Giant tun Tonna galea, copyright Stefano Guerrieri.

Belongs within: Cassoidea.

The Tonnidae, tun shells, are a group of generally large sand-dwelling gastropods most commonly found in deeper water (Wilson & Gillett 1971).

Charactgers (from Wilson & Gillett 1971): Shell generally large, rather thin and inflated; strong spiral ribs present, axial sculpture and varices absent; periostracum thin, smooth, greenish. Anterior canal a U-shaped notch. Umbilicus deep or absent; columellar callus often present, may be reflected over umbilicus and parietal wall to form shield.

Tonnidae [Doliaria, Doliidae]
    |  i. s.: Eudolium pyriforme (Sowerby 1914) MG-H11
    |--Oocorythinae [Oocorythidae] BR05
    |    |--Galeocorys leucodoma HA59
    |    `--Oocorys Fischer 1884 BR05
    |         `--O. tosaensis Habe & Azuma 1959 HA59
    `--Tonninae BR05
         |--Macgillivrayia Forbes 1852 [Macgillivrayiidae] BR05
         |--Galeodolium Sacco 1891 [Galeodoliidae] BR05
         |--Malea Valenciennes 1832 P89
         |    |--M. camura Guppy 1866 P89
         |    |--M. densecostata (Rutsch 1934) P89
         |    |--M. goliath Pilsbry & Johnson 1917 P89
         |    |--M. mareana Weisbord 1962 P89
         |    |--M. petiti Petuch 1989 P89
         |    |--M. pomum (Linnaeus 1758) BW09
         |    `--M. springi Petuch 1989 P89
         `--Tonna Brünnich 1772 [incl. Dolium Lamarck 1801] BR05
              |--*T. galea (Linné 1758) P61, S11 [=Buccinum galea P61]
              |--T. allium Dillwyn 1817 [incl. T. costata Menke 1828] WG71
              |--T. ampullacea Philippi 1845 WG71
              |--T. canaliculata Linnaeus 1758 WG71
              |--T. cepa (Röding 1798) BW09
              |--T. cerevisina Hedley 1919 WG71
              |--T. chinensis (Dillwyn 1817) WG71 [=Buccinum chinense H09]
              |--T. cumingii (Reeve 1849) [=Dolium cumingii] H09
              |--T. dolium (Linnaeus 1758) P61
              |--‘Dolium’ fasciatum F66
              |--T. haurakiensis Hedley 1919 P61 [=T. cerevisina haurakiensis P27]
              |--T. maculosa (Dillwyn 1817) S11
              |--T. maoria Powell 1938 P61
              |--T. melanostoma (Jay 1839) [=Dolium melanostomum] BC01
              |--T. olearium ZLK11
              |--T. perdix (Linnaeus 1758) BW09 [=Buccinum perdix H09]
              |--T. pomum (Linnaeus 1758) [=Buccinum pomum] H09
              |--T. tessellata Lamarck 1816 WG71
              |--T. tetracotula Hedley 1919 P27
              `--T. variegata (Lamarck 1822) MG-H11 [=Dolium variegatum H09]

*Type species of generic name indicated


[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[BW09] Bryce, C., & C. Whisson. 2009. The macromolluscs of Mermaid (Rowley Shoals), Scott and Seringapatam Reefs, Western Australia. Records of the Western Australian Museum Supplement 77: 177–208.

[F66] Fraser, L. 1866. Communication of a list of mollusks collected by R. Swinhoe, Esq., in Formosa. Proceedings of the Zoological Society of London 1866: 146.

[HA59] Habe, T., & M. Azuma. 1959. A new Oocorys from Japan. Nautilus 72 (4): 116.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O’Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

[P89] Petuch, E. J. 1989. New species of Malea (Gastropoda: Tonnidae) from the Pleistocene of southern Florida. Nautilus 103 (3): 92–95.

[P27] Powell, A. W. B. 1927. On a large Tonna and two other gasteropods of Australian origin. Transactions and Proceedings of the New Zealand Institute 57: 559–562.

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4th ed. Whitcombe and Tombs Limited: Christchurch.

[S11] Simone, L. R. L. 2011. Phylogeny of the Caenogastropoda (Mollusca), based on comparative morphology. Arquivos de Zoologia 42 (4): 161–323.

[WG71] Wilson, B. R., & K. Gillett. 1971. Australian Shells: illustrating and describing 600 species of marine gastropods found in Australian waters. A. H. & A. W. Reed: Sydney.

[ZLK11] Zou, S., Q. Li & L. Kong. 2011. Additional gene data and increased sampling give new insights into the phylogenetic relationships of Neogastropoda, within the caenogastropod phylogenetic framework. Molecular Phylogenetics and Evolution 61: 425–435.

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