Caconemobius fori, copyright Alan Cressler.

Belongs within: Grylloidea.

The Nemobiinae, pygmy field crickets, are small ground-dwelling crickets found in moist habitats (Rentz 1996). Members of the tribe Thetellini are flightless and usually associated with coastal habitats such as rocky or coral beaches or mangroves (Otte 1994). Members of the tribe Pteronemobiini, found worldwide, are characterised by the presence of a glandular spur on the hind tibiae in males. The wood cricket Nemobius sylvestris is a species with reduced wings found in forest margins in Europe and northern Africa.

Characters (from Rentz 1996): With or without tegmina. Head with dorsal bristles. Hind tibia shorter than hind femur, with 2-4 internal and external subapical spurs, without spines between spurs, and with three internal and three external apical spurs. Second tarsal segment laterally compressed, without large pads.

<==Nemobiinae R96
    |--Pteronemobius R96 [Pteronemobiini O94]
    |    |--P. bigelowi M83
    |    |--P. heydeni D37
    |    |--P. nigrovus M83
    |    |--P. nundra R96
    |    |--P. regulus R96
    |    `--P. truncatus R96
    |--Nemobius BM76 [Nemobiini O94]
    |    |  i. s.: N. allardi BM76
    |    |         N. csikii Bolivar 1901 B01
    |    |         N. fasciatus BM76
    |    |           |--N. f. fasciatus RH15
    |    |           `--N. f. socius RH15
    |    |         N. heydeni B01
    |    |         N. infernalis B01
    |    |         N. nitidus Bolivar 1901 B01
    |    |         N. sylvestris I92
    |    `--N. (Argizala) R15
    |         |--N. (A.) brasiliensis (Walker 1869) R15, R18 [=Argizala brasiliensis R18]
    |         `--N. (A.) hebardi Rehn 1915 R15 [incl. N. argentinus Bruner 1916 R18]
    `--Thetellini [Burcini] O94
         |--Thetella tarnis Otte & Alexander 1983 [incl. Burcus koblovae] O94
         |--Apteronemobius O94
         |--Fikola O94
         |--Ionemobius O94
         |--Orintia O94
         |--Parapteronemobius O94
         `--Caconemobius Kirby 1906 [=Paranemobius Alfken 1901 non Saussure 1877] O94
              |--*C. schauinslandi (Alfken 1901) [=*Paranemobius schauinslandi] O94
              |--C. albus Otte 1994 O94
              |--C. anahulu Otte 1994 O94
              |--C. fori Gurney & Rentz 1978 O94
              |--C. howarthi Gurney & Rentz 1978 O94
              |--C. nihoensis Otte 1994 O94
              |--C. paralbus Otte 1994 O94
              |--C. sandwichensis Otte 1994 O94
              |--C. uuku Otte 1994 O94
              `--C. varius Gurney & Rentz 1978 O94

Nemobiinae incertae sedis:
  Bobilla R96
    |--B. neobivittata R91
    `--B. victoriae R96
  Narellina R96
  Tincanita R96
  Silvinella R96
  Nambungia R96
  Specnia R96
  Dictyonemobius R96
    |--D. lateralis R96
    `--D. pacificus R96

*Type species of generic name indicated


[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[B01] Bolivar, I. 1901. Orthopterák [Orthoptères]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 223–243. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[D37] Dobzhansky, T. 1937. Genetics and the Origin of Species. Columbia University Press: New York.

[I92] Imes, R. 1992. The Practical Entomologist. Aurum Press: London.

[M83] Martin, N. A. 1983. Miscellaneous observations on a pasture fauna: an annotated species list. DSIR Entomology Division Report 3: 1–98.

[O94] Otte, D. 1994. The Crickets of Hawaii: origin, systematics and evolution. The Orthopterists' Society: The Academy of Natural Sciences of Philadelphia.

[R15] Rehn, J. A. G. 1915. A further contribution to the knowledge of the Orthoptera of Argentina. Proceedings of the Academy of Natural Sciences of Philadelphia 67 (2): 270–292.

[R18] Rehn, J. A. G. 1918. On a collection of Orthoptera from the State of Pará, Brazil. Proceedings of the Academy of Natural Sciences of Philadelphia 70: 144–236, pls 1–2.

[RH15] Rehn, J. A. G., & M. Hebard. 1915. The genus Gryllus (Orthoptera) as found in America. Proceedings of the Academy of Natural Sciences of Philadelphia 67 (2): 293–322.

[R91] Rentz, D. C. F. 1991. Orthoptera (grasshoppers, locusts, katydids, crickets). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1 pp. 369–393. Melbourne University Press: Carlton (Victoria).

[R96] Rentz, D. 1996. Grasshopper Country: The abundant orthopteroid insects of Australia. University of New South Wales Press: Sydney.

Last updated: 23 May 2022.

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