Belongs within: Coreoidea.
Contains: Leptoscelis, Colpura, Leptoglossus, Coreus.
The Coreidae, squash bugs, are a group of large plant-feeding bugs that typically produce strong repellent odours (Carver et al. 1991).
Characters (from Carver et al. 1991): Stout, usually robust bugs with relatively thick legs; bucculae extending to, and usually behind, level of antennifers; head usually much less than half as wide as base of pronotum; ocelli present; metapleural scent gland openings with distinct peritremes; posterior margins of T4 and T5 produced posteriorly in mid line; membrane of fore wing with longitudinal accessory veins; corium without a large hyaline area.
<==Coreidae [Corizidae]
|--Agriopocorinae CGW91
| |--Agriopocoris CGW91
| `--Tylocryptus egenus CGW91
|--Cloresminae D01a
| |--Notobitus abdominalis Distant 1901 D01a
| `--Cloresmus D01a
| |--C. khasianus Distant 1901 D01a
| `--C. modestus Distant 1901 D01a
|--Physomerinae D01b
| |--Rhyticoris spinipes [=Lygaeus spinipes; incl. Piezogaster patulus Walker 1871] D01b
| |--Physomerus D01b
| | |--P. grossipes (Fabr. 1794) [=Lygaeus grossipes; incl. P. delineatus Walker 1871] D01b
| | `--P. subargenteus Walker 1871 D01b
| `--Acanthocoris D01b
| |--A. anticus Walker 1871 D01b
| |--A. esau Distant 1901 D01b
| `--A. obscuricornis Dall. 1852 [incl. A. tarsalis Walker 1871] D01b
|--Pseudophloeinae CGW91
| |--Gralliclava CGW91
| |--Clavigralloides CGW91
| `--Clavigralla D01b
| |--C. acantharis (Fabr. 1803) [=Lygaeus acantharis; incl. C. tuberculata Dall. 1852] D01b
| |--C. aliena (Walker 1871) [=Cletus alienus] D01b
| |--C. dispar Walker 1872 (n. d.) D01b
| |--C. gibbosa G01
| |--C. horrens D01b
| |--C. indecora (Walker 1871) [=Cletus indecorus] D01b
| `--C. spinigera Walker 1872 (n. d.) D01b
|--Discogastrinae D01b
| |--Savius Stål 1862 [incl. Homoeoceroides Walker 1871] D01b
| | `--‘Homoeocerus’ diversicornis [=*Homoeoceroides diversicornis] D01b
| |--Cnemomis D01b
| | |--C. cognata Stål 1865 [incl. Hypselonotus signatus Walker 1871] D01b
| | `--C. dubia (Dall. 1852) [=Paryphes dubius; incl. Hypselonotus mundus Walker 1871] D01b
| |--Scamurius D01b
| | |--S. amabilis (Stål 1855) [=Paryphes amabilis; incl. Homoeoceroides incongruus Walker 1871] D01b
| | `--S. scutellaris (Walker 1871) [=Hypselonotus scutellaris] D01b
| `--Discogaster Burmeister 1835 [incl. Leptornytus Walker 1871] D01b
| |--D. drewseni D01b
| `--D. rufocornis (Walker 1871) [=Leptornytus rufocornis] D01b
|--Spartocerinae D01b
| |--Sephina D01b
| | |--S. humeralis Distant 1901 D01b
| | |--S. limbata [incl. S. atra Walker 1871] D01b
| | `--S. vinula [incl. Spartocera lampyroides Walker 1871] D01b
| `--Spartocera D01b
| |--S. batatas (Fabr. 1798) [=Lygaeus batatas] D01b
| |--S. brevicornis E66
| |--S. fusca (Thunb. 1783) [=Cimex fuscus; incl. Spartocerus diffusus] D01b
| |--S. gigantea D01b
| |--S. grandis Distant 1901 D01b
| |--S. pantomima (Dist. 1881) [=Sephina pantomima; incl. Sp. ortonedai Montand. 1897] D01b
| `--S. rubicunda Spin. in Gay 1852 [incl. S. chilensis Walker 1871] D01b
|--Gonocerinae D01b
| |--Gonocerus L-SSY04
| | |--G. annulipes Lucas 1847 E12
| | `--G. juniperi Herrich-Schäffer 1839 L-SSY04
| |--Cletomorpha D01b
| | |--C. kirbyi Distant 1901 [=C. denticulata Kirby 1891 (preoc.)] D01b
| | |--C. raja Distant 1901 D01b
| | `--C. walkeri Kirby 1891 [incl. C. benita Kirby 1891] D01b
| `--Plinacthus D01b
| |--P. acicularis (Fabr. 1803) [=Alydus acicularis; incl. Cletus conspicuus Walker 1871] D01b
| |--P. dubius (Herr.-Schäff. 1842) [=Gonocerus dubius; incl. Cletus clarus Walker 1871] D01b
| |--P. pungens (Thunb. 1783) [=Cimex pungens; incl. Gonocerus luridus Dall. 1852] D01b
| `--P. spinosus Stål 1859 [incl. Cletus mundus Walker 1871] D01b
|--Acanthocephalinae D01a
| |--Acanthocephala D01a
| | |--A. affinis (Walker 1871) [=Metapodius affinis] D01a
| | |--A. angustipes (Westw. in Hope 1842) [=Metapodius angustipes; incl. M. constrictus Walk. 1871] D01a
| | |--A. femorata L86
| | |--A. latipes (Drury 1782) [=Cimex latipes; incl. Metapodius sericeicollis Walk. 1871] D01a
| | `--A. unicolor (Westw. 1842) [=Metapodius unicolor; incl. M. distinctus Walk. 1871] D01a
| `--Laminiceps D01a
| |--L. cardinalis D01a
| |--L. ducalis D01a
| |--L. fasciatus D01a
| |--L. festivus Distant 1901 D01a
| |--L. megaera (Burm. 1835) [=Paryphes megaera] D01a
| `--L. sumptuosus Distant 1901 D01a
|--Centroscelinae D01b
| |--Acidomeria strigata (Walker 1871) [=Gonocerus strigatus] D01b
| |--Namacus annulicornis Stål 1870 [incl. N. rufescens Walker 1872] D01b
| |--Margus D01b
| | |--M. inornatus D01b
| | `--M. obscurator (Fabr. 1803) [=Coreus obscurator] D01b
| |--Catorhintha D01b
| | |--C. guttula (Fabr. 1794) [=Lygaeus guttula] D01b
| | `--C. semialba (Walker 1872) [=Namacus semialbus] D01b
| |--Paryphes D01b
| | |--P. festivus Costa 1863 [incl. P. gloriosus Walker 1871] D01b
| | `--P. viridipes Walker 1871 (n. d.) D01b
| `--Hypselonotus D01b
| |--H. concinnus [incl. H. propinquus Walker 1871] D01b
| |--H. interruptus Hahn 1831 [incl. Jadera subvittata Walker 1871] D01b
| `--H. lanceolatus Walker 1871 D01b
|--Mictinae D01b
| |--Saguntus lobulatus Stål 1865 [incl. Hypselonotus crassifemur Walker 1871] D01b
| |--Quintius pallens (Walker 1871) [=Hirileus pallens; incl. H. pallidus Walker 1871 D01b
| |--Molchina Amyot & Serv. 1843 [incl. Euplatycoris Walker 1871] D01b
| | `--M. granulata Stål 1870 [incl. Euplatycoris bellicornis Walker 1871] D01b
| |--Zoreva D01b
| | |--Z. marginalis (Walker 1871) [=Hypselonotus marginalis] D01b
| | `--Z. spinifera D01b
| |--Archimerus D01b
| | |--A. alternatus E66
| | |--A. calcarator E66
| | `--A. pallens (Dall. 1852) [=Physomerus pallens, Piezogaster pallens] D01b
| `--Nematopus D01b
| |--N. fasciatus Westw. in Hope 1842 [incl. Hypselonotus fascicollis Walker 1871] D01b
| |--N. indus (Linnaeus 1758) [=Cimex indus; incl. Hypselonotus pectoralis Walker 1871] D01b
| |--N. lepidus Stål 1862 [incl. Hypselonotus armatus Walker 1871] D01b
| `--N. ruficrus (Perty 1830) [=Anisoscelis ruficrus; incl. Hypselonotus ventralis Walker 1871] D01b
|--Leptoscelidini [Leptoscelinae] D01b
| |--Leptoscelis D01b
| |--Malvana minax (Walker 1871) [=Hypselonotus minax] D01b
| |--Amblyomia B96a
| |--Coribergia B96a
| |--Dalmatomammurius B96a
| |--Kalinckascelis B96a
| |--Malvanaioides B96a
| |--Plunentis B96a
| |--Sephinioides Brailovsky 1996 B96a
| | `--*S. ugaldei Brailovsky 1996 B96a
| `--Phthia D01b
| |--P. affinis Distant 1901 D01b
| |--P. cantharidina D01b
| |--P. decorata D01b
| |--P. lunata [incl. P. concinna Walker 1871] D01b
| |--P. picta D01b
| |--P. smaragdina (Walker 1871) [=Sphictyrtus smaragdinus] D01b
| `--P. ventralis D01b
|--Corizinae D01b
| |--Corizus H01
| | |--C. crassicornis [incl. C. crassicornis var. abutilon] H01
| | |--C. maculatus H01
| | |--C. parumpunctatus H01
| | `--C. subrufus [incl. C. capitatus] H01
| |--Jadera D01b
| | |--J. abdominalis Walker 1871 (n. d.) D01b
| | |--J. aeola (Dallas 1852) [=Serinetha aeola; incl. Lygaeus conspersus Walker 1872] D01b
| | |--J. antica (Walker 1872) [=Lygaeus anticus] D01b
| | |--J. haematoloma [incl. Lygaeus marginalis Walker 1872] D01c
| | `--J. sanguinolenta (Fabr. 1775) [=Cimex sanguinolentus; incl. Lygaeus rufoculis Kirby 1890] D01b
| `--Serinetha D01b
| |--S. abdominalis (Fabr. 1803) [=Lygaeus abdominalis; incl. S. taprobanensis Dallas 1852] D01b
| |--S. aethiops Dallas 1901 D01b
| |--S. fraterna D01b
| |--S. lurida Dallas 1852 D01b
| |--S. subrufescens (Kirby 1888) [=Lygaeus subrufescens] D01c
| `--S. vicina Dallas 1852 [incl. S. coxalis Kirby 1891, Astacops nigricornis Walker 1872] D01b
|--Colpurinae D01a
| | i. s.: Sciophyrus D01a
| | |--S. anticus (Walker 1871) [=Cletus anticus] D01a
| | `--S. crassus (Walker 1871) [=Cletus crassus] D01a
| | Pachycephalus D01a
| | |--P. opacus D01a
| | `--P. touchei Distant 1901 D01a
| `--Colpurini B96b
| |--Colpura D01a
| |--Agathyrna B96b
| |--Acanthotyla fasciata (Walker 1871) B96b, D01a [=Cletus fasciatus D01a]
| |--Monasavuhygia Brailovsky 1996 B96b
| | `--*M. cordata Brailovsky 1996 B96b
| |--Acarihygia Brailovsky 1993 B96b
| | `--*A. fijiana Brailovsky 1993 B96b
| `--Brachylybas Stål 1870 B96b
| |--*B. variegata (Le Guillou 1841) [=Gonocerus variegatus, B. variegatus] B96b
| |--B. delgadoi Brailovksy 1996 B96b
| |--B. taveuni Brailovsky 1996 B96b
| `--B. vanua Brailovsky 1996 B96b
|--Coreinae CGW91
| |--Mictis profana (Fabr. 1803) CGW91, D14 [=Lygaeus profana D14; incl. M. profanus var. crux K08]
| |--Amblypelta cocophaga CGW91, MHG04
| |--Aulacosternum nigrorubrum CGW91, C91
| |--Pompanotus CGW91
| |--Turrana CGW91
| |--Leptoglossus CGW91
| |--Coreus E12
| |--Amorbus CGW91
| | |--A. alternatus CGW91
| | `--A. rubiginosus G75
| `--Cletus CGW91
| |--C. apicifer Walker 1871 (n. d.) D01b
| |--C. bipunctatus (Westw. in Hope 1842) [=Coreus bipunctatus; incl. Cl. signatus Walker 1871] D01b
| |--C. fuscescens Walker 1871 D01b
| |--C. inconspicuus Walker 1871 D01b
| |--C. punctiger (Dall. 1852) [=Gonocerus punctiger; incl. Homoeocerus minax Walker 1871] D01b
| |--C. punctulatus (Westw. in Hope 1842) [=Coreus punctulatus; incl. Cl. femoralis Kirby 1891] D01b
| |--C. rubidiventris (Wastw. in Hope 1842) [=Coreus rubidiventris; incl. Cl. pallescens Walker 1871] D01b
| |--C. similis WEE70
| `--C. subnotatus Walker 1871 D01b
`--Homoeocerus [Homoeocerinae] D01a
|--H. abbreviatus (Fabr. 1794) [=Lygaeus abbreviatus; incl. H. umbratus Walker 1871] D01a
|--H. abdominalis Distant 1901 D01a
|--H. angulatus West. in Hope 1842 (see below for synonymy) D01a
|--H. annulatus (Thunb. 1822) [=Alydus annulatus; incl. Ceratopachys subsparsus Walker 1871] D01a
|--H. atkinsoni Distant 1901 D01a
|--H. auriculatus D01a
|--H. borneensis Distant 1901 D01a
|--H. concisus Walker 1871 D01a
|--H. graminis (Fabr. 1803) [=Lygaeus graminis; incl. H. turbidus Walker 1871] D01a
|--H. lineatus Walker 1871 (n. d.) D01a
|--H. lucidus Walker 1871 D01a
|--H. magnicornis D01a
|--H. malayensis Distant 1901 D01a
|--H. montanus Distant 1901 D01a
|--H. nigridorsum D01a
|--H. prominulus (Dall 1852) [=Ceratopachys prominulus; incl. C. vicinus Sall 1852] D01a
|--H. serrifer (Westw. in Hope 1842) (see below for synonymy) D01a
|--H. signatus Walker 1871 [incl. H. biplagiatus Stål 1873] D01a
|--H. simplex Walker 1871 D01a
|--H. sinicus Walker 1871 D01a
|--H. smecticus Distant 1901 D01a
|--H. subjectus Walker 1871 D01a
|--H. tinctus D01a
|--H. trimaculatus Distant 1901 D01a
|--H. unipunctatus (Thunb. 1783) [=Cimex unipunctatus; incl. H. chinensis Dall 1852] D01a
|--H. variabilis (Dall 1852) [=Ceratopachys variabilis; incl. H. anticus Walker 1871] D01a
|--H. walkerianus [incl. H. plagiatus Walker 1871 (preoc.)] D01a
`--H. yerburyi Distant 1901 D01a
Coreidae incertae sedis:
Paradasynus spinosus YS01
Thasus gigas GE05, EN20
Narnia inornata I92
Aufeius impressicollis BM76
Syromastes marginatus H01
Therapha hyoscyami H01
Enoplops H01
|--E. scapha H01
`--E. sibirica H01
Chelonidea G75
|--C. tabulata G75
`--C. vittiger G75
Anasa tristis CGW91, RD77
Hebeicoris xinboensis Hong 1985 RJ93
Uranocoris Walker 1871 D01b
`--U. suavis Walker 1871 D01b
Menenotus diminutus Walker 1871 D01b
Derepteryx truncata Walker 1871 D01b
Hyalymenus walkeri [incl. H. ichneumoniformis Walker 1871 (preoc.)] D01b
Verlusia rhombea D01b
Dulichius inflatus RD77
Theraptus RD77
Chariestrus antennator E66
Pachylis Lepeletier & Serville 1825 E66, K18
`--P. argentinus E66
Homoeocerus angulatus West. in Hope 1842 [incl. H. clarus Walker 1871, H. mundus Walker 1871, H. parallelus Walker 1871] D01a
Homoeocerus serrifer (Westw. in Hope 1842) [=Coreus serrifer; incl. H. parvulus Walker 1871, H. unipunctatus Dall 1852 non Cimex unipunctatus Thunb. 1783] D01a
*Type species of generic name indicated
REFERENCES
[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.
[B96a] Brailovsky, H. 1996a. A new genus and a new species of Leptoscelidini (Heteroptera: Coreidae) from Costa Rica. Proceedings of the Entomological Society of Washington 98 (2): 215–220.
[B66b] Brailovsky, H. 1996b. A revision of the tribe Colpurini (Heteroptera: Coreidae) from the Fiji islands. Proceedings of the Entomological Society of Washington 98 (3): 473–490.
[CGW91] Carver, M., G. F. Gross & T. E. Woodward. 1991. Hemiptera (bugs, leafhoppers, cicadas, aphids, scale insects etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers vol. 1 pp. 429–509. Melbourne University Press: Carlton (Victoria).
[C91] CSIRO. 1991. The Insects of Australia: A textbook for students and research workers vol. 1. Melbourne University Press: Carlton (Victoria).
[D01a] Distant, W. L. 1901a. Rhynchotal notes.—VIII. Heteroptera: fam. Coreidae. Annals and Magazine of Natural History, series 7, 7: 6–22.
[D01b] Distant, W. L. 1901b. Rhynchotal notes.—IX. Heteroptera: fam. Coreidae. Annals and Magazine of Natural History, series 7, 7: 416–432.
[D01c] Distant, W. L. 1901c. Rhynchotal notes.—X. Heteroptera: fam. Lygaeidae. Annals and Magazine of Natural History, series 7, 7: 531–541.
[D14] Distant, W. L. 1914. Rhynchota from New Caledonia and the surrounding islands. In: Sarasin, F., & J. Roux (eds) Nova Caledonia: Forschungen in Neu-Caledonian und auf den Loyalty-Inseln. A. Zoologie vol. 1 pt 4 pp. 369–390, pls 11–12. C. W. Kreidels Verlag: Wiesbaden.
[EN20] Eggli, U., & R. Nyffeler (eds) 2020. Illustrated Handbook of Succulent Plants: Monocotyledons 2nd ed. Springer.
[E66] Evans, H. E. 1966. The Comparative Ethology and Evolution of the Sand Wasps. Harvard University Press: Cambridge (Massachusetts).
[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l'Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.
[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.
[G75] Gross, G. F. 1975. Plant-feeding and Other Bugs (Hemiptera) of South Australia. Heteroptera—Part I. Handbook of the Flora and Fauna of South Australia.
[G01] Gupta, S. K. 2001. A conspectus of natural enemies of phytophagous mites and mites as potential biocontrol agents of agricultural pests in India. In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10th International Congress pp. 484–497. CSIRO Publishing: Melbourne.
[H01] Horváth, G. 1901. Hemipterák [Hemiptera]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 245–274. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.
[I92] Imes, R. 1992. The Practical Entomologist. Aurum Press: London.
[K08] Kirkaldy, G. W. 1908. A catalogue of the Hemiptera of Fiji. Proceedings of the Linnean Society of New South Wales 33: 345–391, pl. 4.
[K18] Kury, A. B. 2018. Familial nomina in harvestmen (Arachnida, Opiliones). Bionomina 13: 1–27.
[L86] Lindquist, E. E. 1986. The world genera of Tarsonemidae (Acari: Heterostigmata): a morphological, phylogenetic, and systematic revision, with a reclassification of family-group taxa in the Heterostigmata. Memoirs of the Entomological Society of Canada 118 (S136): 1–517.
[L-SSY04] López-Sebastián, E., J. Selfa & J. Ylla. 2004. Primeros datos del parasitismo de Ooencyrtus pityocampae (Mercet, 1921) sobre Graellsia isabelae (Graells, 1849) en condiciones de laboratorio. Graellsia 60 (1): 121–123.
[MHG04] Maynard, G. V., J. G. Hamilton & J. F. Grimshaw. 2004. Quarantine—phytosanitary, sanitary and incursion management: an Australian entomological perspective. Australian Journal of Entomology 43 (3): 318–328.
[RD77] Richards, O. W., & R. G. Davies. 1977. Imms' General Textbook of Entomology 10th ed. vol. 2. Classification and Biology. Chapman and Hall: London.
[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.
[WEE70] Woodward, T. E., J. W. Evans & V. F. Eastop. 1970. Hemiptera (bugs, leafhoppers, etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 387–457. Melbourne University Press.
[YS01] Yoshizawa, K., & T. Saigusa. 2001. Phylogenetic analysis of paraneopteran orders (Insecta: Neoptera) based on forewing base structure with comments on monophyly of Auchenorrhycha (Hemiptera). Systematic Entomology 26: 1–13.
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