Neides tipularia, copyright Roger Key.

Belongs within: Panheteroptera.
Contains: Alydidae, Coreidae.

The Coreoidea are a group of plant-feeding bugs, most species of which possess ocelli and a series of longitudinal accessory veins in the forewing membrane (Carver et al. 1991). Members of this group include the Berytidae, stilt bugs, extremely slender and long-legged bugs characterised by short and spindle-shaped apical segments on the antennae. The Colobathristidae are also slender and long-legged (albeit not as remarkably so as the Berytidae) with narrow wings that have the venation on the membrane reduced or absent. The Piesmatidae are small bugs with a reduced membrane on the hemelytra, with the pronotum and corium being densely punctate. The Hyocephalidae are a group of seed-eating bugs endemic to Australia characterised by the presence of ovoid, pore-bearing plates ventrolaterally at the base of the abdomen (Carver et al. 1991).

Characters (from Carver et al. 1991): Macropterous or brachypterous; brachypters with antennifers inserted dorsolaterally; ocelli present; membrane of hemelytron in macropters with at least 6 and usually many more longitudinal veins; S6 with 3 or fewer trichobothria on each side.

<==Coreoidea SP02
    |--+--Coreidae G75
    |  `--Hyocephalidae G75
    |       |--Hyocephalus aprugnus G76
    |       `--Maevius indecorus CGW91
    |--Piesmatidae [Piesmatoidea, Piesmidae] SP02
    |    |--Mcateella interioris CGW91, G75
    |    |--Miespa CGW91
    |    `--Piesma CGW91
    |         |--P. capitata H01
    |         `--P. quadratum RD77
    |--Colobathristidae SP02
    |    |--Dayakiellinae SP02
    |    `--Phaenacantha [Colobathristinae] CGW91
    |         |--P. australiae CGW91
    |         |--P. pacifica Horváth 1904 K08
    |         `--P. saccharicida RD77
    `--Berytidae [Berytinae, Neididae] SP02
         |--Berytus clavipes H01
         |--Xenoloma Harris 1943 H96
         |--Phaconotus Harris 1943 H96
         |--Protacanthus Uhler 1893 D01
         |    `--P. halei CGW91
         |--Diabolonotus Henry 1996 H96
         |    `--*D. pilosus Henry 1996 H96
         |--Oedalocanthus Henry 1996 H96
         |    `--*O. ornatus Henry 1996 H96
         |--Metacanthus Costa 1848 D01
         |    |--M. decorus (Uhler 1893) [=Protacanthus decorus; incl. M. capitatus Uhler 1894] D01
         |    `--M. pluto CGW91
         `--Neides CGW91
              |--N. clavipes [=Gerris clavipes] L02
              |--N. maiponga G75
              |--N. propinquus Horváth 1901 H01
              `--N. tipularia G20 [=Gerris tipularius L02]

Coreoidea incertae sedis:
  Pachycoridium RJ93 [Hypocimicidae, Pachymeridiidae SP02, Psychrocoridae, Sisyrocoridae]
    `--P. letum Popov 1986 RJ93
  Idiostolidae [Idiostoloidea] SP02
    |--Idiostolus SS02
    |--Monteithocoris hirsutus CGW91
    `--Trisecus pictus WEE70
  Heterogastridae [Heterogastrinae] SE03
    |--Dinomachus MC13
    |--Woodwardothignus gracilis CGW91
    `--Heterogaster Schilling 1829 SE03
         `--H. urticae (Fabricius 1775) SE03
  Alydidae SP02
  Stenocephalidae CGW91
    |--Dicranocephalus CGW91
    `--Stenocephalus RD77
  Rhopalidae CGW91
    |--Liorhyssus hyalinus CGW91
    |--Rhopalus RD77
    |--Chorosoma macilentum RD77, H01
    |--Myrmus RD77
    |    |--M. glabellus Horváth 1901 H01
    |    |--M. longiventris H01
    |    `--M. miriformis H01
    `--Leptocoris [Serinethinae] CGW91
         |--L. augur K08
         |--L. insularis Kirkaldy 1908 K08
         |--L. mitellatus CGW91
         `--L. rufomarginatus CGW91

*Type species of generic name indicated


[CGW91] Carver, M., G. F. Gross & T. E. Woodward. 1991. Hemiptera (bugs, leafhoppers, cicadas, aphids, scale insects etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1 pp. 429–509. Melbourne University Press: Carlton (Victoria).

[D01] Distant, W. L. 1901. Rhynchotal notes.—IX. Heteroptera: fam. Coreidae. Annals and Magazine of Natural History, series 7, 7: 416–432.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[G75] Gross, G. F. 1975. Plant-feeding and Other Bugs (Hemiptera) of South Australia. Heteroptera—Part I. Handbook of the Flora and Fauna of South Australia.

[G76] Gross, G. F. 1976. Plant-feeding and Other Bugs (Hemiptera) of South Australia. Heteroptera—Part II. Handbook of the Flora and Fauna of South Australia.

[H96] Henry, T. J. 1996. Two new genera and two new species of New World stilt bugs (Heteroptera: Berytidae). Proceedings of the Entomological Society of Washington 98 (3): 533–540.

[H01] Horváth, G. 1901. Hemipterák [Hemiptera]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 245–274. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[K08] Kirkaldy, G. W. 1908. A catalogue of the Hemiptera of Fiji. Proceedings of the Linnean Society of New South Wales 33: 345–391, pl. 4.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[MC13] Majer, J. D., S. K. Callan, K. Edwards, N. R. Gunawardene & C. K. Taylor. 2013. Baseline survey of the terrestrial invertebrate fauna of Barrow Island. Records of the Western Australian Museum Supplement 83: 13–112.

[RD77] Richards, O. W., & R. G. Davies. 1977. Imms' General Textbook of Entomology 10th ed. vol. 2. Classification and Biology. Chapman and Hall: London.

[SE03] Scudder, G. G. E., & A. C. Eyles. 2003. Heterogaster urticae (Hemiptera: Heterogastridae), a new alien species and family to New Zealand. Weta 25: 8–13.

[SP02] Shcherbakov, D. E., & Yu. A. Popov. 2002. Superorder Cimicidea Laicharting, 1781: Order Hemiptera Linné, 1758. The bugs, cicadas, plantlice, scale insects, etc. [=Cimicida Laicharting, 1781, =Homoptera Leach, 1815+Heteroptera Latreille, 1810). In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 143–157. Kluwer Academic Publishers: Dordrecht.

[SS02] Sweet, M. H., & C. W. Schaefer. 2002. Parastrachiinae (Hemiptera: Cydnidae) raised to family level. Annals of the Entomological Society of America 95: 441–448.

[WEE70] Woodward, T. E., J. W. Evans & V. F. Eastop. 1970. Hemiptera (bugs, leafhoppers, etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 387–457. Melbourne University Press.

Last updated: 13 May 2022.

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