Colini

Colus islandicus, copyright Femorale.


Belongs within: Buccinidae.
Contains: Chrysodomus (Chrysodomus), Colus (Aulacofusus).

The Colini is a group of whelks containing species carrying a leaf-shaped operculum with a terminal or subspiral nucleus.

<==Colini (see below for synonymy)
    |--Pyramimitra Conrad 1865 [Pyramimitridae] BR05
    |--Truncaria Adams & Reeve 1850 [Truncariinae] BR05
    |--Metajapelion Goryachev 1987 [Metajapelioninae] BR05
    |--Pomahakia Finlay 1927 F27b
    |    `--*P. aberrans Finlay 1927 [=Fusus plicatilis Hutton 1873 non Wood 1848] F27b
    |--Kelletia Bayle 1884 O27a
    |    `--*K. kelletii (Forbes 1850) [=Fusus kellettii] O27a
    |--Sulcosinus Dall 1894 O27a
    |    `--*S. taphrius (Dall 1891) [=Buccinum taphrium] O27a
    |--Searlesia Harmer 1914 O27a, AV03
    |    `--*S. costiferum (Wood 1848) [=Trophon costiferum] AV03
    |--Microcolus M54
    |    |--M. dunkeri M54
    |    `--M. lincolnensis M54
    |--Chrysodomus Swainson 1840 BR05
    |    |--C. (Chrysodomus) O27a
    |    `--C. (Sulcosipho Dall 1916) O27a
    |         `--C. (S.) tabulatus Baird 1863 O27a
    |--Macron Adams & Adams 1853 O27a
    |    |--M. aethiops O27a
    |    |    |--M. a. aethiops O27a
    |    |    `--*M. a. kellettii (Adams 1853) [=Pseudoliva kellettii] O27a
    |    `--M. lividus (Adams 1854) [=Pseudoliva livida] O27a
    |--Pyrulofusus Mörch 1869 O27a
    |    |--*P. deformis (Reeve 1847) [=Fusus deformis] O27a
    |    `--P. harpa Mörch 1858 O27a
    |         |--P. h. harpa O27a
    |         `--P. h. dexius Dall 1907 O27a
    |--Mohnia Friele 1879 O27a, BR05 [Mohniinae]
    |    |--*M. mohnii [=Fusus mohnii] O27a
    |    |--M. corbis Dall 1913 O27a
    |    |--M. exquisita Dall 1913 O27a
    |    |--M. frielei Dall 1891 O27a
    |    |--M. robusta Dall 1913 O27a
    |    |--M. siphonoidea Dall 1913 O27a
    |    `--M. vernalis Dall 1913 O27a
    |--Verconella Iredale 1914 P27 [=Penion (preoc.) F27b]
    |    |  i. s.: V. crawfordi (Hutton 1873) [=Fusus crawfordi] F27a
    |    |         V. inflata (Hutton 1873) [=Buccinum inflatum] F27a
    |    |--+--V. adusta P27
    |    |  |    |--V. a. adusta P27
    |    |  |    `--V. a. mandarinoides Powell 1927 P27
    |    |  `--V. mandarina [=Siphonalia mandarina] P27
    |    `--+--V. ormesi Powell 1927 P27
    |       `--+--*V. dilatata (Quoy & Gaimard 1833) P27, F27a [=Fusus dilatatus P27, *Penion dilatatus F27b]
    |          |    |--V. d. dilatata P27
    |          |    |--V. d. cuvieriana Powell 1927 P27
    |          |    |--V. d. rex Finlay 1927 F27b
    |          |    `--V. d. rotunda Powell 1927 P27
    |          `--V. elongata Powell 1927 P27
    |--Plicifusus Dall 1902 O27a
    |    |  i. s.: P. arcticus Philippi 1850 O27a
    |    |         P. iljinensis (Sinelnikova in Gladenkov, Sinelnikova & Titova 1984) [=Searlesia iljinensis] AV03
    |    |         P. johanseni Dall 1919 O27a
    |    |         P. verkruzeni Kobelt 1876 O27a
    |    |--*P. (Plicifusus) kroyeri (Möller 1842) [=Sipho kroyeri, Fusus kroyeri] O27a
    |    |--P. (Helicofusus Dall 1916) O27a
    |    |    `--P. (H.) laticordatus Dall 1907 O27a
    |    |--P. (Latifusus Dall 1916) O27a
    |    |    `--P. (L.) griseus Dall 1877 O27a
    |    |--P. (Microfusus Dall 1916) O27a
    |    |    `--P. (M.) brunneus (Dall 1877) [=Chrysodomus brunneus] O27a
    |    `--P. (Retifusus Dall 1916) O27a
    |         |--P. (R.) incisus Dall 1919 O27a
    |         |--P. (R.) oceanodromae Dall 1919 O27a
    |         `--P. (R.) virens (Dall 1877) [=Chrysodomus virens] O27a
    |--Neptunea Röding 1798 BR05
    |    |--N. alaskana (Dall 1902) [=Trophon (Neptunea) alaskanus] O27b
    |    |--N. antiquata (Linnaeus 1758) HP89
    |    |--N. arthritica Bernardi 1858 V89
    |    |    |--N. a. arthritica ZLK11
    |    |    `--N. a. cumingii ZLK11
    |    |--N. avalonensis (Dall 1902) [=Trophon (Neptunea) avalonensis] O27b
    |    |--N. bentleyi (Dall 1908) [=Trophon (Neptunea) bentleyi] O27b
    |    |--N. beringi (Dall 1902) [=Trophon (Neptunea) beringi] O27b
    |    |--N. callicerata (Dall 1919) [=Trophon (Neptunea) callicerata] O27b
    |    |--N. cepula (Sowerby 1880) [=Trophon (Neptunea) cepulus] O27b
    |    |--N. contraria C60
    |    |--N. cymata (Dall 1902) [=Trophon (Neptunea) cymatus] O27b
    |    |--N. disparilis (Dall 1891) [=Trophon (Neptunea) disparilis] O27b
    |    |--N. elegantula (Dall 1907) [=Trophon (Neptunea) elegantulus] O27b
    |    |--N. eucymata (Dall 1902) [=Trophon (Neptunea) eucymata] O27b
    |    |--N. ithitoma (Dall 1919) [=Trophon (Neptunea) ithitoma] O27b
    |    |--N. macouni (Dall 1910) [=Trophon (Neptunea) macouni] O27b
    |    |--N. multicostata (Eschscholtz 1829) [=Trophon (Neptunea) multicostatus] O27b
    |    |--N. orpheus (Gould 1849) [=Trophon (Neptunea) orpheus] O27b
    |    |--N. pacifica (Dall 1902) [=Trophon (Neptunea) pacificus] O27b
    |    |--N. peregrina (Dall 1902) [=Trophon (Neptunea) peregrinus] O27b
    |    |--N. rotundata (Dall 1902) [=Trophon (Neptunea) rotundatus] O27b
    |    |--N. scitula (Dall 1891) [=Trophon (Neptunea) scitulus] O27b
    |    |--N. smithi (Dall 1902) [=Trophon (Neptunea) smithi] O27b
    |    |--N. staphylina (Dall 1919) [=Trophon (Neptunea) staphylinus] O27b
    |    |--N. stilesi Smith 1968 BC01
    |    `--N. stuarti (Smith 1880) [=Trophon (Neptunea) stuarti] O27b
    `--Colus Röding 1798 BR05
         |  i. s.: C. bensoni A27
         |         C. delicatulus [=Verconella delicatula] F27a
         |         C. kaunhoweni Finlay 1927 [=Fusus bicinctus Kaunhowen 1898 non Hutton 1873] F27b
         |         C. solidus A27
         |         ‘Columbarium’ suteri Smith 1915 F27a
         |         C. teschi Finlay 1927 [=Latirus fusiformis Tesch 1915 non Hoernes & Auinger 1891] F27b
         |--*C. (Colus) islandicus [=Murex islandicus] O27a
         |--C. (Anomalosipho Dautzenberg & Fischer 1912) O27a
         |    |--C. (A.) adonis Dall 1919 O27a
         |    |--C. (A.) conulus Aurivillius 1885 O27a
         |    `--C. (A.) martensi Krause 1885 O27a
         |--C. (Aulacofusus) O27a
         |--C. (Latisipho Dall 1916) O27a
         |    |--C. (L.) aphelus Dall 1889 O27a
         |    |--C. (L.) clementinus Dall 1919 O27a
         |    |--C. (L.) dalmasius Dall 1919 O27a
         |    |--C. (L.) errones Dall 1919 O27a
         |    |--C. (L.) halibrectus Dall 1891 O27a
         |    |--C. (L.) halli Dall 1873 O27a
         |    |--C. (L.) hypolispus Dall 1891 O27a
         |    `--C. (L.) jordani Dall 1913 O27a
         `--C. (Limatofusus Dall 1918) O27a
              |--C. (L.) dimidiatus Dall 1919 O27a
              |--C. (L.) halimeris Dall 1919 O27a
              |--C. (L.) morditus Dall 1919 O27a
              |--C. (L.) pulcius Dall 1919 O27a
              |--C. (L.) severinus Dall 1919 O27a
              |--C. (L.) tahwitanus Dall 1918 O27a
              |--C. (L.) timetus Dall 1919 O27a
              |--C. (L.) trombinus Dall 1919 O27a
              `--C. (L.) trophius Dall 1919 O27a

Nomen nudum: Colus modestus A27

Colini [Chrysodomidae, Chrysodominae, Colidae, Colinae, Colusina, Neptuneidae, Neptuneinae, Neptuniidae, Neptuniinae, Verconellidae]

*Type species of generic name indicated

REFERENCES

[A27] Allan, R. S. 1927. The geology and palaeontology of the Lower Waihao Basin, south Canterbury, New Zealand. Transactions and Proceedings of the New Zealand Institute 57: 265–309.

[AV03] Amano, K., & G. J. Vermeij. 2003. Evolutionary adaptation and geographic spread of the Cenozoic buccinid genus Lirabuccinum in the North Pacific. Journal of Paleontology 77 (5): 863–872.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[C60] Cox, L. R. 1960. Gastropoda: general characteristics of Gastropoda. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I84–I169. Geological Society of America, and University of Kansas Press.

[F27a] Finlay, H. J. 1927a. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[F27b] Finlay, H. J. 1927b. New specific names for austral Mollusca. Transactions and Proceedings of the New Zealand Institute 57: 488–533.

[HP89] Harasewych, M. G., & R. E. Petit. 1989. The nomenclatural status and phylogenetic affinities of Syrinx aruanus Linné, 1758 (Prosobranchia: Turbinellidae). Nautilus 103 (2): 83–84.

[M54] Macpherson, J. H. 1954. Molluscs (sea shells and snails). In: Willis, J. M. (ed.) The Archipelago of the Recherche pp. 55–63. Australian Geographical Society: Melbourne.

[O27a] Oldroyd, I. S. 1927a. The Marine Shells of the West Coast of North America vol. II pt I. Stanford University Press: Stanford University (California).

[O27b] Oldroyd, I. S. 1927b. The Marine Shells of the West Coast of North America vol. II pt. II. Stanford University Press: Stanford University (California).

[P27] Powell, A. W. B. 1927. Variation of the molluscan genus Verconella with descriptions of new recent species. Transactions and Proceedings of the New Zealand Institute 57: 549–558.

[V89] Vermeij, G. J. 1989. Habitat and form of Crepidula grandis in Japan, with comments on habitat specialization in calyptraeid gastropods. Nautilus 103 (3): 89–91.

[ZLK11] Zou, S., Q. Li & L. Kong. 2011. Additional gene data and increased sampling give new insights into the phylogenetic relationships of Neogastropoda, within the caenogastropod phylogenetic framework. Molecular Phylogenetics and Evolution 61: 425–435.

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