Gonocerus juniperi, photographed by Ulrich Maurer.

Belongs within: Panheteroptera.

The Coreoidea are a group of plant-feeding bugs, most species of have ocelli and a series of longitudinal accessory veins in the forewing membrane (Carver et al. 1991). Members of the main family, Coreidae, are commonly known as leaf-footed bugs due to the appearance of their enlarged hind tibiae.

Characters (from Carver et al. 1991): Macropterous or brachypterous; brachypters with antennifers inserted dorsolaterally; ocelli present; membrane of hemelytron in macropters with at least 6 and usually many more longitudinal veins; S6 with 3 or fewer trichobothria on each side.

    |--Pachymeridiidae SP02
    |--Idiostolus SS02 [Idiostolidae SP02]
    |--Piesma H01 [Piesmatidae SP02]
    |    `--P. capitata H01
    |--Heterogaster Schilling 1829 [Heterogastridae] SE03
    |    `--H. urticae (Fabricius 1775) SE03
    |--Colobathristidae SP02
    |    |--Dayakiellinae SP02
    |    `--Phaenacantha [Colobathristinae] D14
    |         `--P. pacifica D14
    |--Berytidae SP02
    |    |--Berytus clavipes H01
    |    `--Neides H01
    |         |--N. propinquus Horváth 1901 H01
    |         `--N. tipularius H01
    |--Alydidae [Alydinae] SP02
    |    |--Leptocorisa oratorius R96
    |    |--Mirperus curvidens (Montr. 1858) [=Alydus curvidens] D14
    |    |--Riptortus ZS10
    |    |    |--R. clavatus TC01
    |    |    `--R. serripes ZS10
    |    `--Alydus H01
    |         |--A. calcaratus [incl. A. atratus, A. calcaratus var. hirsutus] H01
    |         |--A. pilosulus H01
    |         `--A. zichyi Horváth 1901 H01
    `--Coreidae [Coreinae] GE05
         |--Paradasynus spinosus YS01
         |--Thasus GE05
         |--Mictis profana (Fabr. 1803) [=Lygaeus profana] D14
         |--Amorbus AY04
         |--Amblypelta cocophaga MHG04
         |--Coreus dufourii Lucas 1847 E12
         |--Narnia inornata I92
         |--Aufeius impressicollis BM76
         |--Syromastes marginatus H01
         |--Therapha hyoscyami H01
         |--Chlorosoma macilentum H01
         |--Gonocerus L-SSY04
         |    |--G. annulipes Lucas 1847 E12
         |    `--G. juniperi Herrich-Schäffer 1839 L-SSY04
         |--Enoplops H01
         |    |--E. scapha H01
         |    `--E. sibirica H01
         |--Megalotomus H01
         |    |--M. junceus [incl. M. limbatus] H01
         |    `--M. ornaticeps H01
         |--Leptoglossus WS04
         |    |--L. bidentatus (Montr. 1855) [=Anisoscelis bidentatus] D14
         |    |--L. clypealis WS04
         |    `--L. oppositus BM76
         |--Myrmus H01
         |    |--M. glabellus Horváth 1901 H01
         |    |--M. longiventris H01
         |    `--M. miriformis H01
         `--Corizus H01
              |--C. crassicornis [incl. C. crassicornis var. abutilon] H01
              |--C. maculatus H01
              |--C. parumpunctatus H01
              `--C. subrufus [incl. C. capitatus] H01

*Type species of generic name indicated


[AY04] Austin, A. D., D. K. Yeates, G. Cassis, M. J. Fletcher, J. La Salle, J. F. Lawrence, P. B. McQuillan, L. A. Mound, D. J. Bickel, P. J. Gullan, D. F. Hales & G. S. Taylor. 2004. Insects 'Down Under'—diversity, endemism and evolution of the Australian insect fauna: examples from select orders. Australian Journal of Entomology 43 (3): 216-234.

[BM76] Bohart, R. M. & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

Carver, M., G. F. Gross & T. E. Woodward. 1991. Hemiptera. In: CSIRO. The Insects of Australia, 2nd ed., vol. 1, pp. 429-509. Melbourne University Press.

[D14] Distant, W. L. 1914. Rhynchota from New Caledonia and the surrounding islands. In Nova Caledonia: Forschungen in Neu-Caledonian und auf den Loyalty-Inseln. A. Zoologie vol. 1, Heft IV (F. Sarasin & J. Roux, eds) pp. 369-390, pl. 11-12. C. W. Kreidels Verlag: Wiesbaden.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l'Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[H01] Horváth, G. 1901. Hemipterák [Hemiptera]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy], Band II. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy], pp. 245-274. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[I92] Imes, R. 1992. The Practical Entomologist. Aurum Press: London.

[L-SSY04] López-Sebastián, E., J. Selfa & J. Ylla. 2004. Primeros datos del parasitismo de Ooencyrtus pityocampae (Mercet, 1921) sobre Graellsia isabelae (Graells, 1849) en condiciones de laboratorio. Graellsia 60 (1): 121-123.

[MHG04] Maynard, G. V., J. G. Hamilton & J. F. Grimshaw. 2004. Quarantine - phytosanitary, sanitary and incursion management: an Australian entomological perspective. Australian Journal of Entomology 43 (3): 318-328.

[R96] Rentz, D. 1996. Grasshopper Country: The abundant orthopteroid insects of Australia. University of New South Wales Press: Sydney.

[SE03] Scudder, G. G. E., & A. C. Eyles. 2003. Heterogaster urticae (Hemiptera: Heterogastridae), a new alien species and family to New Zealand. Weta 25: 8-13.

[SP02] Shcherbakov, D. E., & Yu. A. Popov. 2002. Superorder Cimicidea Laicharting, 1781: Order Hemiptera Linné, 1758. The bugs, cicadas, plantlice, scale insects, etc. [=Cimicida Laicharting, 1781, =Homoptera Leach, 1815 + Heteroptera Latreille, 1810). In History of Insects (A. P. Rasnitsyn & D. L. J. Quicke, eds) pp. 143-157. Kluwer Academic Publishers: Dordrecht.

[SS02] Sweet, M. H. & C. W. Schaefer. 2002. Parastrachiinae (Hemiptera: Cydnidae) raised to family level. Annals of the Entomological Society of America 95: 441-448.

[TC01] Taylor, DeM., & Y. Chinzei. 2001. Vitellogenin and its synthesis in the soft ticks. In Acarology: Proceedings of the 10th International Congress (R. B. Halliday, D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff, eds) pp. 622-627. CSIRO Publishing: Melbourne.

[WS04] Wang, Q., & G. L. Shi. 2004. Mating frequency, duration, and circadian mating rhythm of New Zealand wheat bug Nysius huttoni White (Heteroptera: Lygaeidae). New Zealand Entomologist 27: 113-117.

[YS01] Yoshizawa, K., & T. Saigusa. 2001. Phylogenetic analysis of paraneopteran orders (Insecta: Neoptera) based on forewing base structure with comments on monophyly of Auchenorrhycha (Hemiptera). Systematic Entomology 26: 1-13.

[ZS10] Zborowski, P. & R. Storey. 2010. A Field Guide to Insects in Australia. Reed New Holland: Sydney.

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