Heleomyzidae

Garlic fly Suillia sp., from here.


Belongs within: Schizophora.
Contains: Anthomyzidae.

The Heleomyzidae are a group of flies characterised by strong, convergent postocellar bristles, wing with a subcostal but no humeral break, and a preapical dorsal bristle on each tibia.

<==Heleomyzidae [Cnemospathidae] WT11
    |--Trixoscelidini [Trixoscelididae] MW10
    |    |--Spilochroa ornata MW10
    |    |--Zagonia flavicornis T87
    |    |--Paratrixoscelis T87
    |    |--Psiloplogia T87
    |    `--Trixoscelis WT11
    |         |--T. frontalis T87
    |         `--T. fumipennis M90
    `--+--Anthomyzidae WT11
       `--Suillia [Suilliinae, Suilliini] WT11
            |--S. iniens MW10
            |--S. longipennis M81
            |--S. lurida GP87
            |--S. punctulata MW10
            `--S. variegata WT11

Heleomyzidae incertae sedis:
  Chiropteromyza GE05
  Heleomyzini [Heleomyzinae] MW10
    |--Tephrochlamys rufiventris MW10, M90
    |--Oecothea [incl. Aecothea, Neoecothea] M10
    |    `--O. fenestralis CM70 [incl. Aecothea fidelis GP87]
    |--Scoliocentra [incl. Amoebaleria] MW10
    |    `--S. helvola [=Amoebaleria helvola] MW10
    |--Pseudoleria MW10
    |    |--P. pectinata CM70
    |    `--P. placata MW10 [incl. P. crassata GP87]
    `--Heleomyza M90
         |--H. maculipennis [=Scoliocentra (Leriola) maculipennis; incl. H. difficilis] GP87
         `--H. serrata M90
  Cinderella [Cinderellini] MW10
    `--C. lampra M90
  Borboropsis [Borboropsidae] M90
    `--B. puberula [incl. B. fulviceps] GP87
  Oldenbergiella brumalis GP87
  Lutomyia GP87
    |--L. hemiptera GP87
    `--L. spurca GP87
  Rhinotorini [Rhinotoridae] MW10
    |--Rhinotora MW10
    |--Rhinotoroides M87
    |--Anastomyza M87
    |--Apophoneura M87
    |--Zentula M87
    |--Neorhinotora MW10
    |    |--N. aristalis MW10
    |    `--N. diversa BW09
    `--Cairnsimyia M87
         |--C. excavata M87
         `--C. robusta M87
  Paraneossos [Gephyromyzini] MW10
    `--P. arizonicus GP87
  Prosopantrum [Cnemospathidini] MW10
    `--P. flavifrons MW10
  Tapeigaster [Tapeigastrini] CM91
    `--T. annulipes CM91
  Allophylopsini CM91
    |--Diplogeomyza diaphora CM91
    `--Allophylopsis CM70
  Neossos GP87
  Anorostoma coloradense GP87
  Porsenus johnsoni GP87
  Allophyla atricornis [incl. A. laevis] GP87
  Morpholeria tristis GP87
  Schroederella GP87
    |--S. iners GP87
    `--S. luteoala GP87
  Orbellia GP87
  Eccoptomera GP87
  Neoleria GP87
  Spanoparea GP87
  Anypotacta GP87
  Trichochlamys borealis GP87
  Cephodapedon T87

*Type species of generic name indicated

REFERENCES

[BW09] Buck, M., N. E. Woodley, A. Borkent, D. M. Wood, T. Pape, J. R. Vockeroth, V. Michelsen & S. A. Marshall. 2009. Key to Diptera families—adults. In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 95–156. NRC Research Press: Ottawa.

[CM70] Colless, D. H., & D. K. McAlpine. 1970. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 656–740. Melbourne University Press.

[CM91] Colless, D. H., & D. K. McAlpine. 1991. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 717–786. Melbourne University Press: Carlton (Victoria).

[GP87] Gill, G. D., & B. V. Peterson. 1987. Heleomyzidae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 973–980. Research Branch, Agriculture Canada.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[M81] McAlpine, J. F. 1981. Key to families—adults. In: McAlpine, J. F., B. V. Peterson, G. E. Shewell, H. J. Teskey, J. R. Vockeroth & D. S. Wood (eds) Manual of Nearctic Diptera vol. 1 pp. 89–124. Research Branch, Agriculture Canada.

[M87] McAlpine, J. F. 1987. Rhinotoridae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 989–992. Research Branch, Agriculture Canada.

[M90] McAlpine, J. F. 1990. Insecta: Diptera adults. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1211–1252. John Wiley & Sones: New York.

[MW10] McAlpine, D. K., & N. E. Woodley. 2010. Heleomyzidae (heleomyzid flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 1159–1164. NRC Research Press: Ottawa.

[T87] Teskey, H. J. 1987. Trixoscelididae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 981–984. Research Branch, Agriculture Canada.

[WT11] Wiegmann, B. M., M. D. Trautwein, I. S. Winkler, N. B. Barr, J.-W. Kim, C. Lambkin, M. A. Bertone, B. K. Cassel, K. M. Bayless, A. M. Heimberg, B. M. Wheeler, K. J. Peterson, T. Pape, B. J. Sinclair, J. H. Skevington, V. Blagoderov, J. Caravas, S. N. Kutty, U. Schmidt-Ott, G. E. Kampmeier, F. C. Thompson, D. A. Grimaldi, A. T. Beckenbach, G. W. Courtney, M. Friedrich, R. Meier & D. K. Yeates. 2011. Episodic radiations in the fly tree of life. Proceedings of the National Academy of Sciences of the USA 108 (14): 5690–5695.

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