Water apple Eugenia aquea, photographed by Tai Lung Aik.

Belongs within: Myrteae.

Eugenia is a genus of trees and shrubs that can be distinguished from closely related taxa by its pinnately veined leaves, single-seeded fruit and massive cotyledons (Flora of China). The centre of diversity for the genus is in the Americas, but species are found throughout the tropics. The fruit of some species, such as the water apple or jambu air Eugenia aquea, is edible.

Characters (from Flora of China): Trees or shrubs. Leaves opposite, petiolate; leaf blade pinnately veined. Inflorescences axillary or often lateral below leaves. Flowers bisexual, solitary or clustered. Hypanthium short. Calyx lobes 4. Petals 4. Stamens numerous; anthers parallel, longitudinally dehiscent. Ovary 2- or 3-loculed; ovules many per locule, amphitropous. Fruit a berry, with persistent sepals at apex. Seed usually 1, embryo straight, with 2 fully or partly fused massive cotyledons.

    |--E. aquea [=Syzygium aqueum] P88
    |--E. aromatica ZB01
    |--E. braziliensis MM96
    |--E. buxifolia G17
    |--E. caryophyllus P88
    |--E. christii J87
    |--E. firma S02
    |--E. foetida J87
    |--E. formonica J87
    |--E. glabrata J87
    |--E. grandis P88
    |--E. lineata J87
    |--E. maire A27
    |--E. malaccensis P88 [=Jambosa malaccensis C55, Syzygium malaccense P88]
    |--E. picardae J87
    |--E. smithii B88
    |--E. stictosepala MP00
    |--E. tiburona J87
    `--E. uniflora WO05

*Type species of generic name indicated


[A27] Andersen, J. C. 1927. Popular names of New Zealand plants. Transactions and Proceedings of the New Zealand Institute 57: 905-977.

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[C55] Candolle, A. de. 1855. Géographie Botanique Raisonée: Ou exposition des faits principaux et des lois concernant la distribution géographique des plantes de l’époque actuelle vol. 2. Librairie de Victor Masson: Paris.

[G17] Girault, A. A. 1917. New chalcid flies. Privately published (reprinted Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 72-77).

[J87] Judd, W. S. 1987. Floristic study of Morne La Visite and Pic Macaya National Parks, Haiti. Bulletin of the Florida State Museum—Biological Sciences 32 (1): 1-136.

[MP00] Machado, G., & M. A. Pizo. 2000. The use of fruits by the neotropical harvestman Neosadocus variabilis (Opiliones, Laniatores, Gonyleptidae). Journal of Arachnology 28: 357-360.

[MM96] Mound, L. A., & R. Marullo. 1996. The thrips of Central and South America: an introduction (Insecta: Thysanoptera). Memoirs on Entomology, International 6: 1-487.

[P88] Polunin, I. 1988. Plants and Flowers of Malaysia. Times Editions: Singapore.

[S02] Smetacek, P. 2002. Notes on new records of hooktip moths, Lepidoptera: Drepanidae, from the Kumaon and Garhwal Himalaya. Journal of the Bombay Natural History Society 99 (3): 446-454.

[WO05] Wilson, P. G., M. M. O'Brien, M. M. Heslewood & C. J. Quinn. 2005. Relationships within Myrtaceae sensu lato based on a matK phylogeny. Plant Systematics and Evolution 251: 3-19.

[ZB01] Zhang, N., & M. Blackwell. 2001. Molecular phylogeny of dogwood anthracnose fungus (Discula destructiva) and the Diaporthales. Mycologia 93 (2): 355-365.

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