?Epicyliosoma sp., copyright Imbala.

Belongs within: Diplopoda.

The Pentazonia is a clade including the pill millipedes and related taxa. Members of this clade are united by the presence of telopods and divided sternites (Sierwald et al 2003).

<==Pentazonia [Opisthandria]
    |--Glomeridesmida [Limacomorpha] SS03
    `--Oniscomorpha SS03
         |--Glomerida [Plesiocerata] RHV04
         |    |  i. s.: Corsikomeris remyi Verhoeff 1943 RHV04
         |    `--Glomeridae H90
         |         |--Glomeris L89
         |         |    |--G. connexa L89
         |         |    |--G. inferorum Latzel 1889 L89
         |         |    |--G. marginata KAA-W00
         |         |    `--G. pustulata Latreille 1804 TKM93
         |         |--Onomeris underwoodi H90
         |         `--Glomeroides H90
         |--Amynilyspedida RHV04
         |    |  i. s.: Paraglomeris carbonaria Schlechtendal 1912 RHV04
         |    |--+--Glomeropsis Fritsch 1899 RHV04
         |    |  |--Archiscudderia Fritsch 1899 RHV04
         |    |  `--Palaeosphaeridium Peach 1914 RHV04
         |    `--Amynilyspes Scudder 1882 [Acroglomeridae, Amynilyspedidae] RHV04
         |         |--*A. wortheni Scudder 1882 RHV04
         |         |--A. crescens Fritsch 1899 RHV04
         |         |--A. fatimae Racheboeuf, Hannibal & Vannier 2004 RHV04
         |         `--A. typicus Fritsch 1899 RHV04
         `--Sphaerotheriida RHV04
              |--Sphaeropoeus [Sphaeropoeidae] B13
              `--Sphaerotheriidae MH08
                   |--Cycliosomella furcipatum [=Sechelliosoma furcipatum] H02
                   |--Arthrosphaera brandti Humbert 1865 RHV04
                   |--Procyliosoma Silvestri 1917 MH08
                   |--Cynotelopus Jeekel 1986 MH08
                   |    `--C. notabilis Jeekel 1986 MH08
                   |--Sphaerotherium MH08
                   |    |--S. convexum Koch 1847 MH08
                   |    `--S. libidinosum B13
                   |--Epicyliosoma Silvestri 1917 MH08
                   |    |  i. s.: E. penrithense (Brölemann 1913) MH08
                   |    |         E. sennae (Silvestri 1898) MH08
                   |    |--E. (Epicyliosoma) MH08
                   |    |    |--*E. (E.) albertisii (Silvestri 1895) [=Zephronia albertisii] MH08
                   |    |    `--E. (E.) sarahae Moir & Harvey 2008 MH08
                   |    `--E. (Paracyliosoma Verhoeff 1928) MH08
                   `--Cyliosoma Pocock 1895 B13
                        |--*C. angulatum (Butler 1878) [=Sphaerotherium angulatum] B13
                        |--C. delacyi (White 1859) [=Zephronia delacyi] B13
                        |--C. leiosomum [=Sphaerotherium leiosomum] B13
                        |--C. penrithensis Brölemann 1913 B13
                        |--C. queenslandiae Brölemann 1913 B13
                        |--C. sennae Silvestri 1898 B13
                        |--C. striolatum Pocock 1895 B13
                        |--C. targionii Silvestri 1898 B13
                        `--C. unicolor Silvestri 1897 B13

*Type species of generic name indicated


[B13] Brölemann, H. W. 1913. The Myriapoda in the Australian Museum. Part ii. Diplopoda. Records of the Australian Museum 10 (6): 77-158, plates XIV-XVIII.

[H90] Hoffman, R. L. 1990. Diplopoda. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 835-860. John Wiley & Sones: New York.

[H02] Hunt, D. J. 2002. The African Rhigonematoidea (Nematoda: Rhigonematida). 1. Trachyglossus quintus gen. n., sp. n. (Xustrostomatidae fam. n.) with redescription of Rhigonema multipapillatum (Skrjabin, 1916) and designation of a neotype. Nematology 4: 671-686.

[KAA-W00] Kheirallah, A.-M., A.-N. H. Aly & N. Y. Abdel-Wahed. 2000. Anamorphosis and life-history of the millipede Nopoiulus kochii (Gervais, 1847), new for Egypt. Zoology in the Middle East 21: 159-168.

[L89] Latzel, R. 1889. Sopra alenni miriapodi cavernicoli italiani, raccolti dai Sign. A. Vacca e R. Barberi. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a 7: 360-362.

[MH08] Moir, M. L., & M. S. Harvey. 2008. Discovery of the pill millipede genus Epicyliosoma (Diplopoda: Sphaerotheriida: Sphaerotheriidae) in Western Australia, with the description of a new species. Records of the Western Australian Museum 24 (2): 113-119.

[RHV04] Racheboeuf, P. R., J. T. Hannibal & J. Vannier. 2004. A new species of the diplopod Amynilyspes (Oniscomorpha) from the Stephanian Lagerstätte of Montceau-les-Mines, France. Journal of Paleontology 78 (1): 221-229.

[SS03] Sierwald, P., W. A. Shear, R. M. Shelley & J. E. Bond. 2003. Millipede phylogeny revisited in the light of the enigmatic order Siphoniulida. Journal of Zoological Systematics and Evolutionary Research 41: 87-99.

[TKM93] Thaler, K., B. Knoflach & E. Meyer. 1993. Fragmenta faunistica Tirolensia—X. Berichte des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck 80: 311-325.

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