Tragiscus dimidiatus, photographed by Udo Schmidt.

Belongs within: Scarabaeinae.

The Oniticellini are a tribe of dung beetles found in most parts of the world except Australia. Like the closely related Onthophagini, oniticellins are tunnelers rather than rollers of dung (Philips et al. 2004).

Synapomorphies (from Philips et al. 2004): Wing with distinct and distally truncate to slightly emarginate pigmentation apically; costal margin at wing base without fine striae; epipharynx with anterior median dorsal processtruncate but narrowly widened at apex; maxilla with galea approximately transverse; labium with two palpomeres; lateral edge of mentum slightly sinuate with a shallow notch posterior of middle; longitudinal arms of labium adjacent to distal transverse bridge with narrow parallel-sided arms extending out laterally from triangular base; lateral cervical sclerite with setal brush located mesally near apex short; seventh and eighth antennomeres with small cavity on anterior surface; anterior edge of approximately ventrally directed anterior transverse plate on prescutum narrowly pointed; scutum-scutellum on one level; scutellum width apically 2.0× length; pygidium with transverse ridge absent, dark pigment present at base; genital capsule with transverse darkened portion on ventral plate; sternal region with furcal arm tip projected laterally.

  |  i. s.: Helictopleurus PPS04
  |--Tiniocellus spinipes PPS04
  `--+--Liatongus militaris PPS04
     `--+--+--Scaptocnemis segregis PPS04
        |  `--+--Cyptochirus ambiguus PPS04
        |     `--Drepanocerus reconditus PPS04, L88
        `--+--Euoniticellus intermedius PPS04
           `--+--Oniticellus PPS04
              |    |--O. cubiensis L88
              |    |--O. pallipes [=Scarabaeus pallipes] G89
              |    `--O. pictus PPS04
              `--Tragiscus dimidiatus PPS04

*Type species of generic name indicated


[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880. – IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a 7: 5-72.

[L88] Liebherr, J. K. 1988. General patterns in West Indian insects, and graphical biogeographic analysis of some circum-Caribbean Platynus beetles (Carabidae). Systematic Zoology 37 (4): 385-409.

[PPS04] Philips, T. K., E. Pretorius & C. H. Scholtz. 2004. A phylogenetic analysis of dung beetles (Scarabaeinae: Scarabaeidae): Unrolling an evolutionary history. Invertebrate Systematics 18: 53-88.

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