Mycobates

Mycobates conitus, copyright Monica Young.

Belongs within: Ceratozetoidea.

Mycobates is a genus of oribatid mites found in the Holarctic region (Subías 2004).

Characters (from Balogh & Balogh 1992): Rostrum rounded or bidentate. Prodorsum with lamellae and translamella present, lamellae converging, translamella much longer than cuspides; anterior tectum of notogaster overhanging bothridium, with interlamellar setae originating under tectum. Notogaster with ten pairs of setae or setal alveoli; four pairs of areae porosae; pteromorphae movable. Six pairs of genital setae; one pair of aggenital setae; two pairs of anal setae; three pairs of adanal setae.

<==Mycobates Hull 1916 S04
    |--M. (Mycobates) S04
    |    |--*M. (M.) parmeliae (Michael 1884) (see below for synonymy) S04
    |    |--M. (M.) acuspidatus Behan-Pelletier, Eamer & Clayton 2001 S04
    |    |--M. (M.) austroamericanus Hammer 1958 S04
    |    |--M. (M.) cambricus (Hull 1916) [=Melanozetes cambricus] S04
    |    |--M. (M.) carli (Schweizer 1922) [=Oribata carli] S04
    |    |--M. (M.) corticeus Bvehan-Pelletier, Eamer & Clayton 2001 S04
    |    |--M. (M.) debilis Mihelčič 1957 S04
    |    |--M. (M.) exigualis Behan-Pelletier 1994 S04
    |    |--M. (M.) ezoensis Fujikawa 1982 S04
    |    |--M. (M.) flabelliger (Berlese 1908) [=Sphaerozetes flabelliger] S04
    |    |--M. (M.) hylaeus Behan-Pelletier 1994 S04
    |    |--M. (M.) integer Mihelčič 1957 S04
    |    |--M. (M.) minor Subías, Kahwash & Ruiz 1990 S04
    |    |--M. (M.) monodactylus Shaldybina 1970 S04
    |    |--M. (M.) perates Behan-Pelletier 1994 S04
    |    |--M. (M.) rileyi (Haller 1884) (n. d.) [=Oribata rileyi] S04
    |    |--M. (M.) royi Palacios-Vargas & Vázquez 1988 S04
    |    |--M. (M.) tridentatus Weigmann 1976 S04
    |    `--M. (M.) yukonensis Behan-Pelletier 1994 S04
    `--M. (Calyptozetes Thor 1930) [incl. Permycobates Strenzke 1954] S04
         |--M. (*C.) sarekensis (Trägardh 1910) [=Oribata sarekensis; incl. M. consimilis Hammer 1952] S04
         |--M. (C.) alpinus (Willmann 1951) [=Calyptozetes alpinus] S04
         |--M. (C.) altus Behan-Pelletier 1994 S04
         |--M. (C.) angulatus (Koch 1839) [=Oribata angulata] S04
         |--M. (C.) azaleos Behan-Pelletier 1994 S04
         |--M. (C.) beringianus Behan-Pelletier 1994 S04
         |--M. (C.) bicornis (Strenzke 1954) [=Permycobates bicornis] S04
         |--M. (C.) brevilamellatus Behan-Pelletier 1994 S04
         |--M. (C.) conitus Hammer 1952 S04
         |--M. (C.) cribelliger (Berlese 1904) [=Oribata cribelliger] S04
         |--M. (C.) dryas Behan-Pelletier 1994 S04
         |--M. (C.) hammerae Behan-Pelletier 1994 S04
         |--M. (C.) incurvatus Hammer 1952 S04
         |--M. (C.) monocornis Aoki 1973 S04
         |--M. (C.) occidentalis Behan-Pelletier 1994 S04
         |--M. (C.) patrius Shaldybina 1970 S04
         |--M. (C.) punctatus Hammer 1955 S04
         `--M. (C.) tridactylus Willmann 1929 S04

*Mycobates (Mycobates) parmeliae (Michael 1884) [=Oribata parmeliae; incl. Sphaerozetes (Trichoribates) lenicomus Berlese 1910, Mycobates nadiji Paulitchenko 1991] S04

*Type species of generic name indicated

REFERENCES

Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Chamobates

Chamobates rastratus, copyright R. Penttinen.

Belongs within: Ceratozetoidea.

Chamobates is a genus of oribatid mites found in Eurasia and North America (Subías 2004).

Characters (from Balogh & Balogh 1992): Prodorsum with long lamellae present, translamella absent; lamellar setae originating near rostral region, on surface of prodorsum near end of lamella; interlamellar setae originating on prodorsum near dorsosejugal suture. Notogaster with ten pairs of setal alveoli only; four pairs of areae porosae; pteromorphae immovable. Six pairs of genital setae; one pair of aggenital setae; two pairs of anal setae; three pairs of adanal setae. Legs tridactyle.

<==Chamobates Hull 1916 S04
    |--C. (Chamobates) S04
    |    |--*C. (C.) cuspidatus (Michael 1884) (see below for synonymy) S04
    |    |--C. (C.) alpinus Schweizer 1956 [=C. cuspidatus alpinus] S04
    |    |--C. (C.) apathyi (Tafner 1905) (n. d.) [=Oribata apathyi] S04
    |    |--C. (C.) bispinosus Mahunka 1987 S04
    |    |--C. (C.) caucasicus Shaldybina 1969 S04
    |    |--C. (C.) confusus Subías 2000 (see below for synonymy) S04
    |    |--C. (C.) cuspidatiformis (Trägardh 1904) [=Notaspis cuspidatiformis] S04
    |    |--C. (C.) dentatus Mihelčič 1956 S04
    |    |--C. (C.) dentotutorii Shaldybina 1969 S04
    |    |--C. (C.) egenus (Berlese 1910) [=Euzetes egenus] S04
    |    |--C. (C.) geminus Fujikawa 1997 S04
    |    |--C. (C.) hauseri Mahunka 1980 S04
    |    |--C. (C.) ilinoisensis (Ewing 1909) (n. d.) [=Oribata ilinoisensis] S04
    |    |--C. (C.) interpositus Pschorn-Walcher 1953 S04
    |    |--C. (C.) javensis (Hammer 1979) [=Sphaerozetes javensis; incl. Unguizetes hammerae Balogh & Balogh 2002] S04
    |    |--C. (C.) lapidarius (Lucas 1847) (n. d.) S04, E12 [=Oribates lapidarius E12, Oribata lapidaria S04]
    |    |--C. (C.) longipilis Willmann 1953 S04
    |    |--C. (C.) pedemontanus (Berlese 1910) (n. d.) [=Sphaerozetes pedemontanus] S04
    |    |--C. (C.) perezinigoi Subías 1977 S04
    |    |--C. (C.) pusillus (Berlese 1895) S04 (see below for synonymy)
    |    |--C. (C.) schuetzi (Oudemans 1902) [=Notaspis schuetzi] S04
    |    |--C. (C.) subglobulus (Oudemans 1900) (see below for synonymy) S04
    |    |--C. (C.) tumidisetosus Monson 2002 S04
    |    `--C. (C.) vartismithi Hull 1916 (n. d.) S04
    `--C. (Xiphobates Paulitchenko 1993) S04
         |--C. (*X.) voigtsi (Oudemans 1902) [=Notaspis voigtsi] S04
         |--C. (X.) callipygis Paulitchenko 1991 S04
         |--C. (X.) dactyloscopicus Bernini & Mahunka 1982 S04
         |--C. (X.) depauperatus (Berlese 1886) [=Oribata depauperata] S04
         |--C. (X.) kievensis Shaldybina 1980 S04
         |--C. (X.) rastratus (Hull 1914) [=Oribata rastrata; incl. C. spinosus Sellnick 1928] S04
         `--C. (X.) sergienkae Shaldybina 1980 S04

Chamobates (Chamobates) confusus Subías 2000 [incl. C. angulatus Mihelčič 1957 (n. d.), C. triangularis Mihelčič 1957 (n. d.)] S04

*Chamobates (Chamobates) cuspidatus (Michael 1884) [=Oribata cuspidata; incl. O. dominae Hull 1914, O. lucifer Hull 1914] S04

Chamobates (Chamobates) pusillus (Berlese 1895) S04 [=Oribata pusilla S04; incl. Notaspis cuspidatus borealis Trägardh 1902 S04, Chamobates borealis MH04, C. incisus Hammen 1952 S04]

Chamobates (Chamobates) subglobulus (Oudemans 1900) [=Notaspis subglobula; incl. Oribata facula Koch 1839 (n. d.), Chamobates nasutus Mihelčič 1957, Euzetes nigerrimus Berlese 1908, Oribata omissa Warburton & Pearce 1905] S04

*Type species of generic name indicated

REFERENCES

Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l'Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[MH04] Maraun, M., M. Heethoff, K. Schneider, S. Scheu, G. Weigmann, J. Cianciolo, R. H. Thomas & R. A. Norton. 2004. Molecular phylogeny of oribatid mites (Oribatida, Acari): evidence for multiple radiations of parthenogenetic lineages. Experimental and Applied Acarology 33 (3): 183–201.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Diapterobates

Diapterobates notatus, from the UNIS University Centre in Svalbard.

Belongs within: Ceratozetoidea.

Diapterobates is a genus of oribatid mites found in the Holarctic region (Subías 2004). The type species Diapterobates humeralis has been studied as a control agent for the hemlock woolly adelgid Adelges tsugae, as in the process of feeding on the woolly filaments covering this insect's ovisacs it tends to dislodge and destroy the eggs themselves.

Characters (from Balogh & Balogh 1992): Prodorsum with lamellae and translamella present; sensillus short with dilated head. Notogaster with thirteen pairs of setae; four pairs of areae porosae; areae porosae Aa elongate, thrice or more as long as wide; pteromorphae immovable. Six pairs of genital setae; one pair of aggenital setae; two pairs of anal setae; three pairs of adanal setae. Legs tridactyle.

<==Diapterobates Grandjean 1936 S04
    |--D. humeralis (Hermann 1804) S04 (see below for synonymy)
    |--D. altimontanus Hammer 1977 S04
    |--D. arnoldii Krivolutsky 1966 S04
    |--D. bogduulensis Bayartogtokh & Aoki 1998 S04
    |--D. crinitus Pankov 1982 S04
    |--D. izuensis Suzuki 1971 S04
    |--D. japonicus Aoki 1982 S04
    |--D. montivagus (Hull 1914) (n. d.) [=Oribata montivaga] S04
    |--D. nayoroensis Fujikawa 1984 S04
    |--D. notatus (Thorell 1871) (see below for synonymy) S04
    |--D. oblongus (Koch 1879) [=Oppia oblonga] S04
    |--D. oromurcii Shaldybina 1970 S04
    |--D. pusillus Aoki 1969 S04
    |    |--D. p. pusillus S04
    |    `--D. p. cuspidatus Choi 1986 S04
    |--D. reticulatus (Koch 1879) (see below for synonymy) S04
    |--D. rostralis Shaldybina 1971 S04
    |--D. rostrotuberculatus Shaldybina 1971 S04
    |--D. rotundocuspidatus Shaldybina 1970 [incl. D. siccatus Behan-Pelletier 1986] S04
    |--D. sitnikovae Shaldybina 1970 S04
    |--D. songliensis Choi 1986 S04
    |--D. unimaculatus (Banks 1906) [=Galumna unimaculata] S04
    `--D. variabilis Hammer 1955 S04
         |--D. v. variabilis S04
         |--D. v. altissimus Piffl 1971 S04
         |--D. v. honshuensis Aoki 1982 S04
         `--D. v. yezoensis Aoki 1982 S04

Diapterobates humeralis (Hermann 1804) S04 [=Notaspis humeralis S04, Oribates humeralis CF77; incl. Sphaerozetes (Trichoribates) numerosus Sellnick 1924 S04, *Diapterobates numerosus S04, Oribata picipes Koch 1839 S04]

Diapterobates notatus (Thorell 1871) [=Oribata notata; incl. Claviceps bimaculatus Koch 1879, Sphaerozetes strandi Berlese 1910] S04

Diapterobates reticulatus (Koch 1879) [=Oribata reticulata; incl. Sphaerozetes (Trichoribates) principalis Berlese 1914] S04

*Type species of generic name indicated

REFERENCES

Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[CF77] Canestrini, G., & F. Fanzago. 1877. Intorno agli Acari Italiani. Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti, Serie 5, 4: 69–208, pls 2–7.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Last updated: 27 June 2017.

Anellozetes

Dorsal (left) and ventral views of Anellozetes fusiformis, with close-up of sensillus, from Hammer (1970).

Belongs within: Ceratozetoidea.

Anellozetes is a genus of humerobatid oribatid mites with a scattered distribution in Africa, Asia, Oceania and South America (Subías 2004).

<==Anellozetes Hammer 1962 S04
    |--*A. muscicola Hammer 1962 S04
    |--A. auriculatus (Mahunka 1984) [=Kilimabates auriculatus] S04
    |--A. bicuspidatus (Mahunka 1985) [=Kilimabates bicuspidatus] S04
    |--A. discifer Hammer 1973 S04
    |--A. fusiformis Hammer 1970 S04
    |--A. intermedius Hammer 1967 S04
    |--A. longicaulis Hammer 1967 S04
    |--A. luteus Hammer 1967 S04
    |--A. neonominatus Subías 2004 (see below for synonymy) S04
    `--A. translamellatus (Shaldybina 1971) [=Humerobates translamellatus] S04

Anellozetes neonominatus Subías 2004 [=Kilimabates translamellatus Mahunka 1985 non Humerobates translamellatus Shaldybina 1971] S04

*Type species of generic name indicated

REFERENCES

Hammer, M. 1970. A few oribatid mites from Easter Island. Pacific Insects 12 (2): 279–289.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Africoribates

Dorsal (left) and ventral views of Africoribates australis, from Balogh & Balogh (1992 vol. 2).

Belongs within: Ceratozetoidea.

Africoribates is a genus of oribatid mites found in sub-Saharan Africa (Subías 2004).

Characters (from Balogh & Balogh 1992): Prodorsum with lamellae and translamella present. Notogaster with pairs of setae; four pairs of areae porosae; pteromorphae movable. Six pairs of genital setae; one pair of aggenital setae; two pairs of anal setae; three pairs of adanal setae. Legs tridactyle.

<==Africoribates Evans 1953 S04
    |--*A. ornatus Evans 1953 S04
    |--A. australis Mahunka 1985 S04
    |--A. depilatus (Berlese 1910) [=Peloribates depilatus] S04
    |--A. evansi Balogh 1959 S04
    |--A. glabratus (Berlese 1908) [=Peloribates glabratus] S04
    |--A. macfarlanei Balogh 1959 S04
    |--A. maximus Mahunka 1984 S04
    |--A. pilosus (Mahunka 1985) [=Kilimabates pilosus] S04
    |--A. polygonatus (Mahunka 1985) [=Kilimabates polygonatus] S04
    `--A. undulatus Balogh 1959 S04

*Type species of generic name indicated

REFERENCES

Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Humerobates

Aggregation of Humerobates rostrolamellatus, copyright Theodoor Heijerman.

Belongs within: Ceratozetoidea.

Humerobates is a cosmopolitan genus of oribatid mites in which the pteromorphs may be movable or immovable, and the notogastral setae are represented by alveoli only (Balogh & Balogh 1992). The subgenus Cordylobates is represented by a single species H. (C.) fragilis from Ascension Island in the South Atlantic.

Characters (from Balogh & Balogh 1992): Prodorsum with lamellae present, translamella absent. Notogaster with ten pairs of setal alveoli only; four pairs of areae porosae; pteromorphs well-developed, movable or immovable. Six pairs of genital setae; one pair of aggenital setae; two pairs of anal setae; three pairs of adanal setae. Legs tridactyle.

<==Humerobates Sellnick 1928 S04
    |--H. (Humerobates) (see below for synonymy) S04
    |    |--H. (H.) rostrolamellatus Grandjean 1936 S04 (see below for synonymy)
    |    |    |--H. r. rostrolamellatus S04
    |    |    |--H. r. giganteus Willmann 1939 [incl. H. fungorum giganteus] S04
    |    |    |--H. r. guadarramicus Pérez-Íñigo 1972 [incl. Chamobates magnus Mihelčič 1957 (n. d.)] S04
    |    |    `--H. r. mauritius Jacot 1936 [=H. humeralis mauritius] S04
    |    |--H. (H.) africanus (Mahunka 1984) [=Baloghobates africanus] S04
    |    |--H. (H.) cornutus (Sellnick 1959) S04 [=*Trihumerozetes cornutus BB92]
    |    |--H. (H.) crozetensis (Richters 1908) S04 (see below for synonymy)
    |    |--H. (H.) flechtmanni Pérez-Íñigo & Pérez-Íñigo 1993 S04
    |    |--H. (H.) fungorum (Linnaeus 1758) (n. d.) [=Acarus fungorum] S04
    |    |--H. (H.) nudus (Hammer 1967) S04 [=*Baloghobates nudus CH98]
    |    |--H. (H.) papuanus Balogh 1970 S04
    |    |--H. (H.) parvoglobosus (Hammer 1967) [=Baloghobates parvoglobosus] S04
    |    |--H. (H.) perkinsi Jacot 1934 S04 [=H. humeralis perkinsi S04, H. rostrolamellatus perkinsi SN98]
    |    |--H. (H.) pomboi Pérez-Íñigo 1992 S04
    |    |--H. (H.) setosus Behan-Pelletier & Mahunka 1993 S04
    |    |--H. (H.) taiwanensis Tseng 1984 S04
    |    |--H. (H.) varius Ohkubo 1982 S04
    |    `--H. (H.) zumpti Sellnick 1957 S04
    `--H. (Cordylobates Luxton 1995) S04
         `--H. (*C.) fragilis (Luxton 1995) [=*Cordylobates fragilis] S04

Humerobates Sellnick 1928 (Humerobates) [incl. Antarctozetes Balogh 1961, Baloghobates Hammer 1967, Banksinus Jacot 1938, Jeannelia Dalenius 1958 non Raffray 1913, Trihumerozetes Sellnick 1959] S04

Humerobates (Humerobates) crozetensis (Richters 1908) S04 [=Oribata crozetensis S04, *Antarctozetes crozetensis CH98, *Jeannelia crozetensis CH98]

Humerobates (Humerobates) rostrolamellatus Grandjean 1936 S04 [=Banksinus arboreus rostrolamellatus J38; incl. Oribata arborea Banks 1895 (n. d.) S04, *Banksinus arboreus J38, Humerobates humeralis arborea S59, Sphaerobates dentatus Mihelčič 1957 S04, Notaspis humeralis Sellnick 1928 non Hermann 1804 S04, *Humerobates humeralis S04, Murcia rubra Koch 1841 (n. d.) S04]

*Type species of generic name indicated

REFERENCES

[BB92] Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[CH98] Colloff, M. J., & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

[J38] Jacot, A. P. 1938. Some new western North Carolina moss-mites. Proceedings of the Entomological Society of Washington 40 (1): 10–15.

[S59] Sellnick, M. 1959. Acarina from southeastern Polynesia—II (Oribatidae). Occasional Papers of the Bernice P. Bishop Museum 22 (9): 109–152.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

[SN98] Swift, S. F., & R. A. Norton. 1998. Preliminary report on oribatid mite (Acari: Oribatida) diversity in the Hawaiian Islands. Bishop Museum Occasional Papers 57: 1–44.

Last updated: 15 December 2018.

Protoribates (Protoribates)

Protoribates lophothrichus, copyright R. Penttinen.

Belongs within: Haplozetidae.

The type subgenus of the genus Protoribates is a cosmopolitan group of oribatid mites (Subías 2004), albeit one of uncertain composition owing to confusion over the identity of the type species. True Protoribates have movable pteromorphs and had previously been referred to by the junior name Xylobates (Colloff & Halliday 1998).

Characters (from Balogh & Balogh 1992a, as Protoribates and Xylobates): Prodorsum with lamellae present, translamella absent; interlamellar setae long. Dorsosejugal suture present. Notogaster smooth; ten pairs of notogastral setae short but present; four pairs of areae porosae; areae porosae Aa, A2 and A3 round; pteromorphs well developed, movable. Four to six pairs of genital setae; one pair of aggenital setae; two pairs of anal setae; three pairs of adanal setae. Legs monodactyle.

<==Protoribates Berlese 1908 (Protoribates) (see below for synonymy) S04
    |--*P. (P.) dentatus (Berlese 1883) S04 (see below for synonymy) S04
    |--P. (P.) albidus (Ewing 1908) S04 [=Oribata albida S04, Oribatula albida CH98, *Propescheloribates albidus CH98]
    |--P. (P.) angulatus Choi 1986 S04
    |--P. (P.) antillensis (Mahunka 1985) [=Xylobates antillensis] S04
    |--P. (P.) bayanicus Bayartogtokh 2000 S04
    |--P. (P.) boraboraensis Sellnick 1959 S04
    |--P. (P.) brevisetosus (Fujita 1989) [=Xylobates brevisetosus] S04
    |--P. (P.) capucinus Berlese 1908 S04 [=Xylobates capucinus BB92]
    |    |--P. c. capucinus S04 (see below for synonymy)
    |    |--P. c. bonairensis Willmann 1936 S04
    |    `--P. c. tentaculus Mihelčič 1958 S04
    |--P. (P.) concretus (Kramer 1898) (n. d.) [=Oribata concreta] S04
    |--P. (P.) crassisetiger Choi 1986 S04
    |    |--P. c. crassisetiger S04
    |    `--P. c. nipponicus Fujita 1989 S04
    |--P. (P.) diani (Mahunka 1986) [=Xylobates diani] S04
    |--P. (P.) geonjiensis Choi 1994 S04
    |--P. (P.) gracilis (Aoki 1982) [=Xylobates gracilis] S04
    |--P. (P.) hakonensis Aoki 1994 S04
    |--P. (P.) halleri (Michael 1898) (n. d.) [=Oribata halleri; incl. O. simplex Haller 1884 non Gervais 1849] S04
    |--P. (P.) imperfectus (Banks 1906) [=Galumna imperfecta] S04
    |--P. (P.) iracemae Pérez-Íñigo & Baggio 1994 S04
    |--P. (P.) longiuscula (Koch 1841) (n. d.) [=Zetes longiuscula] S04
    |--P. (P.) lophothrichus (Berlese 1904) S04 (see below for synonymy)
    |--P. (P.) madagascarensis (Balogh 1961) [=Scheloribates madagascarensis] S04
    |--P. (P.) magnus Aoki 1982 S04
    |--P. (P.) mollicoma (Hammer 1973) [=Xylobates mollicoma] S04
    |--P. (P.) myrmecophilus Aoki & Ito 1997 S04
    |--P. (P.) nagaroensis (Fujita 1989) [=Xylobates nagaroensis] S04
    |--P. (P.) natalensis (Pletzen 1963) [=Scheloribates natalensis] S04
    |--P. (P.) oblongus (Ewing 1909) [=Oribata oblonga; incl. O. longa Ewing 1909, Xylobates longisetae Jacot 1937] S04
    |--P. (P.) paracapucinus (Mahunka 1988) [=Xylobates paracapucinus] S04
    |--P. (P.) ‘parvulus’ Willmann 1931 non Berlese 1916 S04
    |--P. (P.) pembertoni (Jacot 1934) [=Xylobates pembertoni] S04
    |--‘Xylobates’ punctulatus (Sellnick 1925) C-R79
    |--P. (P.) robustior (Jacot 1937) (see below for synonymy) S04
    |--P. (P.) rodriguezi (Mahunka 1988) [=Xylobates rodriguezi] S04
    |--P. (P.) rotundus (Aoki 2002) [=Xylobates rotundus] S04
    |--P. (P.) souchnaiensis Abdel-Hamid 1964 S04
    |--P. (P.) tenuis (Wen, Aoki & Wang 1984) [=Xylobates tenuis] S04
    |--P. (P.) triangularis (Hammer 1971) [=Xylobates triangularis] S04
    |--P. (P.) varisetiger (Wen, Aoki & Wang 1984) [=Xylobates varisetiger] S04
    `--P. (P.) yezoensis (Fujikawa 1983) [=Xylobates yezoensis] S04

Protoribates Berlese 1908 (Protoribates) [incl. Alloribates Banks 1947, Propescheloribates Jacot 1936, Styloribates Jacot 1934, Xylobates Jacot 1929] S04

Protoribates (Protoribates) capucinus capucinus Berlese 1908 S04 [incl. Xylobates capucinus angustior Jacot 1937 S04, *Alloribates singularis Banks 1947 CH98, S04]

*Protoribates (Protoribates) dentatus (Berlese 1883) S04 [=Oribata dentata S04, Pr. (Scheloribates) dentatus B16; incl. Oribata monodactyla Haller 1884 S04, Oribates monodactylus S59, Protoribates monodactylus B61, Xylobates monodactylus C-R79, Peloribates vastus Mihelčič 1956 S04]

Protoribates (Protoribates) lophothrichus (Berlese 1904) S04 [=Oribata lophothricha S04, Oribates lophotrichus CH98, *Xylobates lophotrichus CH98; incl. *Styloribates pectinatus Jacot 1934 CH98, S04, Scheloribates pectinatus P63, Xylobates prionotus Woolley 1968 S04]

Protoribates (Protoribates) robustior (Jacot 1937) [=Xylobates oblongus robustior; incl. X. capucinus robustior Jacot 1937 non X. oblongus robustior Jacot 1937] S04

*Type species of generic name indicated

REFERENCES

[B61] Balogh, J. 1961. Identification keys of world oribatid (Acari) families and genera. Acta Zoologica Hungarica 1: 243–344.

[BB92] Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 2. Plates. Hungarian Natural History Museum: Budapest.

[B16] Berlese, A. 1916. Centuria terza di Acari nuovi. Redia 12: 289–338.

[CH98] Colloff, M. J., & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

[C-R79] Corpuz-Raros, L. A. 1979. Philippine Oribatei (Acarina). I. Preliminary list of species and descriptions of forty new species. Philippine Agriculturist 62 (1): 1–82.

[P63] Pletzen, R. van. 1963. Studies on South African Oribatei (Acarina). Acarologia 5 (4): 690–703.

[S59] Sellnick, M. 1959. Acarina from southeastern Polynesia—II (Oribatidae). Occasional Papers of the Bernice P. Bishop Museum 22 (9): 109–152.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Last updated: 19 December 2018.

Haplozetidae

Female Rostrozetes ovulum, copyright B. Eamer.

Belongs within: Oripodoidea.
Contains: Protoribates (Protoribates), Protoribates (Triaunguis), Peloribates (Peloribates), Lauritzenia (Incabates), Indoribates (Haplozetes).

The Haplozetidae are a group of oribatid mites with well-developed, hinged pteromorphs. Members have been divided between separate families Haplozetidae and Xylobatidae or Protoribatidae based on whether the octotaxic system is composed of (respectively) sacculi or areae porosae, but this was not regarded as significant by Norton & Behan-Pelletier (2009). The Oriental species Cribrozetes multiareolatus has the usual four pairs of areae porosae disassembled into 14–18 pairs of areae porosae rougly grouped in the original position. The dorsosejugal suture is absent in the genus Perxylobates (Balogh & Balogh 1992a). Six pairs of genital setae are present in Sicaxylobates, Afroleius, Paraxylobates, Pilobates and Pilobatella; other genera have less.

Characters (from Norton & Behan-Pelletier 2009): Length 250–650µm. Subcapitulum diarthric; chelicerae chelate-dentate. Notogaster with octotaxic system of porose areas or saccules; pteromorphs well developed, with hinge; insertions of adductor muscles for pteromorphs evident on notogaster; posterior notogastral tectum absent. Podotectum II present; circumpedal carina complete. Genital and anal plates separated by more than half length of genital plate; aggenital setae present; postanal porose area absent. Femora III and IV each with one porose area; solenidion σ present on genu III.

<==Haplozetidae [Protoribatidae, Xylobatidae] NB-P09
    |--Paraxylobates Balogh & Mahunka 1969 S04
    |    `--*P. imitans Balogh & Mahunka 1969 S04
    |--Conozetes Balogh & Mahunka 1969 S04
    |    `--*C. arcualis Balogh & Mahunka 1969 S04
    |--Bolkiah Mahunka 1997 S04
    |    `--*B. hauseri Mahunka 1997 S04
    |--Borneozetes Mahunka 1997 S04
    |    `--*B. lanceolatus Mahunka 1997 S04
    |--Cantharozetes Hammer 1961 S04
    |    `--*C. lucens Hammer 1961 S04
    |--Setincabates Lee 1993 S04
    |    `--*S. hypersetosus Lee 1993 S04
    |--Cribrozetes Balogh 1970 S04
    |    `--*C. multiareolatus Balogh 1970 S04
    |--Trixylobates Balogh & Mahunka 1978 S04
    |    `--*T. bidactylus Balogh & Mahunka 1978 S04
    |--Sicaxylobates Luxton 1985 S04
    |    `--*S. sicafer (Hammer 1968) [=Xylobates sicafer] S04
    |--Baloghiella Bulanova-Zachvatkina 1960 S04
    |    |--*B. prima Bulanova-Zachvatkina 1960 S04
    |    `--B. granulata Bayartogtokh & Akrami 2000 S04
    |--Baloghia Mahunka 1994 S04
    |    |--*B. juditae Mahunka 1994 S04
    |    `--B. spinifera Mahunka 1994 S04
    |--Trachyoribates Berlese 1908 S04
    |    |--*T. ampulla (Berlese 1905) [=Oribata ampulla] S04
    |    `--T. chinensis Fukuyama & Aoki 2000 S04
    |--Transoribates Pérez-Íñigo 1992 [incl. Lagenobates Weigmann & Miko 2002] S04
    |    |--T. lagenula (Berlese 1904) (see below for synonymy) S04
    |    `--T. agricola (Nakamura & Aoki 1989) [=Protoribates agricola] S04
    |--Protoribates Berlese 1908 NB-P09, S04
    |    |  i. s.: P. longipilis Sellnick 1931 S93
    |    |--P. (Protoribates) S04
    |    `--P. (Triaunguis) S04
    |--Berlesiella Hammer 1979 S04
    |    |--*B. (Berlesiella) scutata Hammer 1979 S04
    |    `--B. (Comororibula Mahunka 1994) S04
    |         `--B. (*C.) truncata (Mahunka 1994) [=*Comororibula truncata] S04
    |--Afroleius Mahunka 1984 S04
    |    |--*A. deformis Mahunka 1984 S04
    |    |--A. minor Mahunka 1984 S04
    |    `--A. simplex Mahunka 1984 S04
    |--Tuxenia Hammer 1958 S04
    |    |--*T. complicata Hammer 1958 S04
    |    |--T. brevis Covarrubias 1967 [incl. T. brevisetosa Mahunka 1980] S04
    |    `--T. manantialis Hammer 1962 S04
    |--Acutozetes Balogh 1970 S04
    |    |--*A. rostratus Balogh 1970 S04
    |    |--A. bornemisszai Balogh & Balogh 1986 S04
    |    |--A. chibai Aoki 1976 S04
    |    |--A. javensis Hammer 1979 S04
    |    `--A. nadchatrami Balogh & Mahunka 1974 S04
    |--Pilobates Balogh 1960 S04
    |    |--P. (Pilobates) S04
    |    |    |--*P. (P.) pilosellus (Balogh 1958) [=Protoribates pilosellus] S04
    |    |    `--P. (P.) carpentanus Pérez-Íñigo 1969 (see below for synonymy) S04
    |    `--P. (Italobates Mahunka 1994) S04
    |         `--P. (*I.) incisura (Berlese 1916) [=Protoribates (Scheloribates) incisura] S04
    |--Peloribates Berlese 1908 S04
    |    |  i. s.: P. antillensis (Mahunka 1985) S07
    |    |--P. (Peloribates) S04
    |    |--P. (Aokibates Mahunka 1988) S04
    |    |    `--P. (*A.) yoshii (Mahunka 1988) [=*Aokibates yoshii] S04
    |    `--P. (Tentaculozetes Balogh 1970) S04
    |         `--P. (*T.) loksai (Balogh 1970) [=*Tentaculozetes loksai] S04
    |--Pilobatella Balogh & Mahunka 1967 S04
    |    |--*P. punctulata Balogh & Mahunka 1967 S04
    |    |--P. berlesei Bhattacharya & Banerjee 1979 S04
    |    |--P. maurensis Scull 1985 S04
    |    |--P. monstruosa Mahunka 1986 S04
    |    |--P. pseudovermiseta Corpuz-Raros 1979 S04
    |    |--P. schauenbergi Mahunka 1977 S04
    |    `--P. xena Mahunka 1977 S04
    |--Vilhenabates Balogh 1963 S04
    |    |--V. (Vilhenabates) S04
    |    |    |--*V. (V.) minutus (Balogh 1958) [=Peloribates minutus] S04
    |    |    |--V. (V.) hyalinus Tseng 1984 S04
    |    |    `--V. (V.) simplex Balogh 1970 S04
    |    `--V. (Phalacrozetes Aoki 1965) S04
    |         |--V. (*P.) sinatus (Aoki 1965) [=*Phalacrozetes sinatus] S04
    |         `--V. (P.) similis (Mahunka 1988) [=Phalacrozetes similis] S04
    |--Setoxylobates Balogh & Mahunka 1967 S04
    |    |--*S. (Setoxylobates) foveolatus Balogh & Mahunka 1967 S04
    |    |--S. (Plenoxylobates Hammer 1979) S04
    |    |    |--S. (*P.) ramosus (Hammer 1979) [=*Plenoxylobates ramosus] S04
    |    |    |--S. (P.) curtiseta (Hammer 1979) [=Plenoxylobates curtiseta] S04
    |    |    `--S. (P.) mayuloeus (Corpuz-Raros 1979) [=Plenoxylobates mayuloeus] S04
    |    `--S. (Polyxylobates Hammer 1973) S04
    |         |--S. (*P.) diversiporosus (Hammer 1973) [=*Polyxylobates diversiporosus] S04
    |         `--S. (P.) principalis (Berlese 1916) [=Protoribates principalis] S04
    |--Lauritzenia Hammer 1958 S04
    |    |--L. (Lauritzenia) S04
    |    |    |--*L. (L.) longipluma Hammer 1958 S04
    |    |    |--L. (L.) furtadoi (Balogh & Mahunka 1974) [=Haplozetes furtadoi] S04
    |    |    |--L. (L.) glabra Pérez-Íñigo & Baggio 1996 S04
    |    |    |--L. (L.) insignis (Balogh & Mahunka 1966) [=Haplozetes insignis] S04
    |    |    |--L. (L.) loongchiensis (Tseng 1984) [=Haplozetes loongchiensis] S04
    |    |    `--L. (L.) minimicoma (Beck 1964) [=Haplozetes minimicoma] S04
    |    |--L. (Incabates) S04
    |    `--L. (Magnobates Hammer 1967) [incl. Gymnozetes Calugar & Vasiliu 1983] S04
    |         |--L. (*M.) flagellifer (Hammer 1967) [=*Magnobates flagellifer] S04
    |         `--L. (M.) orghidani (Calugar & Vasiliu 1983) S04 [=*Gymnozetes orghidani BB92a]
    |--Perxylobates Hammer 1972 [incl. Octodurozetes Mahunka 1993] S04
    |    |--*P. vermiseta (Balogh & Mahunka 1968) [=Xylobates vermiseta] S04
    |    |--P. barbatus Hammer 1972 S04
    |    |--P. berndhauseri (Mahunka 1993) [=Octodurozetes berndhauseri] S04
    |    |--P. brevisetus Mahunka 1988 S04
    |    |--P. coreanus Coi & Aoki 1993 S04
    |    |--P. guehoi Mahunka 1978 S04
    |    |--P. longissimus (Warburton 1912) [=Oribata longissima] S04
    |    |--P. mahunkai Bayoumi 1980 S04
    |    |--P. mayuloeus Corpuz-Raros 1979 C-R79
    |    |--P. obesus Bayoumi 1977 S04
    |    |--P. paravermiseta Mahunka 1976 S04
    |    |--P. sinensis Wen, Aoki & Wang 1984 S04
    |    |--P. sinlimes (Hammer 1971) C-R79
    |    |--P. taidinchani Mahunka 1976 S04
    |    `--P. vietnamensis (Jeleva & Vu 1987) [=Xylobates vietnamensis] S04
    |--Magyaria Balogh 1963 S04
    |    |--*M. reticulata (Balogh 1958) [=Scheloribates reticulatus] S04
    |    |--M. agusana Corpuz-Raros 1991 S04
    |    |--M. annobonica Pérez-Íñigo 1982 S04
    |    |--M. atlantica Pérez-Íñigo 1983 S04
    |    |--M. breviseta Mahunka 1978 BB92b
    |    |--M. cancellata (Beck 1964) [=Haplozetes cancellatus] S04
    |    |--M. fenestrata Balogh & Mahunka 1974 S04
    |    |--M. filipina Corpuz-Raros 1979 S04
    |    |--M. florida Mahunka 1985 S04
    |    |--M. incisa Balogh & Mahunka 1974 S04
    |    |--M. javensis Hammer 1979 S04
    |    |--M. luzonica Corpuz-Raros 1991 S04
    |    |--M. mindanensis Corpuz-Raros 1979 S04
    |    |--M. ornata Balogh 1963 S04
    |    |--M. pulcherrima Balogh 1970 S04
    |    |--M. strinovichi Balogh 1970 S04
    |    `--M. triungulata Mahunka 1988 S04
    |--Indoribates Jacot 1929 S04
    |    |--I. (Indoribates) [incl. Nixozetes Mahunka 1977, Sundazetes Hammer 1979] S04
    |    |    |--*I. (I.) punctulatus (Sellnick 1925) S04 (see below for synonymy)
    |    |    |--I. (I.) crassisetiger (Fukuyama & Aoki 2000) [=Nixozetes crassisetiger] S04
    |    |    |--I. (I.) gregoryi (Balogh 1970) S04 [=Cosmobates gregoryi S04, Nixozetes gregoryi BB92b]
    |    |    |--I. (I.) japonicus (Aoki 1988) [=Sundazetes japonicus] S04
    |    |    |--I. (I.) javanus (Mahunka 1977) S04 [=*Nixozetes javanus BB92a]
    |    |    |--I. (I.) multisetus (Wen & Zhao 1994) [=Sundazetes multisetus] S04
    |    |    |--I. (I.) nobilis (Golosova 1984) [=Cosmobates nobilis] S04
    |    |    `--I. (I.) panabokkei (Balogh 1970) (see below for synonymy) S04
    |    |--I. (Haplozetes) S04
    |    |--I. (Mancoribates Hammer 1961) [incl. Minasbates Pérez-Íñigo & Baggio 1996] S04
    |    |    |--I. (*M.) rostropilosus (Hammer 1961) [=*Mancoribates rostropilosus] S04
    |    |    |--I. (M.) acutirostrum (Hammer 1968) [=Lauritzenia acutirostrum] S04
    |    |    |--I. (M.) rotundirostrum (Hammer 1968) [=Lauritzenia rotundirostrum] S04
    |    |    `--I. (M.) sphaeroclava (Pérez-Íñigo & Baggio 1996) [=Minasbates sphaeroclava] S04
    |    |--I. (Philippizetes Corpuz-Raros 1979) S04
    |    |    |--I. (*P.) philippinensis (Corpuz-Raros 1979) [=Nixozetes (*P.) philippinensis] S04
    |    |    `--I. (P.) corpusrarosae (Mahunka 1987) [=Nixozetes (P.) corpusrarosae] S04
    |    `--I. (Triungulozetes Subías 2001) [incl. Mixobates Gil-Martín & Subías 1993 non Thor 1905] S04
    |         |--I. (*T.) triangulatus (Beck 1964) [=Haplozetes triangulatus] S04
    |         `--I. (T.) fusifer (Berlese 1908) (see below for synonymy) S04
    `--Rostrozetes Sellnick 1925 NB-P09, S04 [incl. Carabozetes Mihelčič 1957 S04, Zaherizetes Yousef & Nasr 1976 S04]
         |--R. ovulum (Berlese 1908) [=Trachyoribates (Rostrozetes) ovulum) S04
         |    |--R. o. ovulum S04 (see below for synonymy)
         |    |--R. o. appalachicola Jacot 1938 (see below for synonymy) S04
         |    `--R. o. poensis (Mihelčič 1957) [=Carabozetes poensis, Trachyoribates (Rostrozetes) ovulum poensis] S04
         |--R. aegypticus (Yousef & Nasr 1976) [=Zaherizetes aegypticus, Trachyoribates (Rostrozetes) aegypticus] S04
         |--R. africanus (Yousef & Nasr 1976) [=Zaherizetes africanus, Trachyoribates (Rostrozetes) africanus] S04
         |--R. angulifer Balogh & Mahunka 1979 [=Trachyoribates (Rostrozetes) angulifer] S04
         |--R. bothulifer Balogh & Mahunka 1979 [=Trachyoribates (Rostrozetes) bothulifer] S04
         |--R. carinatus Beck 1965 [=Trachyoribates (Rostrozetes) carinatus] S04
         |--R. cristatus Balogh & Mahunka 1969 [=Trachyoribates (Rostrozetes) cristatus] S04
         |--R. cubanus Balogh & Mahunka 1974 [=Trachyoribates (Rostrozetes) cubanus] S04
         |--R. dimorphichaites Higgins 1996 [=Trachyoribates (Rostrozetes) dimorphichaites] S04
         |--R. dispar (Balogh 1958) [=Peloribates dispar, Trachyoribates (Rostrozetes) dispar] S04
         |--R. florens Balogh 1970 [=Trachyoribates (Rostrozetes) florens] S04
         |--R. geminosetosus Balogh & Mahunka 1978 [=Trachyoribates (Rostrozetes) geminosetosus] S04
         |--R. glaber Beck 1965 [=Trachyoribates (Rostrozetes) glaber] S04
         |--R. heterotrichus Pérez-Íñigo & Baggio 1991 [=Trachyoribates (Rostrozetes) heterotrichus] S04
         |--R. inornatus Pérez-Íñigo & Baggio 1991 [=Trachyoribates (Rostrozetes) inornatus] S04
         |--R. irregularis Balogh & Mahunka 1969 [=Trachyoribates (Rostrozetes) irregularis] S04
         |--R. komodensis Mahunka 1977 [=Trachyoribates (Rostrozetes) komodensis] S04
         |--R. mammillatus (Warburton 1912) [=Oribata mammillata, Trachyoribates (Rostrozetes) mamillatus] S04
         |--R. monstruosus Balogh & Mahunka 1969 [=Trachyoribates (Rostrozetes) monstruosus] S04
         |--R. nagaii Mahunka 1987 [=Trachyoribates (Rostrozetes) nagaii] S04
         |--R. nebulosus Beck 1965 PL17 (see below for synonymy)
         |--R. perezinigoi Balogh 1995 [=Trachyoribates (Rostrozetes) perezinigoi] S04
         |--R. pinguis Balogh & Mahunka 1978 [=Trachyoribates (Rostrozetes) pinguis] S04
         |--R. polygonatus Balogh & Mahunka 1969 [=Trachyoribates (Rostrozetes) polygonatus] S04
         |--R. pseudofurcatus Balogh & Mahunka 1968 [=Trachyoribates (Rostrozetes) pseudofurcatus] S04
         |--R. punctatus Karppinen 1966 [=Trachyoribates (Rostrozetes) punctatus] S04
         |--R. rimachensis Beck 1965 [=Trachyoribates (Rostrozetes) rimachensis] S04
         |--R. schalleri Beck 1965 [=Trachyoribates (Rostrozetes) schalleri] S04
         |--R. sexclavatus Sellnick 1925 [=Trachyoribates (Rostrozetes) sexclavatus] S04
         `--R. shibai Aoki 1976 [=Trachyoribates (Rostrozetes) shibai] S04

Indoribates (Indoribates) panabokkei (Balogh 1970) [=Cosmobates panabokkei; incl Nixozetes apoensis Corpuz-Raros 1979] S04

*Indoribates (Indoribates) punctulatus (Sellnick 1925) S04 [=Protoribates punctulatus S04; incl. Sundazetes crispus Hammer 1979 S04, Indoribates crispus BB92b]

Indoribates (Triungulozetes) fusifer (Berlese 1908) [=Protoribates (Scheloribates) fusifer; incl. Haplozetes (Mixobates) insularis Pérez-Íñigo & Peña 1996] S04

Pilobates (Pilobates) carpentanus Pérez-Íñigo 1969 [incl. Protoribates elongatus Mihelčič 1956 (n. d.), Pr. longior Mihelčič 1957 (n. d.) non Berlese 1908] S04

Rostrozetes nebulosus Beck 1965 PL17 [=Rostrozetes foveolatus nebulosus S04; incl. R. nebulosus tropica Beck 1963 (n. n.) B65]

Rostrozetes ovulum appalachicola Jacot 1938 [=R. foveolatus appalachicola, Trachyoribates (Rostrozetes) ovulum appalachicola] S04

Rostrozetes ovulum ovulum (Berlese 1908) S04 [incl. Peloribates areolatus Balogh 1958 S04, Trachyoribates areolatus B65, T. dorsalis Balogh 1958 S04, Rostrozetes flavus Woodring 1965 S04, *R. foveolatus Sellnick 1925 S04, R. geneuxi Mahunka 1985 S04, Trachyoribates nodosus Hammer 1958 S04, R. nodosus B65, R. pulcherrimus Balogh 1961 S04, Peloribates punctulatus Balogh 1958 S04, Trachyoribates punctulatus B65, R. punctulifer Balogh & Mahunka 1979 S04, R. trimorphus Balogh & Mahunka 1979 S04]

Transoribates lagenula (Berlese 1904) [=Oribata lagenula; incl. Protoribates latus Mihelčič 1965, *Transoribates latus, P. pupula Berlese 1910] S04

*Type species of generic name indicated

REFERENCES

[BB92a] Balogh, J., & P. Balogh. 1992a. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[BB92b] Balogh, J., & P. Balogh. 1992b. The Oribatid Mites Genera of the World vol. 2. Plates. Hungarian Natural History Museum: Budapest.

[B65] Beck, L. 1965. Über Variabilität und Wertigkeit morphologischer Merkmale bei adulten Oribatiden (Arachnida, Acari) am Beispiel der Gattung Rostrozetes Sellnick 1925. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft 508: 1–64.

[C-R79] Corpuz-Raros, L. A. 1979. Philippine Oribatei (Acarina). I. Preliminary list of species and descriptions of forty new species. Philippine Agriculturist 62 (1): 1–82.

[NB-P09] Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3^rd ed. pp. 430–564. Texas Tech University Press.

[PL17] Pachl, P., A. C. Lindl, A. Krause, S. Scheu, I. Schaefer & M. Maraun. 2017. The tropics as an ancient cradle of oribatid mite diversity. Acarologia 57 (2): 309–322.

[S07] Schatz, H. 2007. Biogeography of oribatid mites (Acari, Oribatida) from the Cordillera de Talamanca, Costa Rica and Panama. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 151–167. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[S93] Selden, P. A. 1993. Arthropoda (Aglaspidida, Pycnogonida and Chelicerata). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 297–320. Chapman & Hall: London.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Last updated: 24 February 2019.

Paraphauloppia (Paraphauloppia)

Dorsal (left) and ventral views of Paraphauloppia novaezealandica, from Balogh & Balogh (1992 vol. 2).

Belongs within: Oribatulidae.

The type subgenus of the genus Paraphauloppia is a group of oribatulid mites found in Australasia and the Americas (Subías 2004).

Characters (from Balogh & Balogh 1992): Prodorsum with well developed lamellae; translamella absent; sublamellar line absent. Dorsosejugal suture medially interrupted; basal part of dorsosejugal suture directed inward, almost horizontal. Notogaster with ten pairs of setae; four pairs of areae porosae; pteromorphae absent. Genital plates with three pairs of extremely small alveoli; one pair of aggenital setae; two pairs of anal setae; three pairs of adanal setae. Legs tridactyle.

<==Paraphauloppia Hammer 1967 (Paraphauloppia) S04
    |--*P. (P.) novaezealandica Hammer 1967 S04
    |--P. (P.) acutinotata Lee & Birchby 1991 S04
    |--P. (P.) altimontana (Hammer 1958) [=Oribatula altimontana] S04
    |--P. (P.) altimontanoides (Hammer 1958) [=Oribatula altimontanoides] S04
    |--P. (P.) australis (Hammer 1962) [=Eporibatula australis] S04
    |--P. (P.) cordylinosa Higgins & Woolley 1975 S04
    |--P. (P.) globata Lee & Birchby 1991 S04
    |--P. (P.) gracilis (Hammer 1958) [=Eporibatula gracilis] S04
    |--P. (P.) magniporosa (Hammer 1958) [=Oribatula magniporosa] S04
    |--P. (P.) morenoi (Hammer 1962) [=Oribatula morenoi] S04
    |--P. (P.) obtusinotata Lee & Birchby 1991 S04
    `--P. (P.) triforata Lee & Birchby 1991 S04

*Type species of generic name indicated

REFERENCES

Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Oribatula (Zygoribatula)

Oribatula exilis, copyright R. Penttinen.

Belongs within: Oribatulidae.

The subgenus Zygoribatula of the genus Oribatula is a cosmopolitan group of oribatulid mites, typically distinguished from the closely allied Oribatula by the presence of a translamella connecting the prodorsal lamellae. However, the reliability of this feature has been called into question (and its presence may vary between individuals of a single species) so the composition of Zygoribatula requires re-investigation. The genus Fovoribatula, previously recognised as a distinct genus for two Australian species with reduced leg chaetotaxy, was not accepted as distinct by Subías (2004), albeit the latter did not explain his reasons for this decision.

Characters (from Lee 1992): Proteronotum with lamellate translamella and lamella, subparallel pair of lamellae, medium size to long, distance between lamellar seta (z1) and dorsosejugal furrow greater than 1.5 times distance between interlamellar seta j2 and bothridium of sensory seta z2. Thirteen or fourteen pairs of short to medium, smooth or weakly ciliate hysteronotal setae; humeral setae (Z1, S1) similar in size and shape to other hysteronotal setae (rarely Z1 stouter and more ciliate, but still subequal in length or smaller). Genital shield with four pairs of setae.

<==Oribatula (Zygoribatula Berlese 1916) (see below for synonymy) S04
    |--O. (*Z.) connexa Berlese 1904 S04
    |    |--O. c. connexa (see below for synonymy) S04
    |    |--O. c. substriata (Grobler & Kok 1993) [=Zyoribatula substriata] S04
    |    `--O. c. ucrainica (Iordansky 1990) [=Zygoribatula terricola ucrainica] S04
    |--O. (Z.) acuminata Wallwork 1964 S04
    |    |--O. a. acuminata S04
    |    `--O. a. mariana Aoki 1994 S04
    |--O. (Z.) agaveae (Aoki & Wang 1986) [=Zygoribatula agaveae] S04
    |--O. (Z.) andrianovae (Bulanova-Zachvatkina 1967) [=Zygoribatula andrianovae] S04
    |--O. (Z.) arcuata (Hammer 1977) [=Zygoribatula arcuata] S04
    |--O. (Z.) arcuatissima Berlese 1916 (see below for synonymy) S04
    |--O. (Z.) baloghi (Mahunka 1986) (see below for synonymy) S04
    |--O. (Z.) bonairensis (Willmann 1936) [=Zygoribatula bonairensis] S04
    |    |--O. b. bonairensis [incl. Zygoribatula heterotricha Mahunka 1979] S04
    |    `--O. b. chilensis (Covarrubias & Toro 1997) [=Zygoribatula bonairensis chilensis] S04
    |--O. (Z.) brevisetosa (Ewing 1909) S04 [=Notaspis brevisetosa S04, *Neoribatula brevisetosa CH98]
    |--O. (Z.) capitata (Banks 1910) [=Liacarus capitatus] S04
    |--O. (Z.) clavata Ewing 1917 S04
    |--O. (Z.) cognata (Oudemans 1902) (see below for synonymy) S04
    |--O. (Z.) concolor (Banks 1895) [=Scutovertex concolor] S04
    |--O. (Z.) contracta (Grobler 1994) [=Zygoribatula contracta] S04
    |--O. (Z.) cycloporosa (Lee 1992) S04 [=Zygoribatula cycloporosa L92]
    |--O. (Z.) dactylaris (Subías, Ruiz & Kahwash 1990) S04 [=Zygoribatula dactylaris S04, Phauloppiella dactilaris L92]
    |--O. (Z.) debilitranslamellata (Kulijev 1962) [=Zygoribatula debilitranslamellata] S04
    |--O. (Z.) dentata (Balogh 1958) (n. d.) [=Zygoribatula dentata] S04
    |--O. (Z.) egelida (Tseng 1984) [=Zygoribatula egelida] S04
    |--O. (Z.) endroedyi (Mahunka 1986) [=Zygoribatula endroedyi] S04
    |--O. (Z.) eucalla (Wen & Zhang 1988) [=Zygoribatula eucalla] S04
    |--O. (Z.) exarata Berlese 1916 (see below for synonymy) S04
    |--O. (Z.) exilis (Nicolet 1855) S04 (see below for synonymy)
    |--O. (Z.) floridana (Fritz 1982) [=Zygoribatula floridana] S04
    |--O. (Z.) frisiae (Oudemans 1900) S04 (see below for synonymy)
    |--O. (Z.) fusca (Ewing 1913) [=Notaspis pyrostigmata fusca; incl. Zygoribatula rostrata Jacot 1938] S04
    |--O. (Z.) galula (Mahunka 2001) [=Zygoribatula galula] S04
    |--O. (Z.) glabra (Michael 1890) (see below for synonymy) S04
    |--O. (Z.) globosa (Mahunka 2000) [=Zygoribatula globosa] S04
    |--O. (Z.) gracilata (Grobler & Kok 1993) [=Zygoribatula gracilata] S04
    |--O. (Z.) grandjeani (Nawar & Borolossy 1990) [=Zygoribatula grandjeani] S04
    |--O. (Z.) granulata (Kunst 1958) [=Zygoribatula granulata] S04
    |--O. (Z.) gratiosa (Tseng 1984) [=Zygoribatula gratiosa] S04
    |--O. (Z.) guadarramica (Pérez-Íñigo 1978) [=Zygoribatula guadarramica] S04
    |--O. (Z.) hailongensis (Wen & Zhang 1988) [=Zygoribatula hailongensis] S04
    |--O. (Z.) heterochaeta (Feider, Vasiliu & Calugar 1970) [=Zygoribatula heterochaeta] S04
    |--O. (Z.) hispanica Subías & Arillo 1998 S04
    |--O. (Z.) kelbadjarica (Kulijev 1962) [=Zygoribatula kelbadjarica] S04
    |--O. (Z.) keralensis (Ramani & Haq 1999) [=Zygoribatula keralensis] S04
    |--O. (Z.) knighti (Luxton 1987) [=Zygoribatula knighti] S04
    |--O. (Z.) koki (Balogh & Balogh 2002) [=Zygoribatula koki] S04
    |--O. (Z.) lata (Hammer 1961) [=Zygoribatula lata; incl. Z. elongata Hammer 1961, Z. oblonga Hammer 1961] S04
    |--O. (Z.) lenticulata (Mínguez & Subías 1986) [=Zygoribatula lenticulata] S04
    |--O. (Z.) levigata (Wen & Bu 1988) [=Zygoribatula levigata] S04
    |--O. (Z.) lineaporosa (Grobler 1993) [=Zygoribatula lineaporosa] S04
    |--O. (Z.) lineata (Hammer 1979) [=Zygoribatula lineata] S04
    |--O. (Z.) longicuspis (Balogh 1966) [=Zygoribatula longicuspis] S04
    |--O. (Z.) longilinea (Wang & Li 1997) [=Zygoribatula longilinea] S04
    |--O. (Z.) longisensilla (Djaparidze 1985) [=Zygoribatula longisensilla] S04
    |--O. (Z.) longiseta (Golosova 1970) [=Zygoribatula longiseta] S04
    |--O. (Z.) mabar (Mahunka 2000) [=Zygoribatula mabar] S04
    |--O. (Z.) mariehammerae (Feider, Vasiliu & Calugar 1970) [=Zygoribatula mariehammerae] S04
    |--O. (Z.) marina (Fujikawa 1972) [=Zygoribatula marina] S04
    |--O. (Z.) meruensis (Mahunka 1969) [=Zygoribatula meruensis] S04
    |--O. (Z.) microporosa (Bulanova-Zachvatkina 1967) [=Zygoribatula microporosa] S04
    |--O. (Z.) niliaca (Bayoumi 1980) [=Zygoribatula niliaca] S04
    |--O. (Z.) novazealandica (Hammer 1967) [=Zygoribatula novazealandica] S04
    |--O. (Z.) obsessa Subías 2004 S04 (see below for synonymy)
    |--O. (Z.) oceana (Hammer 1972) [=Zygoribatula oceana] S04
    |--O. (Z.) pennata (Grobler 1993) [=Zygoribatula pennata] S04
    |--O. (Z.) prodorsissima (Feider, Vasiliu & Calugar 1970) [=Zygoribatula prodorsissima] S04
    |--O. (Z.) ‘robusta’ (Grobler 1993) non Iordansky 1991 [=Zygoribatula robusta] S04
    |--O. (Z.) ruchljadevi (Bulanova-Zachvatkina 1967) [=Zygoribatula ruchljadevi] S04
    |--O. (Z.) sabulosa (Balogh 1966) [=Zygoribatula sabulosa] S04
    |--O. (Z.) salina (Balogh 1966) [=Zygoribatula salina] S04
    |--O. (Z.) sayedi (Elbadry & Nasr 1974) [=Zygoribatula sayedi] S04
    |--O. (Z.) schauenbergi (Mahunka 1978) [=Zygoribatula schauenbergi] S04
    |--O. (Z.) sculpturata (Mahunka 1989) [=Phauloppia sculpturata] S04
    |--O. (Z.) setosa Evans 1953 S04
    |--O. (Z.) skrjabini (Bulanova-Zachvatkina 1967) [=Zygoribatula skrjabini] S04
    |--O. (Z.) smirnovi (Bulanova-Zachvatkina 1978) [=Zygoribatula smirnovi] S04
    |--O. (Z.) socotrensis (Mahunka 2000) [=Zygoribatula socotrensis] S04
    |--O. (Z.) spherisensilla (Djaparidze 1985) [=Zygoribatula spherisensilla] S04
    |--O. (Z.) subantarctica (Pletzen & Kok 1971) [=Zygoribatula subantarctica] S04
    |--O. (Z.) talis (Tseng 1984) [=Zygoribatula talis] S04
    |--O. (Z.) tameyei (Elbadry & Nasr 1974) [=Zygoribatula tameyei] S04
    |--O. (Z.) tenuiseta (Hammer 1977) [=Zygoribatula tenuiseta] S04
    |--O. (Z.) thalassophila Grandjean 1935 [incl. O. travei Bernini 1969] S04
    |--O. (Z.) tortilis (Hammer 1977) [=Zygoribatula tortilis] S04
    |--O. (Z.) trigonella (Bulanova-Zachvatkina 1967) [=Zygoribatula trigonella] S04
    |--O. (Z.) tritici (Elbadry & Nasr 1974) [=Zygoribatula tritici] S04
    |--O. (Z.) truncata (Aoki 1961) [=Zygoribatula truncata] S04
    |--O. (Z.) undulata Berlese 1916 (see below for synonymy) S04
    |--O. (Z.) vanharteni (Mahunka 2000) [=Zygoribatula vanharteni] S04
    |--O. (Z.) vegeta (Tseng 1984) [=Zygoribatula vegeta] S04
    |--O. (Z.) vulgaris (Bulanova-Zachvatkina 1967) [=Zygoribatula vulgaris] S04
    `--O. (Z.) zicsii (Bayoumi 1979) [=Zygoribatula zicsii] S04

Oribatula (Zygoribatula Berlese 1916) [incl. Fovoribatula Lee & Birchby 1991, Neoribatula Ewing 1917, Zetobelba Hull 1916] S04

Oribatula (Zygoribatula) arcuatissima Berlese 1916 [incl. Notaspis aequalis Michael 1890 (n. d.), Zygoribatula gozmanyi Mahunka 1980] S04

Oribatula (Zygoribatula) baloghi (Mahunka 1986) [=Zygoribatula baloghi; incl. Z. undulata Balogh 1966 non O. undulata Berlese 1916] S04

Oribatula (Zygoribatula) cognata (Oudemans 1902) [=Eremaeus cognatus; incl. Zygoribatula diversa Mihelčič 1956, Oribatula (Zygoribatula) socia Berlese 1916] S04

Oribatula (Zygoribatula) connexa connexa Berlese 1904 [incl. Zygoribatula concinna Iordansky 1990, O. (Z.) excavata Berlese 1916, Z. magna Ramsay 1966, Z. striatissima Hammer 1962, Z. terricola Hammen 1952, Z. trichosa Mihelčič 1956] S04

Oribatula (Zygoribatula) exarata Berlese 1916 [incl. O. rugifrons Sellnick 1943, O. rugosa Kulijev 1979, O. rugifrons striata Mihelčič 1955] S04

Oribatula (Zygoribatula) exilis (Nicolet 1855) S04 [=Notaspis exilis S04, Oppia exilis CF77; incl. Zygoribatula clavatotrichobothria Kulijev 1962 S04, Z. extremodentata Kulijev 1962 S04, Dameosoma jugorum Hull 1914 S04, *Zetobelba jugorum CH98, Oribatula (Zygoribatula) lineola Berlese 1916 S04, Notaspis exilis var. maculata Schweizer 1922 S22]

Oribatula (Zygoribatula) frisiae (Oudemans 1900) S04 [=Eremaeus frisiae S04; incl. Zygoribatula frisiae insularis Travé 1961 S04, Notaspis pyrostigmata Ewing 1909 S04, Z. tenuelamellata Mihelčič 1956 S04, Oribatula variabilis Berlese 1908 S04, O. (Zygoribatula) variabilis B16]

Oribatula (Zygoribatula) glabra (Michael 1890) [=Notaspis glabra; incl. O. (Z.) angulata Berlese 1916, Eremaeus brauni Sellnick 1908, Zygoribatula laubieri canariensis Pérez-Íñigo & Peña 1997, Z. hortobagyensis Mahunka 1983, Z. laubieri Travé 1961, Z. matritensis Mihelčič 1966, Z. laubieri meridionalis Travé 1961, Eremaeus propinquus Oudemans 1902] S04

Oribatula (Zygoribatula) obsessa Subías 2004 S04 [=*Fovoribatula brevisetosa Lee & Birchby 1991 non Notaspis brevisetosa Ewing 1909 CH98]

Oribatula (Zygoribatula) undulata Berlese 1916 [incl. Zygoribatula dubita Coetzer 1968, Z. heteroporosa Wallwork 1972, Z. longiporosa Hammer 1953, Z. tadrosi Popp 1960] S04

*Type species of generic name indicated

REFERENCES

[B16] Berlese, A. 1916. Centuria terza di Acari nuovi. Redia 12: 289–338.

[CF77] Canestrini, G., & F. Fanzago. 1877. Intorno agli Acari Italiani. Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti, Serie 5, 4: 69–208, pls 2–7.

[CH98] Colloff, M. J., & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

[L92] Lee, D. C. 1992. New species of Oribatulidae (Acarida: Cryptostigmata: Planofissurae) from South Australian soils, with a review of subfamilies and Australian records. Records of the South Australian Museum 26 (1): 37–49.

[S22] Schweizer, J. 1922. Beitrag zur Kenntnis der terrestrischen Milbenfauna der Schweiz. Verhandl. Naturf. Ges. Basel 33: 23–112, 4 pls.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Last updated: 16 December 2018.

Lamellobates

Anterior dorsal view of Lamellobates reticulatus, copyright B. Eamer.

Belongs within: Ceratozetoidea.

Lamellobates is a pantropical genus of oribatid mites of uncertain affinities.

Characters (from Balogh & Balogh 1992 vol. 1): Prodorsum covered by two large lamellae. Notogaster with ten pairs of setae; four pairs of sacculi; pteromorphae immovable. Six pairs of genital setae; one pair of aggenital setae; two pairs of anal setae; two pairs of adanal setae (ad2 and ad3). Legs monodactyle.

<==Lamellobates Hammer 1958 [Lamellobatidae] S04
    |--L. (Lamellobates) S04
    |    |--L. (L.) molecula (Berlese 1916) [=Achipteria molecula] S04
    |    |    |--L. m. molecula (see below for synonymy) S04
    |    |    `--L. m. aegypticus Bayoumi 1979 [=L. hauseri aegypticus] S04
    |    |--L. (L.) botari Balogh & Mahunka 1977 S04
    |    |--L. (L.) engelbrechti Mahunka 1989 S04
    |    |--L. (L.) gyoergyi Balogh & Mahunka 1977 S04
    |    |--L. (L.) intermedius Nübel-Reidelbach & Woas 1992 S04
    |    |--L. (L.) nitidus Calugar 1987 S04
    |    |--L. (L.) ocularis Jeleva & Vu 1987 S04
    |    |--L. (L.) orientalis Csiszár 1961 S04
    |    |--L. (L.) quadricornis Pérez-Íñigo & Baggio 1985 S04
    |    `--L. (L.) reticulatus Behan-Pelletier 1998 S04
    `--L. (Paralamellobates Bhaduri & Raychaudhuri 1968) S04
         |--L. (*P.) bengalensis Bhaduri & Raychaudhuri 1968 S04
         |--L. (P.) misella (Berlese 1910) S04 (see below for synonymy)
         `--L. (P.) striatus (Behan-Pelletier 1998) [=Paralamellobates striatus] S04

Lamellobates (Lamellobates) molecula molecula (Berlese 1916) [incl. L. angolensis Balogh 1958, L. hauseri Mahunka 1977, *L. palustris Hammer 1958, L. rotundatus Scull, Jeleva & Cruz 1984] S04

Lamellobates (Paralamellobates) misella (Berlese 1910) S04 [=Oribatella misella S04; incl. O. ceylanica Oudemans 1915 S04, O. schoutedeni Balogh 1959 S04, Paralamellobates schoutedeni BB92]

*Type species of generic name indicated

REFERENCES

[BB92] Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 2. Plates. Hungarian Natural History Museum: Budapest.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.