Reconstruction of Karabasia evansi, from Grimaldi & Engel (2005).

Belongs within: Hemiptera.
Contains: Heteroptera.

The clade Heteropteroidea unites the true bugs of the Heteroptera with their living sister taxon, the Southern Hemisphere Peloridiidae (the only living members of the Coleorrhyncha). The peloridiids are flattened, broad-bodied bugs with widely-separated eyes on short peduncles, paranotal lobes on the pronotum, and forewings with net-like veins. They are found amongst moss and leaf litter in South America and Australasia (Grimaldi & Engel 2005).

Synapomorphies (from Grimaldi & Engel 2005): Antennae clavate; gula present on underside of head; thoracic scent glands present in adults; wings overlapping, lying flat against abdomen, with specialised wing-coupling devices; abdomen flattened, with spiracles on ventral surface.

    |--+--Heteroptera GE05
    |  `--Scytinopteroidea SP02
    |       |--Scytinopteridae SP02
    |       |--Serpentivenidae SP02
    |       |--Ipsviciidae SP02
    |       |--Stenovicia F71 [Stenoviciidae SP02]
    |       |    `--S. angustata Evans 1943 F71
    |       `--Paraknightia F71 [Paraknightiidae SP02]
    |            `--P. magnifica Evans 1943 F71
    `--Coleorrhyncha [Peloridiina] GE05
         |--Mesoscytina [Cicadocorinae, Progonocimicidae, Progonocimicoidea] SP02
         `--Peloridioidea SP02
              |--Karabasia GE05 [Karabasiidae SP02, Karabasiinae]
              |    `--K. evansi GE05
              |--Hoploridium [Hoploridiidae, Hoploridiinae] GE05
              |    `--H. dollingi SP02
              `--Peloridiidae GE05
                   |--Peloridora GE05
                   |--Howeria AY04
                   `--Peloridium hammoniorum V73

*Type species of generic name indicated


[AY04] Austin, A. D., D. K. Yeates, G. Cassis, M. J. Fletcher, J. La Salle, J. F. Lawrence, P. B. McQuillan, L. A. Mound, D. J. Bickel, P. J. Gullan, D. F. Hales & G. S. Taylor. 2004. Insects 'Down Under' - diversity, endemism and evolution of the Australian insect fauna: examples from select orders. Australian Journal of Entomology 43 (3): 216-234.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1-167.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[SP02] Shcherbakov, D. E., & Yu. A. Popov. 2002. Superorder Cimicidea Laicharting, 1781: Order Hemiptera Linné, 1758. The bugs, cicadas, plantlice, scale insects, etc. [=Cimicida Laicharting, 1781, =Homoptera Leach, 1815 + Heteroptera Latreille, 1810). In History of Insects (A. P. Rasnitsyn & D. L. J. Quicke, eds) pp. 143-157. Kluwer Academic Publishers: Dordrecht.

[V73] Vandel, A. 1973. Les isopodes terrestres de l'Australie. Étude systématique et biogéographique. Mémoires du Muséum National d'Histoire Naturelle, Série A, Zoologie 82: 1-171.

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