Dorsal and ventral views of Propalticus specimen, from here.

Belongs within: Coleoptera.
Contains: Monotomidae, Nitidulidae, Xenoscelinae, Pharaxonothinae, Erotylinae, Cryptophilinae, Languriinae, Loberinae, Cucujidae, Cryptophagidae, Silvanidae, Phalacridae, Laemophloeidae.

The Cucujoidea is an assemblage of beetles characterised by the presence of a ring-type (cucujoid) aedeagus in the male and a more or less cylindrical, pygopod-like tenth abdominal segment in the larva. Larvae also usually have a mandible with a prostheca and an often falcate mala, and may have six stemmata (Lawrence & Britton 1991).

Cucujoids include a number of fungus-feeding beetle families, such as the Erotylidae, Cryptophagidae and Corylophidae. The Erotylidae also includes a number of plant-feeding species that were previously treated as a separate family Languriidae (Leschen 2003). The Hobartiidae are a family of oblong, pubescent fungus-feeding beetles found in Australia and South America, closely related to the pollen-feeding Boganiidae of Australia and southern Africa. Members of these latter two families possess dorsal tubercles on the mandible that fit against lateral cavities on the clypeus (Lawrence & Britton 1991). They are also closely related to the palmetto beetles of the genus Smicrips, a Neotropical group of tiny beetles associated with vegetation, particularly with palms (Bouchard 2014).

<==Cucujoidea [Aphidiphagi, Clavicornia, Cucujipes, Erotilenae]
    |--+--+--Monotomidae MW15
    |  |  `--Nitidulidae MW15
    |  `--+--Hobartiidae MW15
    |     |    |--Hydnobioides pubescens LB91
    |     |    `--Hobartius B14
    |     |         |--H. chilensis Tomaszewska & Ślipiński 1995 B14
    |     |         `--H. tasmanicus LB91
    |     `--+--Smicrips [Smicripidae] MW15
    |        |    `--S. palmicola LeConte 1878 B14
    |        `--Boganiidae MW15
    |             |  i. s.: Athertonium LB91
    |             |--+--Boganium B98
    |             |  `--Afroboganium B98
    |             `--Paracucujinae CN91
    |                  |--Metacucujus encephalarti B98
    |                  `--Paracucujus rostratus Sen Gupta & Crowson 1966 B14
    `--+--+--Erotylidae [Languriidae] MW15
       |  |    |  i. s.: Hedista [Triplacinae] LB91
       |  |    |         Pselaphacus MW15
       |  |    |--Xenoscelinae L03
       |  |    `--+--Pharaxonothinae L03
       |  |       `--+--+--Erotylinae L03
       |  |          |  `--Cryptophilinae L03
       |  |          `--+--Languriinae L03
       |  |             `--Loberinae L03
       |  `--+--Protosphindus [Protosphindinae] MW15
       |     |    |--P. bellus B14
       |     |    `--P. chilensis Sen Gupta & Crowson 1977 B14
       |     `--+--Helotidae MW15
       |        |    |--Neohelota B14
       |        |    `--Helota B14
       |        |         |--H. fulviventris Kolbe 1886 B14
       |        |         |--H. gemmata B14
       |        |         `--H. gorhami B14
       |        `--+--Ericmodes MW15 [incl. Protocucujus B74; Protocucujidae MW15]
       |           |    |--E. australis LB91
       |           |    |--E. fuscitarsis Reitter 1878 B14
       |           |    `--E. nigris B14
       |           `--Sphindidae MW15
       |                |--Sphindus MW15
       |                `--Aspidiphorus LB91
       |                     |--A. humeralis LB91
       |                     `--A. orbiculatus V09
       `--+--+--Phloeostichidae MW15
          |  |    |--Rhopalobrachium AY04
          |  |    |--Myriabola [Myriabolinae] L03
          |  |    |    `--M. grouvelliana L03
          |  |    `--Hymaea [Hymaeinae] LB91
          |  |         |--H. laticollis Carter 1908 C08
          |  |         |--H. magna LB91
          |  |         `--H. succinifera Pascoe 1869 M86
          |  `--+--Cucujidae MW15
          |     `--+--Agapytho [Agapythidae] MW15
          |        |    `--A. foveicollis Broun 1921 B14
          |        `--Priasilphidae [Priasilphinae] MW15
          |             |--Priastichus tasmanicus Crowson 1973 B14
          |             |--Chileosilpha B14
          |             `--Priasilpha B14
          |                  |--P. angulata Leschen, Lawrence & Ślipiński 2005 B14
          |                  `--P. obscura LB91
          `--+--+--Cryptophagidae MW15
             |  `--+--Cavognathidae MW15
             |     |    |--Taphropiestes pullivora (Crowson 1964) B14
             |     |    |--Cavognatha pullivora LB91
             |     |    |--Neocercus LB91
             |     |    `--Zeonidicola E02
             |     |         |--Z. chathamensis Watt 1980 E02
             |     |         `--Z. dumbletonii LB91
             |     `--+--Silvanidae MW15
             |        `--+--Paratomaria MW15
             |           `--Atomaria MW15
             |                |--A. analis C01
             |                |--A. fuscata C01
             |                `--A. ruficornis L03
             `--+--+--Phalacridae MW15
                |  `--+--Laemophloeidae MW15
                |     `--Propalticidae MW15
                |          |--Slipinskogenia B14
                |          `--Propalticus B14
                |               |--P. doddi B74
                |               |--P. oculatus Sharp 1879 B14
                |               `--P. simplex LB91
                `--+--+--Cyclaxyra [Cyclaxyridae] MW15
                   |  |    |--C. jelineki Gimmel, Leschen & Ślipiński 2009 B14
                   |  |    `--C. politula B14
                   |  `--Uleiotinae LB91
                   |       |--Dendrophagus australis LB91
                   |       |--Brontopriscus LB91
                   |       |--Psammoecus LB91
                   |       |--Cryptamorpha desjadinsi LB91
                   |       `--Uleiota Latreille 1802 MW15, L02
                   |            `--*U. flavipes [=Brontes flavipes] L02
                   `--+--Myrabolia [Myraboliidae] MW15
                      |    `--M. brevicornis (Erichson 1842) B14 [=Silvanus brevicornis H73]
                      `--Passandridae [Passandrinae] MW15
                           |--Catogenus A71
                           |--Ancistria LB91
                           |--Aulonosoma LB91
                           `--Passandra [incl. Hectarthrum] LB91
                                |--P. heros LB91 [=Hectarthrum heros B70]
                                `--P. simplex (Murray 1867) B14 [=Hectarthrum simplex G89]

Cucujoidea incertae sedis:
  Tasmosalpingus [Tasmosalpingidae, Tasmosalpinginae] B14
    `--T. quadrispilotus Lea 1919 B14
  Phloiophilus B14 [Phloiophilidae MW15]
    `--P. edwardsii Stephens 1830 B14
  Cybocephalus LB91 [Cybocephalidae MW15, Cybocephalinae]
  Lamingtonium [Lamingtoniidae] L03
    |--L. binnaburense L03
    |--L. loebli Lawrence & Leschen 2003 B14
    `--L. thayerae L03
  Engis australis [=Episcapha australis] B35
  Sinisilvana Hong 2002 [Sinisilvanidae] H02
    `--*S. fushunensis Hong 2002 H02

*Type species of generic name indicated


[A71] Askew, R. R. 1971. Parasitic Insects. Heinemann Educational Books: London.

[AY04] Austin, A. D., D. K. Yeates, G. Cassis, M. J. Fletcher, J. La Salle, J. F. Lawrence, P. B. McQuillan, L. A. Mound, D. J. Bickel, P. J. Gullan, D. F. Hales & G. S. Taylor. 2004. Insects ‘Down Under’—diversity, endemism and evolution of the Australian insect fauna: examples from select orders. Australian Journal of Entomology 43 (3): 216–234.

[B98] Bellés, X. 1998. A new subgenus and two new species of Sphaericus (Coleoptera: Ptinidae) from Western Australia. European Journal of Entomology 95: 263–268.

[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d'Urville. Faune entomologique de l'océan Pacifique, avec l'illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.

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[B70] Britton, E. B. 1970. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 495–621. Melbourne University Press.

[B74] Britton, E. B. 1974. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers. Supplement 1974 pp. 62–89. Melbourne University Press.

[C08] Carter, H. J. 1908. Revision of the genus Seirotrana, together with descriptions of new species of other Australian Coleoptera. Proceedings of the Linnean Society of New South Wales 33: 392–422.

[CN91] Cranston, P. S., & I. D. Naumann. 1991. Biogeography. In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1 pp. 180–197. Melbourne University Press: Carlton (Victoria).

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[E02] Emberson, R. M. 2002. The beetle (Coleoptera) fauna of the Chatham Islands: additions and corrections. New Zealand Entomologist 25: 69–77.

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 5–72.

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[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LB91] Lawrence, J. F., & E. B. Britton. 1991. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 543–683. Melbourne University Press: Carlton (Victoria).

[L03] Leschen, R. A. B. 2003. Erotylidae (Insecta: Coleoptera: Cucujoidea): phylogeny and review. Fauna of New Zealand 47: 1–108.

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[MW15] McKenna, D. D., A. L. Wild, K. Kanda, C. L. Bellamy, R. G. Beutel, M. S. Caterino, C. W. Farnum, D. C. Hawks, M. A. Ivie, M. L. Jameson, R. A. B. Leschen, A. E. Marvaldi, J. V. McHugh, A. F. Newton, J. A. Robertson, M. K. Thayer, M. F. Whiting, J. F. Lawrence, A. Ślipiński, D. R. Maddison & B. D. Farrell. 2015. The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Systematic Entomology 40 (4): 835–880.

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Last updated: 6 October 2018.

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