Saccamminidae

Pelosina variabilis, copyright Jan Pawlowski.

Belongs within: Foraminifera.

The Saccamminidae are a group of Foraminifera with agglutinated tests known from the Ordovician to the present (Loeblich & Tappan 1964); it is likely that this represents a polyphyletic assemblage of relatively simple-bodied forams.

See also: A small bag of grains (Taxon of the Week: Saccamminidae); Three random foram genera (Taxon of the Week: Pelosininae).

Characters (from Loeblich & Tappan 1964): Test free or attached, subglobular, or in groups; aperture absent, single, or multiple.

<==Saccamminidae (see below for synonymy)
    |--Diffusilininae LT64
    |    |--Diffusilina Heron-Allen & Earland 1924 LT64
    |    |    `--*D. humilis Heron-Allen & Earland 1924 LT64
    |    |--Crithionina Goës 1894 [=Arcrithionum Rhumbler 1913] LT64
    |    |    `--*C. mamilla Goës 1894 [=*Arcrithionum mamilla] LT64
    |    |--Daitrona Loeblich & Tappan 1961 LT64
    |    |    `--*D. lens (Goës 1896) [=Crithionina lens] LT64
    |    |--Discobotellina Collins 1958 LT64
    |    |    `--*D. biperforata Collins 1958 LT64
    |    |--Kerionammina Moreman 1933 LT64
    |    |    `--*K. favus Moreman 1933 LT64
    |    |--Masonella Brady 1889 [=Armasonellum Rhumbler 1913] LT64
    |    |    `--*M. planulata Brady 1889 [=*Armasonellum planulatum] LT64
    |    |--Oryctoderma Loeblich & Tappan 1961 LT64
    |    |    |--*O. rotundata (Cushman 1910) [=Crithionina rotundata] LT64
    |    |    |--O. pisum (Goës 1896) [=Crithionina pisum] LT61
    |    |    |--O. rugosa (Goës 1896) [=Crithionina rugosa] LT61
    |    |    `--O. teicherti (Parr 1942) [=Crithionina teicherti] LT61
    |    |--Pseudowebbinella Shchedrina 1962 LT64
    |    |    `--*P. goesi (Höglund 1947) [=Crithionina goesi] LT64
    |    |--Thuramminoides Plummer 1945 LT64
    |    |    `--*T. sphaeroidalis Plummer 1945 LT64
    |    |--Verrucina Goës 1896 [=Arverrucum Rhumbler 1913] LT64
    |    |    `--*V. rudis Goës 1896 [=*Arverrucum rude] LT64
    |    `--Weikkoella Summerson 1958 LT64
    |         `--*W. sphaerica Summerson 1958 LT64
    |--Psammosphaerinae [Stegnammininae, Thekammininae] LT64
    |    |--Blastammina Eisenack 1932 LT64
    |    |    `--*B. polymorpha Eisenack 1932 LT64
    |    |--Ceratammina Ireland 1939 LT64
    |    |    `--*C. cornucopia Ireland 1939 LT64
    |    |--Amphifenestrella Rhumbler 1935 LT64
    |    |    `--*A. wiesneri Rhumbler 1935 LT64
    |    |--Storthosphaera Schulze 1875 [=Arstorthosphaerum Rhumbler 1913; incl. Titanopsis de Folin 1887] LT64
    |    |    |--*S. albida Schulze 1875 [=*Arstorthosphaerum albidum] LT64
    |    |    `--‘*Titanopsis’ irregularis de Folin 1887 LT64
    |    |--Pseudastrorhiza Eisenack 1932 [incl. Parvistellites Wetzel 1951] LT64
    |    |    |--*P. silurica Eisenack 1932 LT64
    |    |    `--P. hospitalis (Wetzel 1951) [=*Parvistellites hospitalis] LT64
    |    |--Stegnammina Moreman 1930 [incl. Raibosammina Moreman 1930, Thekammina Dunn 1942] LT64
    |    |    |--*S. cylindrica Moreman 1930 LT64
    |    |    |--S. mica (Moreman 1930) [=*Raibosammina mica] LT64
    |    |    `--S. quadrangularis (Dunn 1942) [=*Thekammina quadrangularis] LT64
    |    |--Sorosphaera Brady 1879 (see below for synonymy) LT64
    |    |    |--*S. confusa Brady 1879 [=*Arsorophaerum confusum] LT64
    |    |    |--‘*Thuramminopsis’ canaliculata Haeusler 1883 LT64
    |    |    |--S. consociata (Rhumbler in Wiesner 1931) [=*Psammophax consociata] LT64
    |    |    |--S. gracilis (Frentzen 1944) [=*Danubica gracilis] LT64
    |    |    |--S. perforata (Shchedrina 1939) [=*Arenosphaera perforata] LT64
    |    |    `--S. robusta LT64
    |    `--Psammosphaera Schulze 1875 (see below for synonymy) LT64
    |         |--*P. fusca Schulze 1875 [=*Arpsammosphaerum fuscum] LT64
    |         |--P. bowmanni C40
    |         |--P. carnata [=Proteonina carnata] H03
    |         |--P. exigua (Rhumbler 1935) [=*Pilalla exigua] LT64
    |         |--P. frankei (Rhumbler 1935) (see below for synonymy) LT64
    |         |--P. parva C40
    |         `--P. rustica C40
    `--Saccammininae [Pelosininae] LT64
         |--Causia Rhumbler 1938 LT64
         |    `--*C. injudicata Rhumbler 1938 LT64
         |--Pilulina Carpenter 1870 [=Arpilulum Rhumbler 1913] LT64
         |    `--*P. jeffreysii Carpenter 1870 [=*Arpilulum jeffreysii] LT64
         |--Pelosphaera Heron-Allen & Earland 1932 LT64
         |    `--*P. cornuta Heron-Allen & Earland 1932 LT64
         |--Saccamminoides Geroch 1955 LT64
         |    `--*S. carpathicus Geroch 1955 LT64
         |--Stomasphaera Mound 1961 LT64
         |    `--*S. brassfieldensis Mound 1961 LT64
         |--Thurammina Brady 1879 [=Arthyrammum Rhumbler 1913, Thyrammina Rhumbler 1904] LT64
         |    |--*T. papillata Brady 1879 [=*Arthyrammum papillatum, *Thyrammina papillata] LT64
         |    `--T. echinata LT64
         |--Lagenammina Rhumbler 1911 [=Arlagenammum Rhumbler 1913] LT64
         |    |--*L. laguncula Rhumbler 1911 [=*Arlagenammum laguncula] LT64
         |    `--L. compressa (Cushman & McCulloch 1939) [=Proteonina compressa] H03
         |--Technitella Norman 1878 (see below for synonymy) LT64
         |    |--*T. legumen Norman 1878 [=*Artechnitum legumen] LT64
         |    `--T. richardi (de Folin 1887) [=*Dioxeia richardi] LT64
         |--Brachysiphon Chapman 1906 (see below for synonymy) LT64
         |    |--*B. corbuliformis Chapman 1906 [=*Arbrachysiphum corbuliforme] LT64
         |    |--B. australe (Crespin 1958) [=*Sacculinella australe] LT64
         |    `--B. rudis (Parr 1942) [=Hyperammina rudis, *Hyperamminita rudis] LT64
         |--Saccammina Sars in Carpenter 1869 (see below for synonymy) LT64
         |    |--*S. sphaerica Brady 1871 [=*Arsaccammum sphaericum] LT64
         |    |--S. lathrami BL79
         |    `--‘Reophax’ placenta Grzybowski 1897 [=*Placentammina placenta] LT64
         |--Ordovicina Eisenack 1938 (see below for synonymy) LT64
         |    |--*O. oligostoma Eisenack 1938 LT64
         |    |--O. bicuspidata (Dunn 1942) [=*Shidelerella bicuspidata] LT64
         |    |--O. citroniforma (Eisenack 1938) [=*Amphitremoida citroniforma, *Amphitremoidea citroniforma] LT64
         |    |--O. typa (Dunn 1942) [=*Croneisella typa] LT64
         |    `--O. williamsae (Dunn 1942) [=*Gastroammina williamsae] LT64
         `--Pelosina Brady 1879 (see below for synonymy) LT64
              |--*P. variabilis Brady 1879 [=*Arpelosum variabile] LT64
              |--P. arborescens G96
              |--P. bicaudata (Parr 1950) [=*Pelosinella bicaudata] LT64
              |--P. caudata (Montanaro Gallitelli 1955) [=Saccammina caudata, *Caudammina caudata] LT64
              |--P. distoma Millett 1904 [=*Millettina distoma] LT64
              |--P. rotundata C40
              `--P. sphaeriloculum Höglund 1947 [=P. variabilis var. sphaeriloculum, *Globosiphon sphaeriloculum] LT64

Brachysiphon Chapman 1906 [=Arbrachysiphum Rhumbler 1913; incl. Hyperamminita Crespin 1958, Sacculinella Crespin 1958] LT64

Ordovicina Eisenack 1938 [incl. Amphitremoida Eisenack 1938, Amphitremoidea Thalmann 1941, Croneisella Dunn 1942, Gastroammina Dunn 1942, Shidelerella Dunn 1942] LT64

Pelosina Brady 1879 [=Arpelosum Rhumbler 1913; incl. Caudammina Montanaro Gallitelli 1955, Globosiphon Avnimelech 1952, Millettina Avnimelech 1952, Pelosinella Parr 1950] LT64

Psammosphaera Schulze 1875 [=Arphammosphaerum Rhumbler 1913; incl. Pilalla Rhumbler 1935, Psammella Rhumbler 1935] LT64

Psammosphaera frankei (Rhumbler 1935) [=*Psammella frankei; incl. Psammosphaera frankei f. ellipsoides, Psammosphaera frankei f. sphaeroides] LT64

Saccammina Sars in Carpenter 1869 [=Arsaccammum Rhumbler 1913, Saccamina (l. c.); incl. Placentammina Majzon 1943] LT64

Saccamminidae [Kyphamminidae, Pilulinida, Pilulinidae, Pilulinina, Protocystidae, Psammosphaerida, Psammosphaeridae, Saccamminae, Saccamminina, Stegnamminidae]

Sorosphaera Brady 1879 [=Arsorophaerum Rhumbler 1913, Sorophaera (l. c.); incl. Arenosphaera Shchedrina 1939, Danubica Frentzen 1944, Psammophax Rhumbler in Wiesner 1931, Thuramminopsis Haeusler 1883] LT64

Technitella Norman 1878 [=Artechnitum Rhumbler 1913; incl. Dioxeia de Folin 1887, Hyperamminella de Folin 1887] LT64

*Type species of generic name indicated

REFERENCES

[BL79] Basov, V. A., B. G. Lopatin, I. S. Gramberg, A. I. Danjushevskaya, V. Ya. Kaban’kov, V. M. Lazurkin & D. K. Patrunov. 1979. Lower Cretaceous lithostratigraphy near Galicia Bank. Initial Reports of the Deep Sea Drilling Project 47: 683–717.

[C40] Cushman, J. A. 1940. Foraminifera: Their classification and economic use 3^rd ed. Harvard University Press: Cambridge (Massachusetts).

[G96] Gooday, A. J. 1996. Xenophyophores (Protista), including two new species, from two abyssal sites in the northeast Atlantic Ocean. Journal of Foraminiferal Research 26 (3): 193–208.

[H03] Hanagata, S. 2003. Miocene-Pliocene Foraminifera from the Niigata oil-fields region, northeastern Japan. Micropaleontology 49 (4): 293–340.

[LT61] Loeblich, A. R., Jr & H. Tappan. 1961. Remarks on the systematics of the Sarkodina (Protozoa), renamed homonyms and new and validated genera. Proceedings of the Biological Society of Washington 74: 213–234.

[LT64] Loeblich, A. R., Jr & H. Tappan. 1964. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 1. The Geological Society of America and The University of Kansas Press.