Amobia signata, copyright Jeremy R.

Belongs within: Oestroidea.
Contains: Metopiini, Miltogrammatini.

The Miltogramminae are a group of flies whose larvae develop in the brood cells of burrowing Hymenoptera, feeding on the host's stored food reserves and/or the host larva itself (Shewell 1987).

Characters (from Shewell 1987, as Miltogrammini): Tegula usually black, contrasting with pale basicosta. Infrasquamal hairs absent. Meron with row of setae only. Coxopleural streak present. Mid femur of male lacking apical ctenidium. Cephalopharyngeal skeleton of first instar larva with single labral hook.

    |--+--Chorezmomyiini F92
    |  `--Metopiini F92
    `--+--Miltogrammatini F92
       `--+--Amobia Robineau-Desvoidy 1830 [incl. Pachyphthalmus Brauer & Bergenstamm 1889; Amobiini] F92
          |    |--*A. signata (Meigen 1824) [=Tachina signata] F92
          |    |--A. aurifrons BM76
          |    |--A. distorta (AllĂ©n 1926) [=Pachyophthalmus distortus] F92
          |    `--A. floridensis BM76
          `--Phyllotelini [Phyllotelina] F92
               |--Synorbitomyia Townsend 1932 F92
               |    `--*S. linearis (Villeneuve 1929) [=Hoplocephala linearis] F92
               `--Phylloteles Loew 1844 F92
                    |--*P. pictipennis Loew 1844 F92
                    `--P. stackelbergi Rohdendorf 1975 F92

Miltogramminae incertae sedis:
  Pachygraphomyia spinosa BM76
  Eumacronychia [Eumacronychiini] PD10
    |--E. polita [=Gymnoprosopa polita] PD10
    `--E. sternalis P09
  Macronychia Rondani 1859 PD10, F92 [Macronychiinae, Macronychiini]
    |--M. striginervis (Zetterstedt 1844) (see below for synonymy) F92
    |--M. aurata PD10
    |--M. griseola (Rondani 1859) [=Tachina griseola] F92
    `--M. polyodon (Meigen 1824) F92
  Opsidia PD10
    |--O. diademata PD10
    |--O. gonioides PD10
    `--O. metopioides PD10
  Euphytomima nomiivora S87
  Euphyto S87
  Eusenotainia rufiventris S87
  Gymnopsidia inflaticornis S87
  Opsidiotrophus micidus S87
  Opsidiopsis oblata S87
  Anicia campestris S87
  Aenigmetopia MD13

Macronychia striginervis (Zetterstedt 1844) [=Xysta striginervis; incl. *Macronychia agrestis Rondani 1859, Miltogramma ungulans Pandelle 1895, Ma. ungulans] F92

*Type species of generic name indicated


[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[F92] Fan Z. 1992. Key to the Common Flies of China 2nd ed. Science Press: Beijing.

[MD13] Meiklejohn, K. A., M. Dowton, T. Pape & J. F. Wallman. 2013. A key to the Australian Sarcophagidae (Diptera) with special emphasis on Sarcophaga (sensu lato). Zootaxa 3680 (1): 148–189.

[P09] Pape, T. 2009. Economic importance of Diptera. In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera, vol. 1 pp. 65–77. NRC Research Press: Ottawa.

[PD10] Pape, T., & G. A. Dahlem. 2010. Sarcophagidae (flesh flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 1313–1335. NRC Research Press: Ottawa.

[S87] Shewell, G. E. 1987. Sarcophagidae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 1159–1186. Research Branch, Agriculture Canada.

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