Isanda coronata, copyright Femorale.

Belongs within: Trochidae.
Contains: Lirularia, Ethminolia.

The Umboniinae are a group of small top shells found mostly on soft substrates in shallower waters. Genera related to Umbonium and Lirularia have a monopectinate ctenidium but the ctenidium remains bipectinate in basal genera such as Monilea and Ethalia (Herbert 1992).

Characters (from Herbert 1992): Radula with teeth of central field reduced, marginals well-developed and numerous, latero-marginal plate present; protoconch small (<250 µm in diameter) and beaked; teleoconch often with scratch-like axial microsculpture; snout with subterminal papillae; forehead frequently narrow or absent and snout reduced in size; eye stalks more or less elongate, curving up under shell margin in life, eyes usually large, terminal; left neck lobe variously modified to form particle exclusion filter, sometimes greatly hypertrophied; foot anteriorly bifid; ctenidium long, extending whole length of mantle cavity; ctenidial filaments elongate, with swollen tips and prominent bursicles, filaments on left side of ctenidial axis reduced or absent; afferent ctenidial membrane long, ctenidium sometimes attached to mantle skirt throughout its length.

Umboniinae [Monileini, Rotelladae, Rotellinae, Umboniidae, Umboniini] BR17
    |  i. s.: ‘Minolia’ edithae Melvill 1891 H92
    |         ‘Minolia’ biangulosa (Adams 1855) H92
    |         ‘Solariella’ dereimsi Dollfus 1911 H92
    |         ‘Solariella’ canaliculata Smith 1872 H92
    |--Callumbonella Thiele 1924 [=Umbotrochus Thiele 1924 non Perner 1903] KC60
    |    `--*C. gorgonarum (Fischer 1883) [=Gibbula gorgonarum, *Umbotrochus gorgonarum] KC60
    |--Inkaba Herbert 1992 H92
    |    `--*I. tonga Herbert 1992 H92
    |--Zethalia Finlay 1926 H92 [incl. Ethaliopsis Cossmann 1918 non Schepman 1908 KC60]
    |    `--*Z. zelandicum (Adams 1854) P61 (see below for synonymy)
    |--Antisolarium Finlay 1926 H92
    |    |--*A. egenum (Gould 1849) P61 (see below for synonymy)
    |    `--A. stoliczkai (Zittel 1865) [=Solariella stoliczkai] F27
    |--Bankivia Krauss 1848 [Bankiviini] BR05
    |    |--B. (Bankivia) fasciata (Menke 1830) H92 (see below for synonymy)
    |    `--B. (Leiopyrga Adams & Adams 1863) KC60
    |         `--B. (L.) lineolaris (Gould 1861) KC60 (see below for synonymy)
    |--Pseudominolia Herbert 1992 H92
    |    |--*P. splendens (Sowerby 1897) [=Solariella splendens; incl. Minolia variegata Odhner 1919] H92
    |    |--‘Solariella’ algoensis Thiele 1925 (see below for synonymy) H92
    |    `--‘Minolia’ climacota Melvill 1897 H92
    |--Talopena Iredale 1918 H92
    |    |--*T. incerta (Iredale 1913) H92 [=Monilea incerta KC60]
    |    |--T. gloriola Iredale 1929 H92
    |    |--T. lifuana (Fischer 1878) H92 [=Trochus lifuanus H92, Monilea lifuana H09]
    |    |--T. vernicosa (Gould 1861) MG-H11
    |    `--T. vitiliginea (Menke 1843) F27, H09 [=Trochus vitiligineus H09, Monilea vitiliginea H09]
    |--Umbonium Link 1807 H92 [=Globulus Schumacher 1817 KC60, Rotella Lamarck 1822 BR05]
    |    |  i. s.: U. costatum (Valenciennes in Kiener 1838–1839) H92
    |    |         U. eloiseae Dance, Moolenbeek & Dekker 1992 BC01
    |    |         *Rotella’ lineolata Lamarck 1822 BR17
    |    |         ‘Rotella’ neozelanica H86
    |    |--*U. (U.) vestiarium (Linnaeus 1758) [=Trochus vestiarius, *Globulus vestiarius, Rotella vestiaria] KC60
    |    |--U. (Ethaliella Pilsbry 1905) KC60
    |    |    `--U. (*E.) floccata (Sowerby 1903) [=Ethalia floccata] KC60
    |    |--U. (Protorotella Makiyama 1925) KC60
    |    |    `--U. (*P.) yuantaniensis (Makiyama 1925) [=*Protorotella yuantaniensis] KC60
    |    `--U. (Suchium Makiyama 1925) KC60
    |         `--U. (*S.) suchiense Yokoyama 1923 KC60
    |--Ethminolia Iredale 1924 [incl. Conotalopia Iredale 1929, Sericominolia Kuroda & Habe 1952] H92
    |    |--*E. probabilis Iredale 1924 H92
    |    |--E. durbanensis (Kilburn 1977) [=Solariella durbanensis] H92
    |    |--E. eilikrines (Melvill 1891) H92
    |    |--E. eudeli (Deshayes 1863) [=Trochus eudeli] H92
    |    |--E. gravieri (Lamy 1909) [=Cyclostrema gravieri] H92
    |    |--‘Minolia’ henniana Melvill 1891 H92 [=Isanda (*Conotalopia) henniana KC60, Monilea henniana H09]
    |    |--‘Solariella’ illustris Sturany 1903 H92
    |    |--E. impressa (Nevill & Nevill 1869) (see below for synonymy) H92
    |    |--E. nektonica (Okutani 1961) (see below for synonymy) H92
    |    |--E. sculpta (Sowerby 1897) [=Solariella sculpta, Minolia sculpta] H92
    |    |--E. stearnsii (Pilsbry 1895) [=Minolia stearnsii, *Sericominolia stearnsii, Solariella stearnsi] H92
    |    `--E. vitiliginea (Menke 1843) H92
    |--Lirularia Dall 1909 H92, BR05 [Lirulariinae]
    |    |--*L. lirulata (Carpenter 1863) BR17 [=Margarites lirulata BR17; incl. Margarita lirulata var. conica C64]
    |    |    |--L. l. lirulata O27
    |    |    |--L. l. obsoleta (Carpenter 1864) [=Margarites lirulata obsoleta] O27
    |    |    `--L. l. subelevata (Carpenter 1864) [=Margarites lirulata subelevata] O27
    |    |--L. acuticostata (Carpenter 1864) O27 [=Margarites (Lirularia) acuticostatus B56]
    |    |--L. althorpensis (Dall 1919) [=Margarites (Lirularia) althorpensis] O27
    |    |--L. antoniae Rubio & Rolán 1997 BC01
    |    |--L. inflatula (Dall 1920) [=Margarites (Lirularia) inflatula] O27
    |    |--L. iridescens (Schrenck 1863) H84
    |    |--L. lacunata (Carpenter 1864) [=Gibbula lacunata, Margarites (Lirularia) lacunata] O27
    |    |--L. pedroana [=Margarites (Lirularia) pedroana] B56
    |    |--L. smithi (Bartsch 1927) [=Margarites (Lirularia) smithi] O27
    |    `--L. succincta (Carpenter 1864) [=Gibbula succincta, Margarites (Lirularia) succincta] O27
    |--Ethalia Adams & Adams 1854 [incl. Liotrochus Fischer 1878] H92
    |    |--*E. guamensis (Quoy & Gaimard 1834) H92 (see below for synonymy)
    |    |    |--E. g. guamensis H92
    |    |    `--E. g. selenomphala Pilsbry 1905 H92
    |    |--E. bysma Herbert 1992 H92
    |    |--E. carneolata Melvill 1897 [=Rotella carneolata; incl. E. carneolata var. rubrostrigata Melvill 1904] H92
    |    |--E. electra Herbert 1992 H92
    |    |--E. gilchristae Herbert 1992 H92
    |    |--E. lamprus (Watson 1880) [=Trochus (Solariella) lamprus] H92
    |    |--E. minolina Melvill 1897 [incl. E. minolina var. infralaevior Schepman 1908] H92
    |    |--E. minuta [=Vitrinella minuta] C64
    |    |--E. pulchella (Adams 1855) [=Isanda pulchella] H09
    |    |--E. regularis [=Vitrinella regularis] C64
    |    |--E. striolata (Adams 1855) [incl. E. trilobata Sowerby in Reeve 1875-1878] H92
    |    |--E. supravallata Carpenter 1864 C64
    |    `--‘Vitrinella’ valvatoides C64
    |--Monilea Swainson 1840 [=Talopia Gray 1842; Talopiidae, Talopiini] BR05
    |    |  i. s.: M. angulata Adams 1851 H08
    |    |         M. corrugata [=Trochus corrugatus] B79
    |    |         M. glaphyrella (Melvill & Standen 1895) [=Minolia glaphyrella] H09
    |    |         M. lentiginosa Adams 1852 H09
    |    |         M. patricia (Philippi 1851) H92
    |    |         M. philippii Adams 1855 [=Ethalia philippii] H92
    |    |         M. pudibunda (Fischer 1878) [=Trochus pudibundus] H09
    |    |         M. stoliczkai H86
    |    |         M. tropicalis Hedley 1907 H09
    |    |         M. vernicosa Gould 1861 H92
    |    |--*M. (Monilea) callifera (Lamarck 1822) KC60 [=Trochus calliferus KC60, *Talopia callifera BR17]
    |    |--M. (Priotrochus Fischer 1879) KC60, H92 [=Aphanotrochus Martens 1880 KC60]
    |    |    `--M. (*P.) obscura (Wood 1828) [=Trochus obscurus, *Aphanotrochus obscurus] KC60
    |    `--M. (Rossiteria Brazier 1895) [=Solanderia Fischer in Kiener 1879 non Duchassaing & Michelin 1846] KC60
    |         `--M. (*R.) nuclea (Philippi 1849) [=Trochus nucleus] KC60
    `--Isanda Adams & Adams 1854 [Isandini] BR05
         |  i. s.: I. hornungi (Bisacchi 1931) H92
         |--*I. (Isanda) coronata Adams 1854 KC60
         |--I. (Archiminolia Iredale 1929) KC60
         |    `--I. (*A.) oleacea (Hedley & Petterd 1905) [=Monilea oleacea] KC60
         |--I. (Conominolia Finlay 1927) KC60
         |    |--I. (*C.) conica (Marshall 1917) [=Heliacus conicus] KC60
         |    `--‘Conominolia’ sulcatina (Suter 1917) [=Solariella sulcatina] F27
         |--I. (Parminolia Iredale 1929) KC60
         |    `--I. (P.) apicina (Gould 1861) (see below for synonymy) KC60
         |--I. (Umbonella Adams 1863) KC60
         |    `--I. (*U.) murrea (Reeve 1848) [=Turbo murreus] KC60
         |--I. (Vanitrochus Iredale 1929) [incl. Conotrochus Pilsbry 1889 non Schröter 1863] KC60
         |    `--I. (*V.) tragema (Melvill & Standen 1869) [=Solariella tragema] KC60
         `--I. (Waisiolia Beets 1942) KC60
              `--I. (*W.) jucanda Beets 1942 KC60

*Antisolarium egenum (Gould 1849) P61 [=Solarium (Torinia) egenum H92, Minolia egena F27, Solariella egena F27; incl. Margarita dilecta Adams 1855 H92, Solariella dilecta H92, Trochus (Solariella) dilectus H92]

Bankivia (Bankivia) fasciata (Menke 1830) H92 [=Phasianella fasciata KC60, Cantharidus fasciatus F27; incl. Bankivia lugubris H08, *B. varians Krauss 1848 KC60]

Bankivia (Leiopyrga) lineolaris (Gould 1861) KC60 [=Cantharidus lineolaris KC60; incl. *Leiopyrga picturata Adams & Adams 1863 KC60, C. picturatus F27]

*Ethalia guamensis (Quoy & Gaimard 1834) H92 [=Rotella guamensis H09, Umbonium (*Ethalia) guamense KC60; incl. R. montrouzieri Souverbie 1860 H92, *Liotrochus montrouzieri H92]

Ethminolia impressa (Nevill & Nevill 1869) [=Solarium impressum, Minolia impressa; incl. M. eucoronata Sowerby 1905] H92

Ethminolia nektonica (Okutani 1961) [=Solariella nektonica; incl. S. nektonica tajimensis Okutani & Sakurai 1968] H92

Isanda (Parminolia) apicina (Gould 1861) [=Monilea apicina; incl. Minolia agapeta Melvill & Standen 1896, *Parminolia agapeta] KC60

‘Solariella’ algoensis Thiele 1925 [incl. Margarita articulata Gould 1861 non Adams 1853, Gibbula articulata, Minolia articulata, Pseudominolia articulata, Mi. bleeki Thiele 1925] H92

*Zethalia zelandica (Adams 1854) P61 [=Umbonium zelandicum P61, Ethalia zelandica Rousseau 1854 F27; incl. Globulus anguliferus Philippi 1854 non Sowerby 1840 F27, *Ethaliopsis angulifera F27, Umbonium anguliferum F27]

*Type species of generic name indicated


[B56] Berry, S. S. 1956. Mollusca dredged by the Orca off the Santa Barbara Islands, California, in 1951. Journal of the Washington Academy of Sciences 46 (5): 150–157.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[B79] Brazier, J. 1879. List of marine shells collected on Fitzroy Island, north coast of Australia; with notes on their geographical range. Journal of Conchology 2: 186–199.

[C64] Carpenter, P. P. 1864. Supplementary report on the present state of our knowledge with regard to the Mollusca of the west coast of North America. Report of the British Association for the Advancement of Science 33: 517–686.

[F27] Finlay, H. J. 1927. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[H08] Hedley, C. 1908. Studies on Australian Mollusca. Part X. Proceedings of the Linnean Society of New South Wales 33: 456–489, pls 7–10.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[H92] Herbert, D. G. 1992. Revision of the Umboniinae in southern Africa and Mozambique (Mollusca: Prosobranchia: Trochidae). Annals of the Natal Museum 33 (2): 379–459.

[H84] Hickman, C. S. 1984. Implications of radular tooth-row functional integration for archaeogastropod systematics. Malacologia 25 (1): 143–160.

[H86] Hutton, F. W. 1886. The Mollusca of the Pareora and Oamaru systems of New Zealand. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 205–237.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I169–I331. Geological Society of America, and University of Kansas Press.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O’Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. 2 pt 3. Stanford University Press: Stanford University (California).

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4th ed. Whitcombe and Tombs Limited: Christchurch.

Last updated: 19 October 2021.

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