Geophilomorpha

Geophilus insculptus, copyright Trevor & Dilys Pendleton.


Belongs within: Myriapoda.
Contains: Mecistocephalinae.

The Geophilomorpha are a group of slender, extremely elongate centipedes that live in burrows or crevices.

Characters (from Mundel 1990): Eyes absent; antennae with 14 segments. Number of pedal segments varying from 31 to over 180.

<==Geophilomorpha
    |--Adesmata BFM03
    |    |  i. s.: Stigmatogaster gracilis Meinert 1870 BFM03
    |    |--Geophilidae BFM03
    |    |    |--Pachymerium [Pachymeriidae] BFM03
    |    |    |    `--‘Geophilus’ attenuatus Say 1821 [incl. G. ferrugineus Koch 1835] BFM03
    |    |    `--Geophilus BFM03
    |    |         |--G. flavidus C01
    |    |         |    |--G. f. flavidus A01
    |    |         |    `--G. f. escherichii Verh. 1896 A01
    |    |         |--G. ganonotus Attems 1901 A01
    |    |         |--G. infossulatus Attems 1901 A01
    |    |         |--G. insculptus Attems 1895 BFM03
    |    |         `--G. sibiricus A01
    |    `--Chilenophilidae [Chilenophilinae] K54
    |         |--Ribautia GEW01
    |         |--Lestophilus K54
    |         |--Nesidiphilus Chamberlin 1915 [incl. Suturodes Chamberlin 1922] K54
    |         |    |--N. montecristi Kraus 1954 K54
    |         |    `--N. tardus (Chamberlin 1922) K54
    |         `--Polycricus Saussure & Humbert 1872 K54
    |              |--P. aequatorialis Silvestri 1902 K54
    |              `--P. toltecus (Humbert & Saussure 1869) (see below for synonymy) K54
    `--Mecistocephalidae [Dicellophilidae] BFM03
         |--Arrupinae BFM03
         |    |  i. s.: Partygarrupius Verhoeff 1939 BFM03
         |    |--Agnostrup Foddai, Bonato et al. 2003 BFM03
         |    |    `--A. paucipes (Miyosi 1955) BFM03
         |    `--+--Nannarrup Foddai, Bonato et al. 2003 BFM03
         |       |    `--*N. hoffmani Foddai, Bonato et al. 2003 BFM03
         |       `--Arrup Chamberlin 1912 [incl. Nodocephalus Attems 1928, Prolamnonyx Silvestri 1919] BFM03
         |           |--A. holstii (Pocock 1895) BFM03
         |           `--+--*A. pylorus Chamberlin 1912 BFM03
         |              `--A. dentatus (Takakuwa 1934) BFM03
         `--+--Mecistocephalinae BFM03
            `--Dicellophilinae BFM03
                 |--Proterotaiwanella Bonato, Foddai & Minelli 2002 BFM03
                 |    |--*P. tanabei Bonato, Foddai & Minelli 2002 BFM03
                 |    `--P. sculptulata (Takakuwa 1936) BFM03
                 `--+--Anarrup Chamberlin 1920 [incl. Sundarrup Attems 1930] BFM03
                    |    |--*A. nesiotes Chamberlin 1920 BFM03
                    |    `--A. flavipes (Attems 1930) [=*Sundarrup flavipes] BFM03
                    `--Dicellophilus Cook 1896 BFM03
                         |--D. carniolensis (Koch 1847) BFM03
                         `--+--D. latifrons Takakuwa 1934 BFM03
                            `--+--*D. limatus (Wood 1862) BFM03
                               `--D. anomalus (Chamberlin 1904) BFM03

Geophilidae incertae sedis:
  Oryidae K55
    |--Orphnaeus brevilabiatus (Newport 1845) [=Geophilus brevilabiatus] K54
    `--Notiphilides maximilani (Humbert & Saussure 1870) (see below for synonymy) K55
  Schendylurus [Schendylidae, Schendylinae] K55
    `--S. pallidus Kraus 1955 K55
  Strigamia [Linotaeniidae] BFM03
  Zelanion antipodus MG06
  Soniphilidae M90
  Sogonidae M90
  Tampiyidae M90
  Aphilodontidae M90
  Eriphantidae M90
  Dignathodontidae M90
  Himantaridae M90
  Ballophilidae M90
  Neogeophilidae M90
  Gonibregmatidae M90

Notiphilides maximilani (Humbert & Saussure 1870) [=Notiphilus maximilani; incl. Keporya miranda Chamberlin 1941, Notiphilides maximilani miranda] K55

Polycricus toltecus (Humbert & Saussure 1869) [=Arthronomalus toltecus, A. (Polycricus) toltecus, Geophilus toltecus] K54

*Type species of generic name indicated

REFERENCES

[A01] Attems, C. Graf. 1901. Myriopodák [Myriopoden]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy], Band II. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy], pp. 275-310. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[BFM03] Bonato, L., D. Foddai & A. Minelli. 2003. Evolutionary trends and patterns in centipede segment number based on a cladistic analysis of Mecistocephalidae (Chilopoda: Geophilomorpha). Systematic Entomology 28: 539-579.

[C01] Csiki, E. 1901. Utivázlat [Reiseskizze]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy], Band II. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy], pp. xii-xli. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[GEW01] Giribet, G., G. D. Edgecombe & W. C. Wheeler. 2001. Arthropod phylogeny based on eight molecular loci and morphology. Nature 413: 157-161.

[K54] Kraus, O. 1954. Myriapoden aus El Salvador. Senckenbergiana Biologica 35 (5-6): 293-349.

[K55] Kraus, O. 1955. Myriapoden aus Peru, III. Senckenbergiana Biologica 36 (3-4): 173-200.

[MG06] Mallatt, J., & G. Giribet. 2006. Further use of nearly complete 28S and 18S rRNA genes to classify Ecdysozoa: 37 more arthropods and a kinorhynch. Molecular Phylogenetics and Evolution 40: 772-794.

[M90] Mundel, P. 1990. Chilopoda. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 819-833. John Wiley & Sones: New York.

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