Myrtales

Pau-terra Qualea grandiflora, from here.


Belongs within: Malvidae.
Contains: Combretaceae, Onagraceae, Lythraceae, Melastomatineae, Myrtaceae.

The Myrtales are a clade of flowering plants, most members of which have flaky bark and opposite leaves with colleters (Angiosperm Phylogeny Website).

Synapomorphies (from Angiosperm Phylogeny Website): Bark flaky; flavonols only, myricetin, methylated ellagic acid present; cork cambium deep seated; pits vestured; libriform fibres septate; tension wood present; secondary phloem stratified; vascular bundles bicollateral, internal phloem present; nodes 1:1; cuticle waxes often absent; branching from current flush; leaves opposite, lamina with brochidodromous venation, colleters present; inflorescence racemose; hypanthium present, nectariferous; calyx valvate, corolla clawed; anthers incurved in bud; pollen with pseudocolpi; gynoecium inferior, style long, minor stylar bundles present, stigma wet; ovules many/carpel, micropyle bistomal and zig-zag, inner integument ca 2 cells across; antipodal cells ephemeral; exotegmen cells tracheidal, endotesta crystalliferous; endosperm at most slight.

<==Myrtales [Myrtanae, Myrtiflorae, Sonneratiaceae]
    |  i. s.: Xenodendron YY22
    |         Paleomyrtinaea WM09
    |--Lythrineae [Onagrineae] T00
    |    |--Combretaceae WM09
    |    `--+--Onagraceae WM09
    |       `--Lythraceae WM09
    `--+--Melastomatineae WM09
       `--Myrtineae T00
            |--Myrtaceae WM09
            `--Qualea [Vochysiaceae] WM09
                 |--Q. grandiflora WO05
                 `--Q. parviflora WM09

*Type species of generic name indicated

REFERENCES

[T00] Thorne, R. F. 2000. The classification and geography of the flowering plants: Dicotyledons of the class Angiospermae (subclasses Magnoliidae, Ranunculidae, Caryophyllidae, Dilleniidae, Rosidae, Asteridae, and Lamiidae). The Botanical Review 66: 441-647.

[WM09] Wang, H., M. J. Moore, P. S. Soltis, C. D. Bell, S. F. Brockington, R. Alexandre, C. C. Davis, M. Latvis, S. R. Manchester & D. E. Soltis. 2009. Rosid radiation and the rapid rise of angiosperm-dominated forests. Proceedings of the National Academy of Sciences of the USA 106 (10): 3853-3858.

[WO05] Wilson, P. G., M. M. O'Brien, M. M. Heslewood & C. J. Quinn. 2005. Relationships within Myrtaceae sensu lato based on a matK phylogeny. Plant Systematics and Evolution 251: 3-19.

[YY22] Yampolsky, C., & H. Yampolsky. 1922. Distribution of sex forms in the phanerogamic flora. Bibliotheca Genetica 3: 1-62.

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