Leucosiidae

Ebalia tumefacta, copyright B. E. Picton.

Belongs within: Heterotremata.

The Leucosiidae, purse or pebble crabs, are a group of crabs with a rounded or pentagonal, usually very convex, carapace.

<==Leucosiidae
    |--Heteronucia venusta Nobili 1906 TSH09
    |--Myrine kessleri (Paul’son 1875) MG-H11
    |--Oreophorus reticulatus Adams & White 1849 MG-H11
    |--Parilia major Sakai 1962 MG-H11
    |--Philyra MG-H11
    |--Merocryptus Milne Edwards 1873 B64
    |    `--M. lambriformis Milne Edwards 1873 [incl. Ebalia rugulosa Yokoya 1933] B64
    |--Myra MG-H11
    |    |--M. curtimana Galil 2001 MG-H11
    |    |--M. fugax P71
    |    `--M. subgranulata (Fabricius 1798) KK03
    |--Randallia MG-H11
    |    |--R. eburnea Alcock 1896 MG-H11
    |    `--R. pustuloides Sakai 1961 MG-H11
    |--Persephona Leach 1817 BF07
    |    |--P. lichtensteinii Leach 1817 BF07
    |    |--P. mediterranea (Herbst 1794) BF07
    |    `--P. punctata (Linnaeus 1758) BF07
    |--Leucosia MG-H11
    |    |--L. haematosticta Adams & White 1848 MG-H11
    |    |--L. ocellata Bell 1855 MG-H11
    |    |--L. signata Paulson 1975 KK03
    |    `--L. whitei Bell 1855 MG-H11
    |--Arcania MG-H11
    |    |--A. cornuta (MacGilchrist 1905) MG-H11
    |    |--A. elongata Yokoya 1933 MG-H11
    |    |--A. gracilis (Henderson 1898) MG-H11
    |    |--A. muricata Galil 2002 MG-H11
    |    |--A. novemspinosa (Adams & White 1849) MG-H11
    |    `--A. septemspinosa (Fabricius 1787) MG-H11
    `--Ebalia Leach 1817 B64
         |--E. algirica Lucas 1846 E12
         |--E. cranchii Leach 1817 KK03
         |--E. deshayesii Lucas 1846 E12
         |--E. edwardsii Costa 1838 KK03
         |--E. granulosa Milne-Edwards 1837 KK03
         |--E. insignis Lucas 1846 E12
         |--E. laevis (Bell 1855) [=Phlyxia laevis; incl. P. cheesmani Filhol 1886, E. cheesmani] B64
         |--E. nux Milne-Edwards 1883 KK03
         |--E. tuberculosa (Milne Edwards 1873) [=Persephona tuberculosa; incl. Phlyxia granulosa Haswell 1880] B64
         |--E. tuberosa (Pennant 1777) KK03
         `--E. tumefacta M62

*Type species of generic name indicated

REFERENCES

[B64] Bennett, E. W. 1964. The marine fauna of New Zealand: Crustacea Brachyura. New Zealand Department of Scientific and Industrial Research Bulletin 153: 1–120.

[BF07] Braga, A. A., A. Fransozo, G. Bertini & P. B. Fumis. 2007. Bathymetric distribution and recruitment of the spider crab Libinia spinosa H. Milne Edwards 1834 in the Ubatuba and Caraguatatuba regions, northern coast of São Paulo, Brazil (Crustacea, Brachyura, Majoidea, Pisidae). Senckenbergiana Biologica 87 (1): 7–16.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l'Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[KK03] Kocataş, A., & T. Katağan. 2003. The decapod crustacean fauna of the Turkish seas. Zoology in the Middle East 29: 63–74.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O'Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

[M62] Monniot, F. 1962. Recherches sur les graviers a Amphioxus de la région de Banyuls-sur-Mer. Vie et Milieu 13: 231–322.

[P71] Por, F. D. 1971. One hundred years of Suez Canal—a century of Lessepsian migration: retrospect and viewpoints. Systematic Zoology 20 (2): 138–159.

[TSH09] Titelius, M. A., A. Sampey & C. G. Hass. 2009. Crustaceans of Mermaid (Rowley Shoals), Scott and Seringapatam Reefs, north Western Australia. Records of the Western Australian Museum Supplement 77: 145–176.