Dionycha

Male Agroeca brunnea, copyright Benedikt.


Belongs within: Araneomorphae.
Contains: Olios, Heteropoda, Pseudopoda, Eusparassinae, Sparassinae, Thelcticopis, Thanatus, Philodromus, Diaea, Ozyptila, Xysticus, Thomisus, Salticus, Attus, Maratus, Gallieniellidae, Lamponidae, Gnaphosidae, Prodidomidae, Clubiona.

The Dionycha are a diverse clade of spiders united by the loss of the unpaired leg tarsal claw (Jocqué & Dippenaar-Schoeman 2007) Major subgroups of the Dionycha include the Salticidae, jumping spiders (referred to in many older publications as the Attidae), a distinctive cosmopolitan group of hunting spiders in which the anterior median eyes are much larger than the other eyes and are directed forwards on the carapace. Salticids have much better vision than many other spiders, with most species using their vision to actively track prey. The majority of salticids are included in the subfamily Salticinae, characterised by the absence of a tarsal claw on the female pedipalp or a median apophysis on the male pedipalp, and by a more abrupt gait than in members of other subfamilies.

Dionycha [Laterigradae] JD-S07
    |  i. s.: Botryogaster Doleschall 1859 D59
    |           `--*B. coerulescens Doleschall 1859 D59
    |--Sparassidae [Heteropodidae, Heteropodoidae] JD-S07
    |    |  i. s.: Olios JR08
    |    |         Leucorchestris Lawrence 1962 NHJ08
    |    |           |--*L. arenicola Lawrence 1962 [incl. L. kochi Lawrence 1965] NHJ08
    |    |           |--L. alexandrina Lawrence 1966 NHJ08
    |    |           |--L. sabulosa Lawrence 1966 NHJ08
    |    |           |--L. setifrons Lawrence 1966 NHJ08
    |    |           `--L. steyni Lawrence 1965 NHJ08
    |    |         Typostola magnifica B96
    |    |         Holconia JPV06
    |    |           |--H. immanis (Koch 1867) JPV06 [=Isopeda immanis FF99]
    |    |           `--H. vasta B96
    |    |         Isopeda Koch 1875 PVD10
    |    |           |--I. canberrana Hirst 1992 E07
    |    |           |--I. leishmani Hogg 1903 [=Isopoda (l. c.) leishmani] E07
    |    |           |--I. montana FF99
    |    |           |--I. vastata Strand 1907 J98
    |    |           `--I. villosa Koch 1875 PVD10
    |    |         Neosparassus B96
    |    |           |--N. calligaster [=Olios calligaster] B96
    |    |           |--N. diana [=Olios diana] B96
    |    |           `--N. punctatus [=Olios punctatus] B96
    |    |         Isopedella Hirst 1990 PVD10
    |    |           |--I. conspersa (Koch 1875) [incl. Isopeda maculigastra Strand 1907, Isopedella maculigaster] J98
    |    |           |--I. pessleri (Thorell 1870) [=Isopeda pessleri] E07
    |    |           `--I. victorialis Hirst 1993 PVD10
    |    |         Palystella Lawrence 1928 JK05
    |    |           |--*P. sexmaculatus JK05
    |    |           |--‘Olios’ isongonis Strand 1915 J98
    |    |           |--P. namaquensis Lawrence 1938 JK05
    |    |           `--P. pallida Lawrence 1938 JK05
    |    |         Palystes Koch 1875 JK05
    |    |           |--*P. castaneus (Latreille 1819) JK05
    |    |           |--P. ellioti J98
    |    |           |--P. hoehneli Simon 1890 [incl. P. bornemiszai] JK05
    |    |           |--P. modificus Strand 1906 J98
    |    |           |    |--P. m. modificus J98
    |    |           |    `--P. m. minor Strand 1906 J98
    |    |           |--P. perornatus Pocock 1900 JK05
    |    |           |--P. speciosus Poc. 1898 R13
    |    |           `--P. superciliosus Koch 1875 J98
    |    |         Eodelena melanochelis (Strand 1913) [=Zachria melanochelis] J98
    |    |         Midamus Sim. 1880 T94
    |    |           `--M. lutescens Thorell 1894 T94
    |    |         Origes Simon 1897 JR08
    |    |           |--*O. pollens Simon 1897 JR08
    |    |           |--O. chloroticus Mello-Leitão 1945 JR08
    |    |           `--O. nigrovittatus (Keyserling 1880) [=Sparassus nigrovittatus, Olios nigrovittatus] JR08
    |    |         Prusias Pickard-Cambridge 1892 [incl. Ramnes Pickard-Cambridge 1892] JR08
    |    |           |--*P. nugalis Pickard-Cambridge 1892 JR08
    |    |           |--P. brasiliensis Mello-Leitão 1915 JR08
    |    |           |--P. lanceolatus Simon 1897 JR08
    |    |           `--P. semotus (Pickard-Cambridge 1892) (see below for synonymy) JR08
    |    |         Tibellomma Simon 1903 JR08
    |    |           `--*T. chazaliae (Simon 1898) [=Prusias chazaliae] JR08
    |    |         Paenula Simon 1897 JR08
    |    |           `--*P. paupercula Simon 1897 JR08
    |    |         Polybetes Simon 1896 JR08
    |    |         Adcatomus Karsch 1880 JR08
    |    |         Chrosioderma Simon 1897 JR08
    |    |           |--*C. albidum Simon 1897 JK05
    |    |           `--C. ranomafana Silva-Dávila 2005 JK05
    |    |         Dermochrosia Mello-Leitão 1940 JR08
    |    |         Sparianthina Banks 1929 JR08
    |    |           `--S. selenopoides Banks 1929 JR08
    |    |         Remmius Simon 1897 JR08
    |    |           |--*R. vultuosus Strand 1907 JK05
    |    |           `--R. badius Roewer 1961 JK05
    |    |         Quemedice Mello-Leitão 1942 JR08
    |    |           |--Q. enigmaticus Mello-Leitão 1942 FM11
    |    |           `--Q. piracuruca Rheims, Labarque & Ramírez 2008 FM11
    |    |         Anaptomecus Simon 1903 JR08
    |    |         Arandisa Lawrence 1938 JK05
    |    |           `--*A. deserticola Lawrence 1938 JK05
    |    |         Staianus Simon 1889 JK05
    |    |           `--*S. acuminatus Simon 1889 JK05
    |    |         Carparachne Lawrence 1962 JK05
    |    |           |--*C. alba Lawrence 1962 JK05
    |    |           `--C. aureoflava Lawrence 1966 JK05
    |    |         Cebrennus Simon 1880 JK05
    |    |           |--*C. wagae JK05
    |    |           `--C. aethiopicus Simon 1880 JK05
    |    |         Parapalystes Croeser 1996 JK05
    |    |           |--*P. euphorbiae Croeser 1996 JK05
    |    |           |--P. cultrifer (Pocock 1900) JK05
    |    |           |--P. lycosinus (Pocock 1900) JK05
    |    |           `--P. whiteae (Pocock 1902) JK05
    |    |         Anchonastus Simon 1898 JK05
    |    |           |--*A. caudatus Simon 1898 JK05
    |    |           `--A. plumosus (Pocock 1899) JK05
    |    |         Barylestis Simon 1910 JK05
    |    |           |--*B. blaisei JK05
    |    |           |--B. fagei (Lessert 1929) [=Torania fagei] JK05
    |    |           |--B. montandoni Lessert 1929 [=Torania montandoni] JK05
    |    |           |--B. occidentalis (Simon 1887) [=Torania occidentalis] JK05
    |    |           |--B. scutatus (Pocock 1903) [=Torania scutata] JK05
    |    |           `--B. variatus (Pocock 1899) [=Torania variata] JK05
    |    |         Berlandia Lessert 1921 JK95
    |    |           |--*B. longipes Lessert 1921 JK05
    |    |           `--B. tenebricola Simon & Fage 1922 JK05
    |    |         Cerbalus Simon 1897 JK05
    |    |           |--C. ergensis JK05
    |    |           `--C. pulcherrimus (Simon 1880) (see below for synonymy) JK05
    |    |         Damastes Simon 1880 JK05
    |    |           `--*D. grandidieri JK05
    |    |         Gnathopalystes Rainbow 1899 JK05
    |    |           `--*G. ferox JK05
    |    |         Megaloremmius Simon 1903 JK05
    |    |           `--*M. leo Simon 1903 JK05
    |    |         Microrchestris Lawrence 1962 JK05
    |    |           |--*M. melanogaster Lawrence 1962 JK05
    |    |           `--M. scutatus Lawrence 1966 JK05
    |    |         Nisueta Simon 1880 JK05
    |    |           |--*N. quadrispilota Simon 1880 JK05
    |    |           `--N. kolosvaryi Caporiacco 1947 JK05
    |    |         Nonianus Simon 1885 JK05
    |    |           |--*N. pictus JK05
    |    |           `--N. unilateralis Strand 1908 J98
    |    |         Orchestrella Lawrence 1965 JK05
    |    |           `--*O. longipes JK05
    |    |         Panaretella Lawrence 1937 JK05
    |    |           |--*P. distincta (Pocock 1896) JK05
    |    |           |--P. immaculata Lawrence 1952 JK05
    |    |           |--P. minor Lawrence 1952 JK05
    |    |           |--P. scutata (Pocock 1902) JK05
    |    |           `--P. zuluana Lawrence 1937 JK05
    |    |         Pseudomicrommata Järvi 1914 JK05
    |    |           `--*P. longipes (Bösenberg & Strand 1895) JK05
    |    |         Sarotesius Pocock 1898 JK05
    |    |           `--*S. melanognathus Pocock 1898 JK05
    |    |         Rhitymna JK05
    |    |           |--R. hildebrandti Järvi 1914 JK05
    |    |           `--R. saccata Järvi 1912 JK05
    |    |         Stasinoides Berland 1922 JK05
    |    |           `--*S. aethiopica Berland 1922 JK05
    |    |         Irileka iridescens Hirst 1998 T13
    |    |         Zachria Koch 1875 H95
    |    |           |--Z. flavicoma Koch 1875 H95
    |    |           `--Z. oblonga H95
    |    |         Pandercetes Koch 1875 PVD10
    |    |           |--P. isopus S10
    |    |           `--P. peronianus (Walckenaer 1837) PVD10
    |    |--Deleninae [Deleneae] J05
    |    |    |--Delena Walckenaer 1837 M54
    |    |    |    `--D. cancerides Walckenaer 1837 M54
    |    |    `--Pediana Simon 1880 [=Polydamna Thorell 1881] H89
    |    |         |--*P. regina (Koch 1875) [=Heteropoda regina, *Polydamna regina, Polydora (l. c.) regina] H89
    |    |         |--‘Polydamna’ aurichelis Strand 1907 H89
    |    |         |--P. horni (Hogg 1896) [=Isopeda horni] H89
    |    |         |--P. mainae Hirst 1995 H95
    |    |         |--P. occidentalis Hogg 1902 H95
    |    |         `--P. tenuis Hogg 1903 H89
    |    |--Heteropodinae [Heteropodeae] J08
    |    |    |--Heteropoda J08
    |    |    |--Pseudopoda J08
    |    |    |--Yiinthi Davis 1994 J08
    |    |    |--Sinopoda J08
    |    |    |--Bhutaniella J08
    |    |    `--Martensopoda J08
    |    |--Eusparassinae M54
    |    |--Sparassinae RJ08
    |    `--Sparianthinae JK05
    |         |--Thelcticopis J05
    |         |--Pseudosparianthis Simon 1887 J05
    |         |--Sparianthis Simon 1880 J05
    |         |--Pleorotus Simon 1898 J05
    |         |    `--*P. braueri Simon 1898 JK05
    |         |--Rhacocnemis Simon 1897 J05
    |         |    `--*R. guttatus (Blackwall 1877) JK05
    |         |--Stipax Simon 1898 J05
    |         |    `--*S. triangulifer Simon 1898 JK05
    |         |--Thomasettia Hirst 1911 J05
    |         |    `--*T. seychellana Hirst 1911 JK05
    |         `--Stasina Simon 1877 JK05
    |              |--*S. vittata Simon 1877 JK05
    |              |--S. americana K92
    |              |--S. manicata Simon 1897 JK05
    |              |--S. nalandica Karsch 1892 K92
    |              `--S. paripes [=Themeropis paripes] K92
    |--Philodromidae D59
    |    |--Ebo D90
    |    |--Thanatus J98
    |    `--Philodromus JD-S07
    |--Thomisidae [Misumenoidae] BS09
    |    |  i. s.: Cymbachina Bryant 1933 PVD10
    |    |           `--*C. albobrunnea (Urquhart 1893) PVD10 [=Xysticus albobrunnea NS00]
    |    |         Saccodamus formivorus B96
    |    |         Runcinia JD-S07
    |    |           |--R. acuminata B96
    |    |           `--R. flavida JD-S07
    |    |         Synema Simon 1864 PVD10
    |    |           |--S. globosum (Fabricius 1775) K55
    |    |           `--S. suteri Dahl 1907 PVD10
    |    |         Heriaeus hirtus (Latreille 1819) K55
    |    |         Camaricus striatipes (Van Hass 1882) [=Platythomisus striatipes; incl. C. formosus Thorell 1887] T94
    |    |         ‘Pasithea’ Blackwall 1858 non Lea 1833 S69
    |    |         Erissoides argentinus Mello-Leitão 1931 M-L31
    |    |         Tmarus piger K01
    |    |         Bucranium L-M-HM11
    |    |         Tharpyna PVD10
    |    |           |--T. campestrata Koch 1874 E07
    |    |           `--T. munda Koch 1875 PVD10
    |    |         Cymbacha festiva Koch 1874 E07
    |    |         Amyciaea albomaculata ZS10, B96
    |    |         Pistius truncatus K01
    |    |         Rhynchognatha Thorell 1887 K92
    |    |           `--R. cinerascens Thorell 1887 K92
    |    |         Misumena M01
    |    |           |--M. nesiotis Simon 1899 S99
    |    |           |--M. plumbea [=Diaea plumbea] S99
    |    |           |--M. tricuspidata K01
    |    |           `--M. vatia Clerck 1757 M01
    |    |         Daradius Sim. 1886 K92
    |    |           |--D. bipunctatus (Thorell 1877) [=Pistius bipunctatus] K92
    |    |           `--D. callidus Thorell 1890 K92
    |    |         Platythomisus Doleschall 1859 T94
    |    |           |--*P. phryniformis Doleschall 1859 D59
    |    |           `--P. jucundus Thorell 1894 T94
    |    |         Synaema K01
    |    |           |--S. globosum [incl. S. globosum var. nigriventris Kulczyński 1901] K01
    |    |           |--S. japonicum K01
    |    |           `--S. ornatum K01
    |    |         Sidymella Strand 1942 PVD10
    |    |           |--S. angularis (Urquhart 1885) [=Sparassus angularis] NS00
    |    |           |--S. angulata (Urquhart 1885) PVD10 [=Sparassus angulatus NS00]
    |    |           |--S. benhami (Hogg 1910) PVD10
    |    |           |--S. lampei (Strand 1913) [=Sidyma lampei] J98
    |    |           |--S. lobata B96
    |    |           `--S. longipes (Koch 1874) PVD10
    |    |         Misumenops L-M-HM11
    |    |           |--M. asperatus GV09
    |    |           |--M. californicus P74
    |    |           |--M. gabrielensis P74
    |    |           |--M. lepidus MC02
    |    |           |--M. nepenthicola GV09
    |    |           `--M. rothi P74
    |    |         Diaea PVD10
    |    |         Ozyptila K55
    |    |         Monaeses JD-S07
    |    |         Xysticus K01
    |    |         Thomisops pupa JD-S07
    |    |         Pherecydes JD-S07
    |    |         Tibellus K01
    |    |           |--T. maritimus (Menge 1875) BBM02
    |    |           |--T. oblongus Walckenaer 1802 BH02
    |    |           `--T. parallelus K01
    |    |         Misumenoides formosipes E95
    |    |         Pselloninae S99
    |    |           |--Pterelas Simon 1899 S99
    |    |           |    `--*P. schauinslandi Simon 1899 S99
    |    |           `--Psellonus planus S99
    |    |--Borboropactus Simon 1884 [Borboropactidae] JD-S07
    |    `--+--Aphantochilinae JD-S07
    |       |--Bominae JD-S07
    |       |--Dietinae JD-S07
    |       |--Strophius FM11 [Strophiinae JD-S07]
    |       |--Stiphropodinae JD-S07
    |       |--Thomisus JD-S07
    |       `--Stephanopis Cambridge 1869 H09 (see below for synonymy)
    |            |--S. barbipes Koch & Keyserling 1890 KK90
    |            |--S. benhami Hogg 1909 H09
    |            |--S. longipes H09
    |            `--S. ornata B96
    |--Anyphaenidae JD-S07
    |    |  i. s.: Hibana similaris (Banks 1929) S02
    |    |         Osoriella rubella BR07
    |    |         Aysha FM11
    |    |--Malenellinae JD-S07
    |    |--Amaurobioides Pickard-Cambridge 1883 [Amaurobioididae, Amaurobioidinae] PVD10
    |    |    |--*A. maritima Pickard-Cambridge 1883 PVD10
    |    |    |--A. major Forster 1970 PVD10
    |    |    |--A. pallida Forster 1970 PVD10
    |    |    |--A. picuna Forster 1970 PVD10
    |    |    |--A. piscator Hogg 1909 PVD10
    |    |    |--A. pleta Forster 1970 PVD10
    |    |    `--A. pohara Forster 1970 PVD10
    |    `--Anyphaena Sundevall 1833 CH12 [Anyphaeninae JD-S07]
    |         |--*A. accentuata (Walckenaer 1802) CH12
    |         |--A. pacifica MC02
    |         |--A. pugil CH12
    |         |--A. sabina Koch 1866 K55
    |         |--A. taiwanensis Chen & Huang 2010 CH12
    |         |--A. velox B06
    |         `--A. wuyi Zhang, Zhu & Song 2005 CH12
    |--Corinnidae JD-S07
    |    |  i. s.: Supunna Simon 1897 PVD10
    |    |           |--S. funerea Simon 1896 PVD10
    |    |           `--S. picta (Koch 1873) B96
    |    |         Meriola cetiformis (Strand 1908) [=Trachelas cetiformis] J98
    |    |         Abapeba rioclaro Bonaldo 2000 FM11
    |    |         Falconina gracilis (Keyserling 1891) FM11
    |    |         Mazax FM11
    |    |         Paradiestus giganteus (Karsch 1880) FM11
    |    |         Xeropigo FM11
    |    |           |--X. camilae De Souza & Bonaldo 2007 FM11
    |    |           `--X. tridentiger (Pickard-Cambridge 1869) FM11
    |    |         Creugas L-M-HM11
    |    |         Myrmecotypus L-M-HM11
    |    |         Orthobula L-M-HM11
    |    |         Copa JD-S07
    |    |         Pseudocorinna JD-S07
    |    |         Hortipes Bosselaers & Ledoux 1998 JD-S07, J01
    |    |           `--H. merwei JD-S07
    |    |--Castianeira FM11 [Castianeirinae JD-S07]
    |    |--Trachelinae JD-S07
    |    `--Corinna Koch 1841 JD-S07 [Corinninae JD-S07]
    |         |--C. brunneipeltula Strand 1910 S10
    |         `--C. capito (Lucas 1856) FM11
    |--Liocranidae JD-S07
    |    |--Scotina palliardi (Koch 1881) K02
    |    |--Itatsina Kishida 1930 TYM08
    |    |--Cybaeodes [Cybaeodinae] JD-S07
    |    |    `--C. madidus JD-S07
    |    |--Mesiotelus Simon 1897 MG03
    |    |    |--M. kulczynskii Charitonov 1946 MG03
    |    |    `--M. tenuissimus (Koch 1866) K55
    |    |--Phrurolithus K55 [Phrurolithinae JD-S07]
    |    |    |--P. festivus (Koch 1835) K55
    |    |    |--P. flavitarsis (Lucas 1846) K55
    |    |    `--P. minimus Koch 1839 VPB11
    |    |--Liocranum Koch 1866 [Liocraninae] JD-S07
    |    |    |--L. concolor Kraus 1955 K55
    |    |    |--L. giersbergi Kraus 1955 K55
    |    |    |--L. rupicola (Walckenaer 1825) K55
    |    |    |--L. rutilans K55
    |    |    `--L. segmentatum K55
    |    `--Agroeca Westring 1861 MG03
    |         |--A. brunnea (Blackwall 1833) K02
    |         |--A. cuprea Menge 1873 BBM02
    |         |--A. dentigera RKD02
    |         |--A. dubiosissima (Strand 1908) [=Hilke dubiosissima] J98
    |         |--A. inopina V09
    |         |--A. lusatica (Koch 1875) BBM02
    |         |--A. ornata Banks 1892 PS08
    |         `--A. proxima (O.P.-Cambridge 1871) K02
    |--Salticidae [Attidae, Salticoidae, Saltigradae] JD-S07
    |    |  i. s.: Simaetha A02
    |    |           |--S. knowlesi Żabka 1994 DH10
    |    |           |--S. paetula (Keyserling 1882) A02
    |    |           `--S. tenuior (Keyserling 1882) T13
    |    |         Portia Karsch 1878 B08
    |    |           `--P. fimbriata E95
    |    |         Yllenus B08
    |    |           |--Y. arenarius Menge 1868 B08
    |    |           |--Y. elegans [=Attus elegans] S95
    |    |           |--Y. flavociliatus Simon 1895 S95
    |    |           `--Y. hamifer Simon 1895 S95
    |    |         Salticus B02
    |    |         Sitticus D90
    |    |           |--S. finschi (Koch 1879) PS08
    |    |           |--S. pubescens [=Attus pubescens] V09
    |    |           |--S. saltator (Cambridge 1868) BBM02
    |    |           `--S. truncorum (Linnaeus 1758) K55
    |    |         Eris militaris (Walckenaer 1837) TF98a
    |    |         Phanias MC02
    |    |         Menemerus K55
    |    |           |--M. albocinctus Koch & Keyserling 1890 KK90
    |    |           |--M. confusus M-RL88
    |    |           `--M. taeniatus (Koch 1867) K55
    |    |         Marpissa Koch 1846 PVD10
    |    |           |--M. arenaria Urquhart 1888 PVD10
    |    |           |--M. armifera Urquhart 1892 PVD10
    |    |           |--M. cineracea Urquhart 1891 PVD10
    |    |           |--M. leucophaea Urquhart 1888 PVD10
    |    |           |--M. marina (Goyen 1892) PVD10
    |    |           |--M. muscosa E95
    |    |           |--M. nemoralis Urquhart 1892 PVD10
    |    |           `--M. nivoyi E95
    |    |         Holoplatys Simon 1885 W09
    |    |           |--H. appressus (Powell 1873) PVD10
    |    |           |--H. desertina MC13
    |    |           `--H. planissima (Koch 1879) PVD10
    |    |         Opisthoncus necator B96
    |    |         Mopsus mormon (Keyserling 1881) B96
    |    |         Cosmophasis Simon 1901 PVD10
    |    |           |--C. archeyi Berland 1931 PVD10 [=Clynotis archeyi F64]
    |    |           |--C. baehrae T13
    |    |           |--C. bitaeniata B96
    |    |           `--C. micarioides (K. 1880) [=Amycus micarioides] R13
    |    |         Trite Simon 1885 PVD10
    |    |           |--*T. pennata S99
    |    |           |--T. auricoma (Urquhart 1886) PVD10
    |    |           |--T. binotata Simon 1899 S99
    |    |           |--T. herbigrada (Urquhart 1889) PVD10
    |    |           |--T. mustilina (Powell 1873) PVD10
    |    |           |--T. parvula (Bryant 1935) PVD10
    |    |           |--T. planiceps Simon 1899 [incl. T. minax (Powell 1873)] PVD10
    |    |           `--T. urvillei (de Dalmas 1917) PVD10
    |    |         Neon D90
    |    |           |--N. reticulatus RKD02
    |    |           `--N. valentulus Falconer 1912 K02
    |    |         Heliophanus K55
    |    |           |--H. apiatus Simon 1868 K55
    |    |           |--H. corsicus Simon 1871 K55
    |    |           |--H. cupreus (Walckenaer 1802) K55
    |    |           |--H. dubius K01
    |    |           |--H. flavipes (Hahn 1832) BBM02
    |    |           |--H. lineiventris Simon 1868 K55
    |    |           |--H. patagiatus [incl. H. patagiatus var. albolineata Kulczyński 1901] K01
    |    |           `--H. rufithorax Simon 1868 K55
    |    |         Pelegrina PS08
    |    |           |--P. aeneola MC02
    |    |           |--P. flaviceps (Kaston 1973) PS08
    |    |           `--P. flavipes (Peckham & Peckham 1888) PS08
    |    |         Phidippus B08
    |    |           |--P. audax MC02
    |    |           |--P. clarus MC02
    |    |           |--P. comatus MC02
    |    |           |--P. femoratus E95
    |    |           |--P. johnsoni (Peckham & Peckham 1883) B02
    |    |           `--P. regius Koch 1846 B08
    |    |         Homalattus White 1841 K92
    |    |           |--H. cingulatus Karsch 1892 K92
    |    |           |--H. laminatus Karsch 1892 K92
    |    |           `--H. reductus Karsch 1892 K92
    |    |         Linus Peckh. 1885 K92
    |    |           |--L. alticeps Poc. 1898 R13
    |    |           `--L. fimbriatus (Doleschall 1859) K92, D59 [=Salticus fimbriatus K92, Sinis fimbriatus K92]
    |    |         Dischiropus Karsch 1892 K92
    |    |           |--D. alticinctus Karsch 1892 K92
    |    |           `--D. calcaratus [=Plexippus calcaratus] K92
    |    |         Maevia K92
    |    |           |--M. albozonata K92
    |    |           `--M. inclemens R96
    |    |         Icius Simon 1873 K92
    |    |           |--I. discatus Karsch 1892 K92
    |    |           |--I. hamatus (Koch 1846) K55
    |    |           |--I. hartii K62
    |    |           |--I. nebulosus (Simon 1868) K55
    |    |           `--I. notabilis (Simon 1871) K55
    |    |         Plexippus Koch 1846 T94
    |    |           |--P. albolineatus Koch 1846 K92
    |    |           |--P. culicivorus (Doleschall 1859) [=Salticus culicivorus, Menemerus culicivorus] T94
    |    |           |--P. cumulatus Karsch 1892 K92
    |    |           |--P. daemeli S99
    |    |           |--P. paykulli (Aud. in Sav. 1827) [=Attus paykulli, Menemerus paykulli] K92
    |    |           |--P. stridulator Poc. 1898 R13
    |    |           |--P. sylvarus Urquhart 1892 PVD10
    |    |           `--P. validus M-RL88
    |    |         Hasarius Simon 1871 PVD10
    |    |           |--H. adansoni (Audouin 1826) PVD10 [=Attus adansonii K92]
    |    |           |--H. arcigerus Karsch 1892 K92
    |    |           |--H. garetti K. 1881 R13
    |    |           `--H. paykulli V09
    |    |         Leptorchestes mutilliodes (Lucas 1846) K55
    |    |         Evarcha B08
    |    |           |--E. arcuata (Clerk 1757) K55
    |    |           |--E. culicivora (Wesolowska & Jackson 2003) B08
    |    |           |--E. falcata RKD02
    |    |           |--E. jucunda (Lucas 1846) K55
    |    |           `--E. laetabunda RKD02
    |    |         Carrhotus K55
    |    |           |--C. oscitans (Poc. 1898) [=Eustirognathus oscitans] R13
    |    |           `--C. xanthogramma (Walckenaer 1825) K55
    |    |         Saitis Simon 1876 W08
    |    |           `--S. barbipes (Simon 1868) K55
    |    |         Thyene imperialis (Rossi 1846) K55
    |    |         Phlegra bresnieri (Lucas 1846) K55
    |    |         Neaetha membranosa (Simon 1868) K55
    |    |         Aelurillus K55
    |    |           |--A. aeruginosus (Simon 1871) PO08
    |    |           |--A. concolor Kulczyński 1901 K01
    |    |           |--A. conveniens (Pickard-Cambridge 1872) PO08
    |    |           |--A. festivus K01
    |    |           |--A. litera K55
    |    |           |    |--A. l. litera K55
    |    |           |    `--A. l. v-insignitus (Clerck 1757) K55
    |    |           `--A. politiventris (Pickard-Cambridge 1872) PO08
    |    |         Bianor Peckham & Peckham 1885 PVD10
    |    |           |--*B. maculatus (Keyserling 1883) PVD10
    |    |           |--B. albobimaculatus (Lucas 1846) K55
    |    |           `--B. compactus (Urquhart 1885) PVD10
    |    |         Metaphidippus flaviceps JVHN90
    |    |         Corythalia canosa TF98b
    |    |         Araneotanna Özdikman & Kury 2006 [=Tanna Berland 1938 non Distant 1905] OK06
    |    |           `--*A. ornatipes (Berland 1938) [=*Tanna ornatipes] OK06
    |    |         Zenodorus R13
    |    |           |--Z. orbiculatus (Keyserling 1881) DH10
    |    |           `--Z. variatus Poc. 1898 R13
    |    |         Bathippus macroprotopus Poc. 1898 R13
    |    |         Athamus whitmei Cambr. 1877 R13
    |    |         Sigytes diloris (K. 1881) [=Hasarius diloris] R13
    |    |         Euryattus nigriventris (K. 1881) [=Hasarius nigriventris] R13
    |    |         *Synemosyna formica Hentz 1846 [incl. Janus gibberosus Koch 1846] T94
    |    |         Ascalus Thorell 1894 T94
    |    |           |--*A. laeta [=Synemosyna laeta] T94
    |    |           |--A. niger [=Synemosyna nigra] T94
    |    |           `--A. pygmaeus Thorell 1894 T94
    |    |         Janigena melanocephala [=Janus melanocephalus] T94
    |    |         Toxeus T94
    |    |           |--T. mandibularis T94
    |    |           `--T. maxillosus [incl. Synemosyna procera, T. procerus] T94
    |    |         Pergasus Thorell 1894 T94
    |    |           `--*P. formosus [=Salticus formosus] T94
    |    |         Herilus Thorell 1894 T94
    |    |           |--*H. pectorosus [=Salticus pectorosus] T94
    |    |           |--H. alticeps [=Salticus alticeps] T94
    |    |           `--H. radiatus Thorell 1894 T94
    |    |         Sphasus S69
    |    |            |--S. gracilipes White 1849 W49
    |    |            |--S. indicus S69
    |    |            |--S. lepidus Blackw. 1864 S69
    |    |            |--‘Spasus’ signifer Doleschall 1859 D59
    |    |            |--S. similaris Stoliczka 1869 S69
    |    |            |--S. striatus Doleschall 1857 D57
    |    |            |--S. striolatus Doleschall 1859 D59
    |    |            `--S. viridanus Stoliczka 1869 S69
    |    |         Myrmarachne McLeay 1835 C08
    |    |           |--M. assimilis Banks 1930 C08
    |    |           |--M. lupata E95
    |    |           `--M. providens B03
    |    |         Psecas chapoda OMM08
    |    |         Coryphasia OMM08
    |    |         Eustiromastix nativo OMM08
    |    |         Paraplatoides Żabka 1992 W09
    |    |           |--*P. tenerrima (Koch 1879) [=Marptusa tenerrima] W09
    |    |           |--P. caledonica (Berland 1932) [=Holoplatys caledonica] W09
    |    |           `--P. darwini Waldock 2009 W09
    |    |         Zebraplatys Żabka 1992 W09
    |    |           |--Z. fractivittata (Simon 1909) DH10
    |    |           `--Z. keyserlingi Żabka 1992 DH10
    |    |         Paraphidippus aurantius BS09
    |    |         Xenocytaea Berry, Beatty & Prószyński 1998 P11
    |    |           |--*X. triramosa Berry, Beatty & Prószyński 1998 P11
    |    |           |--X. anomala Berry, Beatty & Prószyński 1998 P11
    |    |           |--X. daviesae Berry, Beatty & Prószyński 1998 P11
    |    |           |--X. maddisoni Berry, Beatty & Prószyński 1998 P11
    |    |           |--X. stanislawi Patoleta 2011 P11
    |    |           |--X. taveuniensis Patoleta 2011 P11
    |    |           |--X. victoriensis Patoleta 2011 P11
    |    |           |--X. vonavonensis Patoleta 2011 P11
    |    |           `--X. zabkai Berry, Beatty & Prószyński 1998 P11
    |    |         Cytaea P11
    |    |         Chalcotropis P11
    |    |         Donoessus P11
    |    |         Panysinus nicholsoni [=Hasarius nicholsoni] V09
    |    |         Peckhamia D90
    |    |         Habronattus D90
    |    |         Talavera petrensis D90, RKD02
    |    |         Pseudoeuophrys lanigera [=Euophrys lanigera] V09
    |    |         Cobanus mandibularis E95
    |    |         Pachyballus JD-S07
    |    |         Bacelarella Berland & Millot 1941 JD-S07, J02
    |    |           |--B. conjugans J02
    |    |           |--B. pavida Szûts & Jocqué 2001 J02
    |    |           `--B. tentativa Szûts & Jocqué 2001 J02
    |    |         Orsima JD-S07
    |    |         Viciria pavesii JD-S07
    |    |         Grayenulla MC13
    |    |           |--G. australensis Żabka 1992 DH10
    |    |           |--G. spinimana Żabka & Gray 1992 DH10
    |    |           `--G. waldockae Żabka 1992 DH10
    |    |         Asemonea tenuipes E95
    |    |         Goleba puella E95
    |    |         Jacksonoides queenslandica E95
    |    |         Pseudicius E95
    |    |         Thorellia ensifera E95
    |    |         Astieae Z95
    |    |           |--Megaloastia Żabka 1995 Z95
    |    |           |    `--*M. mainae Żabka 1995 Z95
    |    |           |--Arasia Z95
    |    |           |--Sondra Z95
    |    |           `--Helpis Simon 1901 Z95, PVD10
    |    |                `--*H. minitabunda (Koch 1880) PVD10
    |    |         Philaeus chrysops K01
    |    |         Oedipus aenescens K01
    |    |         Marptusa K01
    |    |           |--M. muscosa K01
    |    |           `--M. radiata K01
    |    |         Clynotis Simon 1901 F64
    |    |           |--C. albobarbatus MC13
    |    |           |--C. barresi Hogg 1909 F64
    |    |           |--C. knoxi Forster 1964 F64
    |    |           |--C. saxatilis (Urquhart 1886) PVD10
    |    |           `--C. severus DH10
    |    |         Ergane K01
    |    |           |--E. arcuata K01
    |    |           |--E. falcata K01
    |    |           |--E. laetabunda K01
    |    |           `--E. signata Koch & Keyserling 1890 KK90
    |    |         Attus PVD10
    |    |         Adoxotoma Simon 1909 PVD10
    |    |           `--A. forsteri Żabka 2004 PVD10
    |    |         Hinewaia Żabka & Pollard 2002 PVD10
    |    |           `--*H. embolica Żabka & Pollard 2002 PVD10
    |    |         Hypoblemum Peckham & Peckham 1885 PVD10
    |    |           `--H. albovittatum (Keyserling 1882) PVD10
    |    |         Jotus Koch 1881 PVD10
    |    |           `--J. ravus (Urquhart 1893) PVD10
    |    |         Laufeia Simon 1889 PVD10
    |    |           `--L. aerihirta (Urquhart 1888) PVD10
    |    |         Ocrisiona Simon 1901 PVD10
    |    |           |--*O. leucocomis (Koch 1879) PVD10
    |    |           |--O. cinerea (Koch 1879) PVD10
    |    |           `--O. yakatunyae Żabka 1990 DH10
    |    |         Cyrba DH10
    |    |         Damoetas DH10
    |    |         Gangus DH10
    |    |         Simaethula DH10
    |    |--Massagrinae JD-S07
    |    |--Marenginae JD-S07
    |    |--Spartaeinae JD-S07
    |    |--Holcolaetinae JD-S07
    |    |--Mogrinae JD-S07
    |    |--Lyssomanes TF98a [Lyssomaninae JD-S07]
    |    |    `--L. viridis E95
    |    |--Dendryphantes K01 [Dendryphantinae JD-S07]
    |    |    `--D. thorellii K01
    |    |--Euophryinae W13
    |    |    |--Maratus W13
    |    |    `--Euophrys V09
    |    |         |--E. frontalis (Walckenaer 1802) BH02
    |    |         |--E. parvula FF99
    |    |         `--E. westringi RKD02
    |    `--Pelleninae M54
    |         |--Habrocestum Simon 1876 M54
    |         |    `--H. punctiventris Keyserling 1883 M54
    |         `--Pellenes K55
    |              |--P. arcigerus (Walckenaer 1837) K55
    |              |--P. bitaeniata (Keyserling 1882) DH10
    |              `--P. simoni Koch 1882 PO08
    |--Gnaphosoidea JD-S07
    |    |  i. s.: Fissarena barrow Platnick 2002 T13
    |    |--Cithaeronidae JD-S07
    |    |    |--Cithaeron P.-Cambridge 1872 JD-S07
    |    |    `--Inthaeron Platnick 1991 JD-S07
    |    |--Trochanteriidae JD-S07
    |    |    |  i. s.: Platyoides JD-S07
    |    |    |           |--P. pictus JD-S07
    |    |    |           `--P. walteri JD-S07
    |    |    |--Morebilus T13 [Morebilinae JD-S07]
    |    |    |    `--M. diversus (Koch 1875) T13
    |    |    |--Trachycosminae JD-S07
    |    |    `--Trochanteria Karsch 1878 [Trochanteriinae] JD-S07
    |    |--Gallieniellidae JD-S07
    |    `--+--Lamponidae JD-S07
    |       |--Ammoxenidae JD-S07
    |       |    |--Rastellus Platnick & Griffin 1990 JD-S07
    |       |    |--Austrammo Platnick 2002 JD-S07
    |       |    |--Barrowammo Platnick 2002 JD-S07
    |       |    |    `--B. waldockae Platnick 2002 T13
    |       |    `--Ammoxenus Simon 1893 JD-S07
    |       |         `--A. amphalodes Dippenaar & Meyer 1980 HD-S02
    |       `--+--Gnaphosidae JD-S07
    |          `--Prodidomidae JD-S07
    |--Selenopidae [Selenopinae] JD-S07
    |    |--Anyphops Benoit 1968 JD-S07
    |    |--Hovops Benoit 1968 JD-S07
    |    |--Garcorops Cotronca 2003 JD-S07
    |    |    `--G. jadis JD-S07
    |    |--Selenops Latreille 1819 M54
    |    |    |--S. aissus B06
    |    |    `--S. australiensis Koch 1871 M54
    |    `--Karaops burbidgei Crews & Harvey 2011 T13
    `--Clubionidae [Clubionoidea] JD-S07
         |--Clubiona J98
         |--Phrurotimpus alarius JVHN90
         |--Wulfila ventralis Banks 1906 B06
         |--Clastes freycineti S10
         |--Scotinella D90
         |--Clubionoides D90
         |--Zatapina Karsch 1892 K92
         |    `--*Z. plumipes (Doleschall 1859) [=Delena plumipes, Tychicus plumipes] K92
         |--Onychocryptus Karsch 1892 K92
         |    `--*O. mutillarius Karsch 1892 K92
         |--Tychichus S10
         |    |--T. genitalis Strand 1910 S10
         |    `--T. longipes K92
         |--Prychia S10
         |    |--P. gracilis K. 1875 R13
         |    `--P. pallidula Strand 1910 S10
         |--Matidia Thorell 1878 CH12
         |    |--*M. virens (Thorell 1878) CH12
         |    `--M. spatulata Chen & Huang 2006 CH12
         `--Leptodrassus Simon 1878 K92
              |--*L. femineus K92
              |--L. scutatus K92
              `--L. seriatus Karsch 1892 K92

Cerbalus pulcherrimus (Simon 1880) [incl. C. angustifrons Denis 1960, C. sahariensis Jézéquel & Junqua 1966] JK05

Prusias semotus (Pickard-Cambridge 1892) [=*Ramnes semotus, Heteropoda simoni Bonnet 1957 (n. n.)] JR08

Stephanopis Cambridge 1869 H09 [=Stephanopsis (l. c.) nec Bedot 1896 nec Lambert 1900 H09; Stephanopseae, Stephanopinae JD-S07]

*Type species of generic name indicated

REFERENCES

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