Heleomyzidae

Garlic fly Suillia sp., from here.

Belongs within: Schizophora.
Contains: Anthomyzidae.

The Heleomyzidae are a group of flies characterised by strong, convergent postocellar bristles, wing with a subcostal but no humeral break, and a preapical dorsal bristle on each tibia. The circumscription of the group has been disputed; as recognised below, it may be paraphyletic to the family Anthomyzidae (Wiegmann et al. 2011) and some authors have divided it between a number of families. Members of the tribe Rhinotorini have unusually tall heads with an excavated vertex and lacking postocellar bristles, and wings without spaced costal spines (McAlpine & Woodley 2010). The Trixoscelidini have ocellar bristles arising outside the ocellar triangle, alongside or in front of the anterior ocellus.

<==Heleomyzidae [Cnemospathidae, Helomyzidae]
    |--Trixoscelidini [Trixoscelidae, Trixoscelididae] MW10
    |    |--Spilochroa ornata MW10
    |    |--Zagonia flavicornis T87
    |    |--Paratrixoscelis T87
    |    |--Psiloplogia T87
    |    `--Trixoscelis WT11
    |         |--T. frontalis T87
    |         `--T. fumipennis MW10
    `--+--Anthomyzidae WT11
       `--Suillia [Suilliinae, Suilliini] WT11
            |--S. iniens MW10
            |--S. longipennis MW10
            |--S. lurida GP87
            |--S. punctulata MW10
            `--S. variegata WT11

Heleomyzidae incertae sedis:
  Chiropteromyza GE05
  Heleomyzini [Heleomyzinae] MW10
    |--Tephrochlamys rufiventris MW10, GP87
    |--Oecothea [incl. Aecothea, Neoecothea] MW10
    |    `--O. fenestralis CM70 [incl. Aecothea fidelis GP87]
    |--Scoliocentra [incl. Amoebaleria] MW10
    |    `--S. helvola [=Amoebaleria helvola] MW10
    |--Pseudoleria MW10
    |    |--P. pectinata CM70
    |    `--P. placata MW10 [incl. P. crassata GP87]
    `--Heleomyza GP87
         |--H. maculipennis [=Scoliocentra (Leriola) maculipennis; incl. H. difficilis] GP87
         `--H. serrata GP87
  Cinderella [Cinderellini] MW10
    `--C. lampra MW10
  Borboropsis [Borboropsidae] GP87
    `--B. puberula [incl. B. fulviceps] GP87
  Oldenbergiella brumalis GP87
  Lutomyia GP87
    |--L. hemiptera GP87
    `--L. spurca GP87
  Rhinotorini [Rhinotoridae] MW10
    |--Rhinotora MW10
    |--Rhinotoroides M87
    |--Anastomyza M87
    |--Apophoneura M87
    |--Zentula M87
    |--Neorhinotora MW10
    |    |--N. aristalis MW10
    |    `--N. diversa MW10
    `--Cairnsimyia M87
         |--C. aroana O98
         |--C. excavata M87
         `--C. robusta M87
  Paraneossos [Gephyromyzini] MW10
    `--P. arizonicus GP87
  Prosopantrum [Cnemospathidini] MW10
    `--P. flavifrons MW10
  Tapeigaster [Tapeigastrini] CM91
    `--T. annulipes CM91
  Allophylopsini CM91
    |--Diplogeomyza diaphora CM91
    `--Allophylopsis CM70
  Neossos T87
  Anorostoma GP87
    |--A. coloradense GP87
    `--A. currani O98
  Porsenus johnsoni GP87
  Allophyla atricornis [incl. A. laevis] GP87
  Morpholeria tristis GP87
  Schroederella GP87
    |--S. iners GP87
    `--S. luteoala GP87
  Orbellia GP87
  Eccoptomera GP87
  Neoleria GP87
  Spanoparea GP87
  Anypotacta GP87
  Trichochlamys borealis GP87
  Cephodapedon T87
  Chaetohelomyza electrica RJ93
  Electroleria P92
  Protosuillia P92

*Type species of generic name indicated

REFERENCES

[CM70] Colless, D. H., & D. K. McAlpine. 1970. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 656–740. Melbourne University Press.

[CM91] Colless, D. H., & D. K. McAlpine. 1991. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2^nd ed. vol. 2 pp. 717–786. Melbourne University Press: Carlton (Victoria).

[GP87] Gill, G. D., & B. V. Peterson. 1987. Heleomyzidae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 973–980. Research Branch, Agriculture Canada.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[M87] McAlpine, J. F. 1987. Rhinotoridae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 989–992. Research Branch, Agriculture Canada.

[MW10] McAlpine, D. K., & N. E. Woodley. 2010. Heleomyzidae (heleomyzid flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 1159–1164. NRC Research Press: Ottawa.

[O98] Oosterbroek, P. 1998. The Families of Diptera of the Malay Archipelago. Brill: Leiden.

[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.

[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.

[T87] Teskey, H. J. 1987. Trixoscelididae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 981–984. Research Branch, Agriculture Canada.

[WT11] Wiegmann, B. M., M. D. Trautwein, I. S. Winkler, N. B. Barr, J.-W. Kim, C. Lambkin, M. A. Bertone, B. K. Cassel, K. M. Bayless, A. M. Heimberg, B. M. Wheeler, K. J. Peterson, T. Pape, B. J. Sinclair, J. H. Skevington, V. Blagoderov, J. Caravas, S. N. Kutty, U. Schmidt-Ott, G. E. Kampmeier, F. C. Thompson, D. A. Grimaldi, A. T. Beckenbach, G. W. Courtney, M. Friedrich, R. Meier & D. K. Yeates. 2011. Episodic radiations in the fly tree of life. Proceedings of the National Academy of Sciences of the USA 108 (14): 5690–5695.

Last updated: 29 June 2021.