Heterorrhina imperatrix, photographed by Les Day.

Belongs within: Scarabaeoidea.

The Cetoniinae are a group of somewhat flattened scarab beetles which are usually strikingly patterned, metallic or enamelled in coloration. Larvae are found in rotting wood or decaying vegetation, while adults are diurnal nectar feeders (Lawrence & Britton 1991).

Characters (from Lawrence & Britton 1991): Tarsal claws equal, more or less fixed; fore coxae conical, produced ventrally; mid coxae transverse or only slightly oblique; mesothoracic epimera usually visible from above; dorsum slightly flattened, usually brightly coloured; elytra without narrow membranous margin; spiracles diverging so that several lie on abdominal sternites; abdomen not shorter than metasternum; antennal insertions exposed; mandibles concealed. Larva with head partly covered by prothorax; apical antennal segment about as wide as penultimate segment, which lacks a sensorium; labrum symmetrical, with deeply pigmented notch on each side of midline; mandible with ventral oval stridulatory area; galea and lacinia entirely fused to form mala; legs short, hind coxa shorter than width of clypeus; anal cleft transverse; segments 9 and 10 partly fused, suture between them incomplete dorsally; S9 without raster composed of bilaterally symmetrical rows of setae.

    |--Plectrone polita Jans. 1884 G89b
    |--Macronota suturalis Snell & Voll 1858 G89b
    |--Valgus modiglianii Gestro 1889 G89b
    |--Heterorrhina G89b
    |    |--H. decora [incl. Coryphocera sexmaculata Mohnike 1871] G89b
    |    |--H. dohrnii Lansb. 1883 G89b
    |    `--H. imperatrix G89b
    |--Glycyphana G89b
    |    |--G. albomaculata Mohnike 1871 G89b
    |    |--G. bisignata Kraatz 1885 G89b
    |    |--G. malayensis Guér. 1840 [=Euryomia malayensis] G89b
    |    |--G. rufovittata Guér. 1840 [=Euryomia rufovittata; incl. G. rufovittata var. melanaria Kraatz 1885] G89b
    |    `--G. sinuata Wall 1868 G89b
    `--Cetonia G89b
         |--C. acuminata [=Protaetia acuminata] G89b
         |--C. ciliata [=Protaetia ciliata] G89b
         |--C. mandarinea [=Protaetia mandarinea] G89b
         |--C. mixta G89b
         |--C. strigicollis Kraatz 1885 G89b
         `--C. thoracica G89a

*Type species of generic name indicated


[G89a] Gestro, R. 1889a. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso «Esploratore» dal 16 Novembre 1879 al 26 Febbraio 1880. – IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a 7: 5-72.

[G89b] Gestro, R. 1889b. Sopra alcune cetonie dell'isola Nias e della costa occidentale di Sumatra raccolte dal Dott. Elio Modigliani. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a 7: 93-99.

Lawrence, J. F., & E. B. Britton. 1991. Coleoptera. In: CSIRO. The Insects of Australia: A textbook for students and research workers, 2nd ed., vol. 2, pp. 543-683. Melbourned University Press.

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