Carabodes (Carabodes)

Carabodes labyrinthicus, copyright Biodiversity Institute of Ontario.

Belongs within: Carabodinae.

The type subgenus of the genus Carabodes is a subcosmopolitan group of carabodid oribatid mites with an evenly arched notogaster. The type species, C. coriaceus, is widespread in forest soils in Europe with less certain records from other parts of the Holarctic region.

Characters (from Balogh & Balogh 1992): Prodorsum and notogaster without high elevation(s) or deep excavation(s); notogaster with ten pairs of well-developed setae. Ventral region not shortened, genital and anal plates normally separated. Genital plates with four setae; aggenital setae usually present.

Carabodes Koch 1835 (Carabodes) [incl. Neocepheus Willmann 1936] S04
    |--C. (C.) marginatus (Michael 1884) SK10, S04 [=Tegeocranus marginatus S04; incl. C. pontiger Berlese 1913 S04]
    `--+--C. femoralis group CA86
       |    |--C. (C.) femoralis (Nicolet 1855) SK10, S04 [=Tegeocranus femoralis S04]
       |    |--C. (C.) rimosus Aoki 1959 CA86
       |    |--C. (C.) rugosior Berlese 1916 CA86, S04 [=C. femoralis var. rugosior S04]
       |    |--C. (C.) transversarius Choi & Aoki 1986 CA86
       |    `--C. (C.) tsushimaensis Aoki 1970 CA86, S04 [=C. rimosus tsushimaensis S04]
       |         |--C. t. tsushimaensis S04
       |         `--C. t. ‘dorsalis’ Choi & Aoki 1986 non C. dorsalis Banks 1896 S04
       `--C. (C.) labyrinthicus (Michael 1879) SK10, S04 (see below for synonymy)

Carabodes (Carabodes) incertae sedis:
  *C. (C.) coriaceus Koch 1835 [incl. C. nepos Hull 1914, C. papillosus Lucas 1849] S04
  C. (C.) affinis Berlese 1913 S04
  C. (C.) apuanicus Bernini 1979 S04
  C. (C.) arduinii Valle 1955 S04
  C. (C.) areolatus Berlese 1916 S04
  C. (C.) auriculatus Mahunka 1987 S04
  C. (C.) azoricus Pérez-Íñigo & Pérez-Íñigo 1996 S04
  C. (C.) basilewskyi Balogh 1958 (n. d.) S04
  C. (C.) bellus Aoki 1959 S04
  C. (C.) berninii Mahunka 1983 S04
  C. (C.) bicolor Balogh 1958 (n. d.) S04
  C. (C.) brevis Banks 1896 S04
  C. (C.) cephalotes Koch 1836 (n. d.) [incl. C. cynocephalus Koch 1839] S04
  C. (C.) cerebrum Fujikawa 1993 S04
  C. (C.) cherokee Reeves 1993 S04
  C. (C.) chirstlus Mahunka 1987 [incl. C. foliatus Morell 1990] S04
  C. (C.) clavatus Warburton 1912 (n. d.) [=C. lalbyrinthicus clavatus] S04
  C. (C.) cochleaformis Reeves 1990 S04
  C. (C.) colorado Reeves & Behan-Pelletier 1998 S04
  C. (C.) comas Kulijev 1979 S04
  C. (C.) coweetaensis Reeves 1995 S04
  C. (C.) dudichi Mahunka & Mahunka-Papp 1999 S04
  C. (C.) egregius Djaparidze 1990 S04
  C. (C.) erectus Reeves 1992 S04
  C. (C.) falcatus Jacot 1937 S04
  C. (C.) floridus Berlese 1913 S04
  C. (C.) gibbiceps Berlese 1916 S04
  C. (C.) grandjeani Bernini 1977 S04
  C. (C.) granulatus Banks 1895 [incl. C. omo Jacot 1937] S04
  C. (C.) gregorioi Gil-Martín & Subías 1997 S04
  C. (C.) guadarramicus Subías & Arillo 2001 S04
  C. (C.) hispanicus Pérez-Íñigo 1966 S04
  C. (C.) hoh Reeves & Behan-Pelletier 1998 S04
  C. (C.) hummelincki (Willmann 1936) S04 [=*Neocepheus hummelincki CH98]
  C. (C.) hungaricus Balogh 1943 S04
  C. (C.) ikeharai Aoki 1988 S04
  C. (C.) interruptus Reeve 1992 S04
  C. (C.) magnus Kunst 1961 S04
  C. (C.) montanus Bernini 1979 S04
  C. (C.) nantahalaensis Reeves 1993 S04
  C. (C.) neonominatus Subías 2004 (see below for synonymy) S04
  C. (C.) niger Banks 1895 S04
  C. (C.) nitens Johnston 1847 (n. d.) S04
  C. (C.) ornatus Storkan 1925 [incl. C. forsslundi Sellnick 1953] S04
  C. (C.) palmifer Berlese 1904 [incl. C. peniculatus Aoki 1970] S04
  C. (C.) paraspinosus Kulijev 1968 S04
  C. (C.) pentasetosus Reeves 1992 S04
  C. (C.) perezinigoi Salinas 1971 S04
  C. (C.) phylliformis Reeves 1993 S04
  C. (C.) pirenaicus Subías & Arillo 2001 S04
  C. (C.) pirenensis Kunst 1961 S04
  C. (C.) polyporetes Reeves 1991 S04
  C. (C.) prunum Fujikawa 1993 S04
  C. (C.) quadrangulus Bernini 1979 [incl. C. flagellifer Mihelčič 1966] S04
  C. (C.) reticulatus Berlese 1913 [=C. coriaceus reticulatus] S04
  C. (C.) spiniformis Reeves 1995 S04
  C. (C.) spinosus Storkan 1925 S04
  C. (C.) subalpinus Thor 1937 S04
  C. (C.) subarcticus Trägardh 1902 S04 [=C. elongatus var. subarctica S22]
  C. (C.) subcarinatus Mahunka & Mahunka-Papp 2000 (see below for synonymy) S04
  C. (C.) tenuis Forsslund 1953 S04
    |--C. t. tenuis S04
    `--C. t. longisetosus Kulijev 1968 S04
  C. (C.) wettsteini Mahunka 1996 S04
  C. (C.) wonalancetanus Reeves 1990 S04

Carabodes (Carabodes) labyrinthicus (Michael 1879) SK10, S04 [=Tegeocranus labyrinthicus S04; incl. Cepheus heimi Oudemans 1903 S04, Carabodes marginepunctatus Trägardh 1902 S04, Ca. scymnus Hull 1914 S04, Ca. vermiculatus Berlese 1916 S04]

Carabodes (Carabodes) neonominatus Subías 2004 [=C. gibbiceps clavata Jacot 1938 non C. labyrinthicus clavatus Warburton 1912] S04

Carabodes (Carabodes) subcarinatus Mahunka & Mahunka-Papp 2000 [incl. C. carinatus Mahunka & Mahunka-Papp 1999 non Wallwork 1977] S04

*Type species of generic name indicated

REFERENCES

Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[CA86] Choi, S. S., & J. Aoki. 1986. The oribatid mites (Acari: Cryptostigmata) of Korea (4). On the femoralis-group of the genus Carabodes C.L. Koch, 1836. Korean Journal of Entomology 16 (1): 27–32.

[CH98] Colloff, M. J., & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

[S22] Schweizer, J. 1922. Beitrag zur Kenntnis der terrestrischen Milbenfauna der Schweiz. Verhandl. Naturf. Ges. Basel 33: 23–112, 4 pls.

[SK10] Schäffer, S., S. Koblmüller, T. Pfingstl, C. Sturmbauer & G. Krisper. 2010. Ancestral state reconstruction reveals multiple independent evolution of diagnostic morphological characters in the "Higher Oribatida" (Acari), conflicting with current classification. BMC Evolutionary Biology 10: 246.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

Last updated: 15 December 2018.