Valves of Rabienella materni, from Wang (1987).

Belongs within: Myodocopida.

The Entomozoidae are a group of fossils known from the Ordovician to the Permian, usually classified as ostracods though this has been questioned on occasion. The carapace bears a long, deep nuchal furrow in most species assigned to the subfamily Entomozoinae but this furrow is slight or absent in most of the Devonian Richterininae. Boucia ornatissima, known from the Upper Silurian of Bohemia, bears a posteroventral sulcus in addition to the nuchal furrow (Benson et al. 1961).

Characters (from Benson et al. 1961): Large; surface often striate; without rostrum, rostral sinus or anterodorsal swelling; often with long, deep nuchal furrow (sometimes slight or absent).

Entomozoidae [Entomidae] W87
    |--Boucia Agnew 1942 [=Basslerella Bouček 1936 nec Kellett 1935 nec Howe 1935; Bouciinae] BB61
    |    `--*B. ornatissima (Bouček 1936) [=*Basslerella ornatissima] BB61
    |--Richterininae W87
    |    |--Fossirichterina Matern 1929 BB61
    |    |    `--*F. intercostata (Matern 1929) [=Richterina (*Fossirichterina) intercostata] BB61
    |    |--Maternella Rabien 1954 BB61
    |    |    |--*M. dichotoma (Paekelmann 1913) [=Richterina costata var. dichotoma] BB61
    |    |    `--M. hemisphaerica H79
    |    `--Richterina Gürich 1896 W87
    |         |  i. s.: R. viltata BB61
    |         |--*R. (Richterina) costata (Richter 1869) [=Cytherina costata] BB61
    |         `--R. (Volkina Rabien 1954) W87
    |              `--R. (*V.) zimmermanni (Volk 1939) W87, BB61 [=Entomis (Nehdentomis) zimmermanni BB61]
    `--Entomozoinae W87
         |--Pseudoentomozoe Přibyl 1951 BB61
         |    `--*P. pteroides (Canavari 1899) [=Entomis pteroides] BB61
         |--Vltavina Bouček 1936 BB61
         |    `--*V. bohemica Bouček 1936 BB61
         |--Bertillonella Stewart & Hendrix 1945 [incl. Waldeckella Rabien 1954] W87
         |    |--*B. subcircularis Stewart & Hendrix 1945 BB61
         |    |--B. cicatricosa (Matern 1929) (see below for synonymy) W87
         |    `--B. erecta (Rabien 1954) W87 [=*Waldeckella erecta BB61]
         |--Entomoprimitia Kummerow 1939 W87 [=Entoprimitia (l. c.) M61]
         |    |--E. nitida (Roemer 1850) (see below for synonymy) BB61
         |    |--E. concentrica (Matern 1929) [=Haploprimitia concentrica] W87
         |    |--E. kayseri (Waldschmidt 1885) [=Cypridina kayseri, Haploprimitia kayseri] W87
         |    |--E. paeckelmanni W87
         |    |--E. splendens BB61
         |    `--E. variostriata (Clarke 1884) [=Primitia variostriata] W87
         |--Rabienella Gründel 1962 W87
         |    |  i. s.: R. barrandei W87
         |    |--R. materni (Volk 1939) (see below for synonymy) W87
         |    `--+--R. volki (Rabien 1958) [=Waldeckella volki, Bertillonella (Rabienella) volki] W87
         |       `--+--R. schmidti (Rabien 1958) [=Waldeckella schmidti, Bertillonella (Rabienella) schmidti] W87
         |          `--+--*R. reichi (Matern 1929) (see below for synonymy) W87
         |             `--+--R. intermedia (Matern 1929) (see below for synonymy) W87
         |                `--R. kegeli (Matern 1929) [=Primitiella kegeli, Waldeckella (Rabienella) kegeli] W87
         `--Entomozoe Přibyl 1951 [=Entomis Jones 1861 non Herrich-Schaefer 1856] BB61
              |  i. s.: E. meneghinii BB61
              |         E. nehdensis H79
              |         E. variostriata H79
              |         E. zoppii BB61
              |--*E. (Entomozoe) tuberosa (Jones 1861) [=*Entomis tuberosa] BB61
              |--E. (Nandania) W87
              |    |--E. (N.) asymmetrica (Koch 1967) W87
              |    `--E. (N.) pseudophthalmus W87
              `--E. (Nehdentomis Matern 1929) W87, BB61
                   |--‘Entomis’ (*N.) nehdensis Matern 1929 BB61
                   |--E. (N.) pseudorichterina (Matern 1929) W87
                   `--E. (N.) tenera W87

Bertillonella cicatricosa (Matern 1929) [=Primitiella cicatricosa, B. (Rabienella) cicatricosa, Waldeckella (R.) cicatricosa] W87

Entomoprimitia nitida (Roemer 1850) [=Cypridina nitida; incl. Primitia hattingensis Matern 1929, *Entomoprimitia hattingensis] BB61

Rabienella intermedia (Matern 1929) [=Primitiella intermedia, Waldeckella (Rabienella) intermedia] W87

Rabienella materni (Volk 1939) [=Primitiella materni, Bertillonella (Rabienella) materni, Waldeckella (R.) materni] W87

*Rabienella reichi (Matern 1929) [=Primitiella reichi, Bertillonella (Rabienella) reichi, Waldeckella (R.) reichi] W87

*Type species of generic name indicated


[BB61] Benson, R. H., J. M. Berdan, W. A. van den Bold, T. Hanai, I. Hessland, H. V. Howe, R. V. Kesling, S. A. Levinson, R. A. Reyment, R. C. Moore, H. W. Scott, R. H. Shaver, I. G. Sohn, L. E. Stover, F. M. Swain & P. C. Sylvester-Bradley. 1961. Systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt Q. Arthropoda 3: Crustacea: Ostracoda pp. Q99–Q421. Geological Society of America and University of Kansas Press.

[H79] House, M. R. 1979. Devonian in the Eastern Hemisphere. In: Robison, R. A., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt A. Introduction. Fossilisation (Taphonomy), Biogeography and Biostratigraphy pp. A183–A217. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

[M61] Moore, R. C. 1961. Supplement—USSR treatise on Ostracoda. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt Q. Arthropoda 3: Crustacea: Ostracoda pp. Q422–Q429. Geological Society of America and University of Kansas Press.

[W87] Wang S.-Q. 1987. Entomozoids (Ostracoda) from Upper Devonian at Hangshan section of Longlin, Guangxi. Acta Palaeontologica Sinica 26 (3): 306–318.

Last updated: 20 October 2020.

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