha) with an emphasis on the superfamily Oestroidea and the position of Mystacinobiidae and McAlpine's fly. Systematic Entomology 35: 614–635.
[PD10] Pape, T., & G. A. Dahlem. 2010. Sarcophagidae (flesh flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 1313–1335. NRC Research Press: Ottawa.
[S87] Shewell, G. E. 1987. Sarcophagidae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 1159–1186. Research Branch, Agriculture Canada.
Last updated: 5 July 2021.
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