Scatopsoidea

Black compost fly Scatopse notata, copyright Miroslav Deml.


Belongs within: Bibionomorpha.

The Scatopsoidea are a clade of flies united by wings with M1+2 fused to Rs for a distance, and M3 and im lost; at least living species also have the larval head greatly reduced (Blagoderov et al. 2002).

<==Scatopsoidea BLM02
    |--Protoscatopsidae BLM02
    |--Synneuridae [Canthyloscelidae] BLM02
    |    |--Exiliscelis californiensis PC81
    |    `--Synneuron PC81
    |         |--S. annulipes PC81
    |         |--S. decipiens PC81
    |         `--S. silvestre PC81
    `--Scatopsidae C81
         |  i. s.: Procolobostema C81
         |         Parmaferia O98
         |--Ectaetia [Ectaetiinae] C81
         |    `--E. clavipes C81
         |--Aspistinae C81
         |    |--Aspistes harti C81
         |    `--Arthria analis C81
         |--Psectrosciarinae C81
         |    |--Anapausis soluta C81
         |    `--Psectrosciara A09
         |         |--P. californica C81
         |         `--P. serrata A09
         `--Scatopsinae C81
              |--Colobostematini A09
              |    |--Colobostema variatum C81
              |    |--Borneoscatopse A09
              |    |--Holoplagia guamensis A09
              |    |--Lumpuria A09
              |    |--Cookella albitarsis A09
              |    `--Ferneiella A09
              |--Scatopsini C81
              |    |--Reichertella C81
              |    |--Apiloscatopse uncinata C81
              |    |--Pharsoreichertella A09
              |    `--Scatopse C81
              |         |--S. fuscipes CM91
              |         |--S. nigra L02
              |         `--S. notata C81
              |--Rhegmoclematini C81
              |    |--Diamphidicus [Diamphidicina] A09
              |    |--Holoclema [Holoclematina] A09
              |    |--Rhegmoclematina A09
              |    |    |--Parmaferia A09
              |    |    `--Thripomorpha truncatum [=Rhegmoclema truncatum] A09
              |    `--Parascatopsina A09
              |         |--Parascatopse wirthi A09
              |         |--Neorhegmoclemina asymmetrica A09
              |         |--Rhegmoclemina bimaculata A09, C81
              |         `--Austroclemina A09
              `--Swammerdamellini C81
                   |--Swammerdamella obtusa C81
                   |--Akorhexoza cactivora A09
                   |--Coboldia fuscipes C81
                   |--Pararhexosa A09
                   |--Hawomersleya A09
                   |--Abrexoza A09
                   |--Cooka A09
                   |--Brahemyia A09
                   |--Quateiella A09
                   |    |--Q. inexpectata A09
                   |    `--Q. minuscula A09
                   `--Rhexoza C81
                        |--R. incisa C81
                        `--R. similis C81

*Type species of generic name indicated

REFERENCES

[A09] Amorim, D. de S. 2009. Scatopsidae (minute black scavenger flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 347–355. NRC Research Press: Ottawa.

[BLM02] Blagoderov, V. A., E. D. Lukashevich & M. B. Mostovski. 2002. Order Diptera Linné, 1758. The true flies (=Muscida Laicharting, 1781). In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 227–240. Kluwer Academic Publishers: Dordrecht.

[CM91] Colless, D. H., & D. K. McAlpine. 1991. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 717–786. Melbourne University Press: Carlton (Victoria).

[C81] Cook, E. F. 1981. Scatopsidae. In: McAlpine, J. F., B. V. Peterson, G. E. Shewell, H. J. Teskey, J. R. Vockeroth & D. S. Wood (eds) Manual of Nearctic Diptera vol. 1 pp. 313–319. Research Branch, Agriculture Canada.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[O98] Oosterbroek, P. 1998. The Families of Diptera of the Malay Archipelago. Brill: Leiden.

[PC81] Peterson, B. V., & E. F. Cook. 1981. Synneuridae. In: McAlpine, J. F., B. V. Peterson, G. E. Shewell, H. J. Teskey, J. R. Vockeroth & D. S. Wood (eds) Manual of Nearctic Diptera vol. 1 pp. 321–324. Research Branch, Agriculture Canada.

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