Gathering of Epinannolene, copyright Karl Questel.

Belongs within: Helminthomorpha.
Contains: Chordeumatida, Polydesmida, Spirobolida, Julida, Spirostreptidae.

The Eugnatha are a clade within the millipedes characterised by modification of the eighth pair of legs in adult males into a pair of functional gonopods, and the fusion of the pleurites to the tergites. Major clades within the Eugnatha include the Nematophora, characterised by the possession of spinnerets, and molar cusps on the mandibles (Sierwald et al. 2003).

    |--Nematophora SS03
    |    |--Stemmiulida [Monocheta] WA04
    |    `--Coelocheta SS03
    |         |--Chordeumatida WA04
    |         `--Callipodida WA04
    |              |--Abacion magnum H00
    |              |--Delophon H90
    |              |--Tetracion H90
    |              |--Texophon H90
    |              |--Tynomma H90
    |              |--Diactis H90
    |              |--Heptium H90
    |              |--Colactis H90
    |              `--Apfelbeckia Verhoeff 1896 H00
    |                   `--A. hessei Verhoeff 1929 UO05
    `--+--Polydesmida WA04
       `--Juliformia [Juloidea] SS03
            |--+--Spirobolida SS03
            |  `--Julida SS03
            `--Spirostreptida SS03
                 |--Spirostreptidae H90
                 |--Epinannolenidea H90
                 |    |--Choctella [Choctellidae] H90
                 |    |    `--C. cummingsi H90
                 |    `--Epinannolene K55
                 |         |--E. alticola (Silvestri 1898) K55
                 |         |--E. bifasciata (Silvestri 1898) K54
                 |         |--E. cylidricaulis Kraus 1954 K54
                 |         `--E. flagellosa Kraus 1955 K55
                 `--Cambalidea H00
                      |  i. s.: Protosilvestria Handschin 1944 H00
                      |--Cambalopsidae H90
                      |    |--Plusioglyphiulus SGS04
                      |    `--Cambalopsis nordquisti A27
                      `--Cambalidae [Leioderidae, Nannolenidae] H90
                           |--Cambala hubrichti H00
                           |--Agastrophus B13
                           |--Hypocambala B13
                           |--Nannolene H90
                           |--Paiteya H90
                           |--Titsona H90
                           |    |--*T. sima C62
                           |    `--T. tida Chamberlin 1962 C62
                           |--Buwatia H90
                           |--Platydere H90
                           |--Pharodere H90
                           |--Endere H90
                           |--Leiodere H90
                           |--Odachurus H90
                           |--Alakene H90
                           |--Doilene H90
                           |--Tigolene H90
                           |--Amastigogonus Brölemann 1913 B13
                           |    `--*A. tasmanianus Brölemann 1913 B13
                           `--Dimerogonus Attems 1903 B13
                                `--*D. orophilus B13

*Type species of generic name indicated


[A27] Attems, C. 1927. Myriopoden von Ambon und anderen Inseln der Banda-See. Zoologische Mededeelingen 10: 61–70.

[B13] Brölemann, H. W. 1913. The Myriapoda in the Australian Museum. Part ii. Diplopoda. Records of the Australian Museum 10 (6): 77–158, pls 14–18.

[C62] Chamberlin, R. V. 1962. Millipeds from the Nevada Test Area. Proceedings of the Biological Society of Washington 75: 53–56.

[H00] Hannibal, J. T. 2000. Hexecontasoma, a new helminthomorph millipede (Hexecontasomatidae n. fam.) from the Mazon Creek, Illinois, fauna (Carboniferous, North America). Fragmenta Faunistica 43 (Suppl): 19–35.

[H90] Hoffman, R. L. 1990. Diplopoda. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 835–860. John Wiley & Sones: New York.

[K54] Kraus, O. 1954. Myriapoden aus Peru, II. Senckenbergiana Biologica 35: 17–55.

[K55] Kraus, O. 1955. Myriapoden aus Peru, III. Senckenbergiana Biologica 36 (3–4): 173–200.

[SGS04] Schwendinger, P. J., G. Giribet & H. Steiner. 2004. A remarkable new cave-dwelling Stylocellus (Opiliones, Cyphophthalmi) from peninsular Malaysia, with a discussion on taxonomic characters in the family Stylocellidae. Journal of Natural History 38: 1421–1435.

[SS03] Sierwald, P., W. A. Shear, R. M. Shelley & J. E. Bond. 2003. Millipede phylogeny revisited in the light of the enigmatic order Siphoniulida. Journal of Zoological Systematics and Evolutionary Research 41: 87–99.

[UO05] Ubick, D., & R. Ozimec. 2005. On the harvestman genus Lola Kratochvíl (Opiliones: Laniatores). Natura Croatica 14 (3): 161–174.

[WA04] Wilson, H. M., & L. I. Anderson. 2004. Morphology and taxonomy of Paleozoic millipedes (Diplopoda: Chilognatha: Archipolypoda) from Scotland. Journal of Paleontology 78 (1): 169–184.

Last updated: 25 May 2017.

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