Zoila friendii, copyright S. & A. Nappo.

Belongs within: Cypraeidae.

Zoila is a genus of medium- to large-sized cowries currently restricted to coasts of Australia, though fossils indicate a wider distribution in the western Pacific in the past. Members of this genus are direct developers, without a planktonic larval stage (Meyer 2003).

<==Zoila Jousseaume 1884 [Zoilinae] BR05
    |  i. s.: Z. gigas (McCoy 1867) [=Cypraea (Zoila) gigas] WG71
    |         Z. mariellae M03
    |         Z. perlae M03
    |--Z. marginata Gaskoin 1849 M03 [=Cypraea (Zoila) marginata WG71]
    |--Z. rosselli Cotton 1948 M03 [=Cypraea (Zoila) rosselli WG71]
    |--Z. venusta Sowerby 1846 M03 [=Cypraea (Zoila) venusta WG71; incl. C. catei Schilder 1963 WG71]
    |    |--Z. v. venusta BC01
    |    `--Z. v. episema (Iredale 1939) BC01 (see below for synonymy)
    `--+--Z. decipiens Smith 1880 M03 [=Cypraea (Zoila) decipiens WG71]
       |--Z. eludens Raybaudi 1991 M03
       `--+--Z. friendii Gray 1831 M03 [=Cypraea (Zoila) friendii WG71; incl. C. friendii f. contraria WG71]
          |    |--Z. f. friendii MG-H11
          |    |--Z. f. jeaniana Cate 1968 MG-H11 [=Cypraea friendii f. jeaniana WG71]
          |    |--Z. ‘jenaiana’ sherylae Raybaudi 1990 M03
          |    `--Z. f. vercoi Schilder 1930 MG-H11 [=Cypraea friendii f. vercoi WG71]
          `--Z. thersites Gaskoin 1849 [=Cypraea (Zoila) friendii thersites] M03

Zoila venusta episema (Iredale 1939) BC01 [=Cypraea venusta episema WG71; incl. Z. venusta sorrentensis Schilder 1963 BC01, C. venusta sorrentensis WG71]

*Type species of generic name indicated


[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1-2): 1-397.

[BC01] Boyko, C. B. & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1-170.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O’Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1-191.

[M03] Meyer, C. P. 2003. Molecular systematics of cowries (Gastropoda: Cypraeidae) and diversification patterns in the tropics. Biological Journal of the Linnean Society 79: 401-459.

[WG71] Wilson, B. R., & K. Gillett. 1971. Australian Shells: illustrating and describing 600 species of marine gastropods found in Australian waters. A. H. & A. W. Reed: Sydney.

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