Neides tipularius, copyright Roger Key.

Belongs within: Panheteroptera.
Contains: Coreidae.

The Coreoidea are a group of plant-feeding bugs, most species of have ocelli and a series of longitudinal accessory veins in the forewing membrane (Carver et al. 1991). Members of this group include the Berytidae, stilt bugs, extremely slender and long-legged bugs characterised by short and spindle-shaped apical segments on the antennae. The Piesmatidae are small bugs with a reduced membrane on the hemelytra, with the pronotum and corium being densely punctate (Carver et al. 1991).

Characters (from Carver et al. 1991): Macropterous or brachypterous; brachypters with antennifers inserted dorsolaterally; ocelli present; membrane of hemelytron in macropters with at least 6 and usually many more longitudinal veins; S6 with 3 or fewer trichobothria on each side.

    |--Piesmatidae [Piesmatoidea] SP02
    |    |--Piesma capitata H01
    |    |--Mcateella interioris G75
    |    `--Miespa CGW91
    |--Colobathristidae SP02
    |    |--Dayakiellinae SP02
    |    `--Phaenacantha [Colobathristinae] D14
    |         |--P. australiae CGW91
    |         `--P. pacifica Horváth 1904 K08
    |--+--Coreidae GE05
    |  `--Hyocephalidae G75
    |       |--Hyocephalus aprugnus G76
    |       `--Maevius indecorus CGW91
    `--Berytidae SP02
         |--Berytus clavipes H01
         |--Protocanthus halei CGW91
         |--Metacanthus pluto G75
         `--Neides H01
              |--N. clavipes [=Gerris clavipes] L02
              |--N. maiponga G75
              |--N. propinquus Horváth 1901 H01
              `--N. tipularius [=Gerris tipularius] L02

Coreoidea incertae sedis:
  Pachymeridiidae SP02
  Idiostolidae [Idiostoloidea] SP02
    |--Idiostolus SS02
    |--Monteithocoris hirsutus CGW91
    `--Trisecus pictus WEE70
  Heterogastridae [Heterogastrinae] SE03
    |--Dinomachus MC13
    |--Woodwardothignus gracilis CGW91
    `--Heterogaster Schilling 1829 SE03
         `--H. urticae (Fabricius 1775) SE03
  Alydidae [Alydinae] SP02
    |--Mirperus curvidens (Montr. 1858) [=Alydus curvidens] D14
    |--Melanacanthus scutellaris MC13
    |--Mutusca brevicornis CGW91
    |--Noliphus CGW91
    |    |--N. erythrocephalus CGW91
    |    `--N. insularis Stal 1873 K08
    |--Leptocorisa CGW91
    |    |--L. acuta CGW91
    |    `--L. oratorius R96
    |--Alydus H01
    |    |--A. calcaratus [incl. A. atratus, A. calcaratus var. hirsutus] H01
    |    |--A. pilosulus H01
    |    `--A. zichyi Horváth 1901 H01
    `--Riptortus WEE70
         |--R. annulicornis K08
         |--R. clavatus TC01
         `--R. serripes ZS10
  Dicranocephalus MC13 [Stenocephalidae CGW91]
  Rhopalidae CGW91
    |--Liorhyssus hyalinus CGW91
    `--Leptocoris [Serinethinae] MC13
         |--L. augur K08
         |--L. insularis Kirkaldy 1908 K08
         |--L. mitellatus G75
         `--L. rufomarginatus CGW91

*Type species of generic name indicated


[CGW91] Carver, M., G. F. Gross & T. E. Woodward. 1991. Hemiptera (bugs, leafhoppers, cicadas, aphids, scale insects etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1 pp. 429–509. Melbourne University Press: Carlton (Victoria).

[D14] Distant, W. L. 1914. Rhynchota from New Caledonia and the surrounding islands. In: Sarasin, F., & J. Roux (eds) Nova Caledonia: Forschungen in Neu-Caledonian und auf den Loyalty-Inseln. A. Zoologie vol. 1 Heft 4 pp. 369–390, pls 11–12. C. W. Kreidels Verlag: Wiesbaden.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[G75] Gross, G. F. 1975. Plant-feeding and Other Bugs (Hemiptera) of South Australia. Heteroptera—Part I. Handbook of the Flora and Fauna of South Australia.

[G76] Gross, G. F. 1976. Plant-feeding and Other Bugs (Hemiptera) of South Australia. Heteroptera—Part II. Handbook of the Flora and Fauna of South Australia.

[H01] Horváth, G. 1901. Hemipterák [Hemiptera]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 245–274. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[K08] Kirkaldy, G. W. 1908. A catalogue of the Hemiptera of Fiji. Proceedings of the Linnean Society of New South Wales 33: 345–391, pl. 4.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[MC13] Majer, J. D., S. K. Callan, K. Edwards, N. R. Gunawardene & C. K. Taylor. 2013. Baseline survey of the terrestrial invertebrate fauna of Barrow Island. Records of the Western Australian Museum Supplement 83: 13–112.

[R96] Rentz, D. 1996. Grasshopper Country: The abundant orthopteroid insects of Australia. University of New South Wales Press: Sydney.

[SE03] Scudder, G. G. E., & A. C. Eyles. 2003. Heterogaster urticae (Hemiptera: Heterogastridae), a new alien species and family to New Zealand. Weta 25: 8–13.

[SP02] Shcherbakov, D. E., & Yu. A. Popov. 2002. Superorder Cimicidea Laicharting, 1781: Order Hemiptera Linné, 1758. The bugs, cicadas, plantlice, scale insects, etc. [=Cimicida Laicharting, 1781, =Homoptera Leach, 1815+Heteroptera Latreille, 1810). In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 143–157. Kluwer Academic Publishers: Dordrecht.

[SS02] Sweet, M. H., & C. W. Schaefer. 2002. Parastrachiinae (Hemiptera: Cydnidae) raised to family level. Annals of the Entomological Society of America 95: 441–448.

[TC01] Taylor, DeM., & Y. Chinzei. 2001. Vitellogenin and its synthesis in the soft ticks. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10th International Congress pp. 622–627. CSIRO Publishing: Melbourne.

[WEE70] Woodward, T. E., J. W. Evans & V. F. Eastop. 1970. Hemiptera (bugs, leafhoppers, etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 387–457. Melbourne University Press.

[ZS10] Zborowski, P., & R. Storey. 2010. A Field Guide to Insects in Australia 3rd ed. Reed New Holland: Sydney.

Last updated: 9 October 2017.

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