Tingoidea

Ceratocader armatus, from here.


Belogs within: Cimicomorpha.

The Tingoidea are a group of mostly plant-feeding bugs. The Tingidae, lace bugs, are distinctive due to the often ornate lace-like ornamentation of the thorax and wings.

Synapomorphies (from Shcherbakov & Popov 2002): Head trichobothria absent; costal fracture reduced; tarsi two-segmented; spermatheca non-functional.

<==Tingoidea
    |--Thaumastocoridae SP02
    |--Joppeicidae SP02
    |--Vianaididae S02
    `--Tingidae GE05
         |  i. s.: Tanybyrsa cumberi Drake 1959 H82
         |         Stephanitis YS01
         |         Corythucha TW05
         |         Vianagramma GE05
         |         Golmonia GE05
         |--Tinginae M09
         |    |--Radinacantha Hacker 1929 [Tingini] M09
         |    |    |--*R. reticulata Hacker 1929 M09
         |    |    |--R. darwini Moir 2009 M09
         |    |    |--R. dondiorum Moir 2009 M09
         |    |    |--R. prudens M09
         |    |    `--R. tasmanica Hacker 1929 M09
         |    |--Cochlochila D55
         |    |    |--C. boxiana Drake 1953 D55
         |    |    `--C. bukobana D55
         |    |--Compseuta D55
         |    |    |--C. bispinosa D55
         |    |    |--C. expleta Drake 1955 D55
         |    |    |--C. holana D55
         |    |    |--C. montandoni D55
         |    |    |--C. ornatella (Stål 1855) (see below for synonymy) D55
         |    |    `--C. picta Schouteden 1923 D55
         |    |--Kapiriella maynei (Schouteden 1919) [=Lembella maynei] D55
         |    |--Eteoneus congolensis Schouteden 1923 D55
         |    |--Angolotingis Drake 1955 D55
         |    |    `--*A. vilhenai Drake 1955 D55
         |    |--Aglotingis nimbana Drake 1954 D55
         |    |--Arushia D55
         |    |--Paracopium stolidum (Horvarth 1912) [=Copium stolidum] D55
         |    `--Phyllontocheila dilatata (Guerin-Meneville 1831) [=Tingis dilatata] D55
         |--Cantacaderinae [Cantacaderidae] SP02
         |    |  i. s.: Sinaldocader SP02
         |    |         Astolphos B27
         |    |--Gonycentrum Bergroth 1898 [Phatnomatini] GM10
         |    `--Cantacaderini GM10
         |         |  i. s.: Cantacader divisus Bergroth 1908 D55
         |         |--+--Carldrakeana Froeschner 1968 GM10
         |         |  |    |--*C. tindalei (Hacker 1928) [=Phatnoma tindalei] GM10
         |         |  |    |--C. engista (Drake & Ruhoff 1961) GM10
         |         |  |    |--C. karli Guilbert & Moir 2010 GM10
         |         |  |    |--C. pallida Lis 2000 GM10
         |         |  |    `--C. socia (Drake & Ruhoff 1961) GM10
         |         |  `--+--Cyperobia Bergroth 1927 GM10
         |         |     |    `--*C. carectorum Bergroth 1927 B27
         |         |     `--Stenocader Drake & Hambleton 1944 GM10
         |         `--+--Nectocader Drake 1928 ML12
         |            |--Caledoderus Guilbert 2012 ML12
         |            `--+--Australocader Lis 1997 ML12
         |               `--Ceratocader Drake 1950 [Ceratocaderini] ML12
         |                    |--*C. armatus (Hacker 1928) ML12
         |                    |--C. bridgettae Moir & Lis 2012 ML12
         |                    |--C. coatesi Moir & Lis 2012 ML12
         |                    |--C. dentatus (Hacker 1928) ML12
         |                    |--C. fulvus Lis 2000 ML12
         |                    |--C. langlandsi Moir & Lis 2012 ML12
         |                    `--C. monteithi Lis 2000 ML12
         `--Serenthia [Serenthiinae] D55
              |--S. angolana Drake 1955 D55
              `--S. longa Drake 1954 D55

Compseuta ornatella (Stål 1855) [=Tropidocheila ornatella, Monanthia (Physatocheila) ornatella; incl. M. nigrosternum Horvarth 1910] D55

*Type species of generic name indicated

REFERENCES

[B27] Bergroth, E. 1927. Hemiptera Heteroptera from New Zealand. Transactions and Proceedings of the New Zealand Institute 57: 671-684.

[D55] Drake, C. J. 1955. Angolan Tingidae (Hemiptera). Diamang Publicações Culturais 27: 83-90.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[GM10] Guilbert, E. & M. L. Moir. 2010. A new species of Carldrakeana (Insecta: Heteroptera: Tingidae) from Western Australia. Records of the Western Australian Museum 25 (4): 382-386.

[H82] Helmore, D. W. 1982. Drawings of New Zealand insects. Bulletin of the Entomological Society of New Zealand 8: 1-52.

[M09] Moir, M. L. 2009. Two new species of the lacebug genus Radinacantha (Hemiptera: Heteroptera: Tingidae) from Australia. Records of the Western Australian Museum 25 (3): 269-275.

[ML12] Moir, M. L. & B. Lis. 2012. Description of three new species of Ceratocader (Hemiptera: Heteroptera: Tingidae) from Western Australia. Records of the Western Australian Museum 27 (2): 148-155.

[SP02] Shcherbakov, D. E., & Yu. A. Popov. 2002. Superorder Cimicidea Laicharting, 1781: Order Hemiptera Linné, 1758. The bugs, cicadas, plantlice, scale insects, etc. [=Cimicida Laicharting, 1781, =Homoptera Leach, 1815 + Heteroptera Latreille, 1810). In History of Insects (A. P. Rasnitsyn & D. L. J. Quicke, eds) pp. 143-157. Kluwer Academic Publishers: Dordrecht.

[TW05] Terry, M. D., & M. F. Whiting. 2005. Mantophasmatodea and phylogeny of the lower neopterous insects. Cladistics 21: 240-257.

[YS01] Yoshizawa, K., & T. Saigusa. 2001. Phylogenetic analysis of paraneopteran orders (Insecta: Neoptera) based on forewing base structure with comments on monophyly of Auchenorrhycha (Hemiptera). Systematic Entomology 26: 1-13.

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