Clathurella rava, copyright Bestshells.

Belongs within: Conoidea.
Contains: Awateria, Scobinella, Glyphostoma (Glyphostoma), Lienardia (Lienardia), Pleurotomoides, Nannodiella, Etrema (Etrema), Etrema (Etremopsis).

The Clathurellidae are a group of conoid gastropods with more or less fusiform shells, usually bearing strong, often cancellate sculpture (Bouchet et al. 2011).

Characters (from Bouchet et al. 2011): Shell small- to medium-sized (5–40, usually 10–20 mm), fusiform to broadly fusiform, subsutural ramp usually unclearly separated. Sculpture mostly strong, usually cancellate, subsutural fold lacking. Shell surface often microgranular. Apertural armature often well developed, in form of pleats and denticles on both inner and outer lips; no true columellar pleats. Siphonal canal well expressed, moderately long. Anal sinus on subsutural ramp, deep, often (sub)tubular. Protoconch typically multispiral, up to six whorls, generally smooth with keeled last whorls; when paucispiral, protoconch usually smooth or spirally striated, sometimes with remaining carination in last portion. Operculum always absent. Radula of hypodermic marginal teeth, usually with relatively small solid basal part, sometimes with weak barb at tip. Distinct ligaments absent.

<==Clathurellidae [Clathurellinae, Defrancianina, Defranciinae]
    |--Clathurella Carpenter 1857 PK11, BR05
    |    |--*C. rava (Hinds 1843) [=Clavatula rava] P66
    |    |--C. affinis Dall 1871 P66
    |    |--C. arctata (Reeve 1846) [=Pleurotoma arctata] H09
    |    |--C. aurea Carpenter 1857 P66
    |    |--C. curtisiana (Hedley 1922) BK11
    |    |--C. daedala (Reeve 1846) [=Pleurotoma daedala] H09
    |    |--C. debilis (Hinds 1843) [=Clavatula debilis] H09
    |    |--C. intercalaris (Carpenter 1856) [=Defrancia intercalaris] P66
    |    |--C. nigrotincta (Montrouzier 1872) PK11
    |    |--C. rigida (Hinds 1843) [=Clavatula rigida] P66
    |    |    |--C. r. rigida P66
    |    |    `--C. r. fuscoligata (Carpenter 1856) [=Mangelia fuscoligata] P66
    |    |--C. serrata (Carpenter 1856) [=Defrancia serrata] P66
    |    `--C. thespesia (Melvill & Standen 1896) [=Daphnella thespesia] H09
    `--+--Nannodiella PK11
       `--Etrema Hedley 1918 PK11, BK11
            |  i. s.: E. tenera (Hedley 1899) BK11
            |--E. (Etrema) P66
            |--E. (Etremopa Oyama 1953) P66
            |    |--E. (*E.) subauriformis (Smith 1879) [=Drillia subauriformis] P66
            |    |--E. (E.) gainesii (Pilsbry 1895) [=Clathurella gainesii; incl. E. sawanensis Yokoyama 1927] P66
            |    |--E. (E.) nassoides (Reeve 1843) [=Pleurotoma nassoides] P66
            |    |--E. (E.) polydesma Hedley 1922 P66
            |    |--E. (E.) saigoensis Makiyama 1927 P66
            |    |--E. (E.) scalarina (Deshayes 1863) [=Pleurotoma scalarina] P66
            |    |--E. (E.) streptonotus (Pilsbry 1904) [=Drillia streptonotus] P66
            |    `--E. (E.) texta (Dunker 1860) [=Defrancia texta] P66
            `--E. (Etremopsis) BK11

Clathurellidae incertae sedis:
  Pleuropyramis Vredenburg 1921 P66
    `--*P. quinqueangularis (Vredenburg 1921) [=Mangillia (Clathurella) quinqueangularis] P66
  Domenginella Vokes 1939 P66
    `--*D. claytonensis (Gabb 1864) [=Turris claytonensis] P66
  Vaughanites Woodring 1928 P66
    `--*V. leptus Woodring 1928 P66
  Eoscobinella Powell 1942 P66
    `--*E. tahuia Powell 1942 P66
  Mitratoma Olsson 1930 P66
    `--*M. bartschi (Olsson 1930) [=Scobinella (*Mitratoma) bartschi] P66
  Endiatoma Cossmann 1896 P66
    `--*E. quadricincta (Cossmann 1883) [=Aphanitoma quadricincta] P66
  Nicolia Gregorio 1880 P66
    `--*N. solitaria (Gregorio 1880) [=Borsonia (*Nicolia) solitaria] P66
  Plentaria Harris 1937 P66
    |--*P. plenta (Aldrich & Harris 1895) [=Pleurotoma (Borsonia) plenta] P66
    `--P. horrida Harris 1937 [=P. plenta horrida] P66
  Cryptoborsonia Powell 1944 P66
    |--*C. pleurotomella Powell 1944 P66
    `--C. rugobela Powell 1944 P66
  Protosurcula Casey 1904 P66
    |--*P. gabbii (Conrad 1865) [=Surcula gabbii] P66
    |--P. platysoma (Heilprin 1880) [=Pleurotoma platysoma] P66
    `--P. tenuirostris Casey 1904 P66
  Eucheilodon Gabb 1860 P66
    |--*E. reticulata Gabb 1860 P66
    |--E. crenocarinata Heilprin 1880 P66
    |--E. gabbianum Casey 1904 P66
    `--E. reticulatoides (Harris 1895) [=Pleurotoma (Eucheilodon) reticulatoides] P66
  Moniliopsis Conrad 1865 P66
    |--*M. elaborata (Conrad 1833) [=Pleurotoma elaborata] P66
    |--M. hammetttensis (Harris 1937) [=Scobinella (Moniliopsis) hammettensis] P66
    |--M. nassiformis (Harris 1937) [=Scobinella (Moniliopsis) nassiformis] P66
    `--M. sculpturata (Aldrich 1911) [=Scobinella sculpturata] P66
  Glyptotoma Casey 1904 P66
    |--*G. crassiplicata (Gabb 1860) [=Pleurotoma crassiplicata, Scobinella crassiplicata] P66
    |    |--G. c. crassiplicata P66
    |    `--G. c. montgomeryensis (Harris 1937) [=Bathytoma (Glyptotoma) montgomeryensis] P66
    |--G. conradiana (Aldrich 1895) [=Borsonia (Scobinella) conradiana] P66
    |--G. fisherana (Harris 1937) [=Bathytoma (Glyptotoma) fisherana] P66
    |--G. parvula Casey 1904 P66
    `--G. rhombica Cooke 1928 P66
  Awateria P66
  Scobinella P66
  Comarmondia Monterosato 1844 BK11 (see below for synonymy)
    `--*C. gracilis (Montagu 1803) (see below for synonymy) P66
  Glyphostoma Gabb 1872 BK11
    |  i. s.: G. aliceae Melvill & Standen 1893 H09
    |         G. conradiana Gabb 1869 O27
    |         G. costicrenata (Cossmann 1900) P66
    |         G. cymodoce Dall 1919 O27
    |         G. huberti Sowerby 1893 H09
    |         G. polynesiense Reeve 1846 H09
    |         G. rostrata Sysoev & Bouchet 2001 BK11
    |         G. strombillum Hervier 1895 H09
    |         G. subcosticrenata MacNeil 1960 P66
    |         G. tricolor (Brazier 1876) [=Clathurella tricolor] H09
    |         G. vultuosum Reeve 1846 H09
    |--G. (Glyphostoma) P66
    |--G. (Euglyphostoma Woodring 1970) BK11
    `--G. (Glyphostomops Bartsch 1934) P66
         |--G. (*G.) hendersoni Bartsch 1934 P66
         |--G. (G.) oenoa (Bartsch 1934) [=Glyphostomops oenoa] P66
         `--G. (G.) pinellasensis (Fargo 1953) [=Glyphostomops pinellasensis] P66
  Lienardia Jousseaume 1884 BK11, P66
    |  i. s.: L. chrysoleuca Melvill 1923 [incl. Glyphostoma permiscere Usticke 1969] BC01
    |         L. globulosa Hervier 1896 BD12
    |         L. jousseaumei (Hervier 1896) BK11
    |         L. nigrotincta (Montrouzier 1872) BK11
    |--L. (Lienardia) P66
    `--L. (Thetidos Hedley 1899) P66
         `--L. (*T.) morsura (Hedley 1899) [=*Thetidos morsura] P66
  Strombinoturris Hertlein & Strong 1951 BK11
    `--*S. crockeri Hertlein & Strong 1951 P66
  Acrista Hedley 1922 BK11
    |--*A. punctilla (Hedley 1922) [=Lienardia (*Acrista) punctilla] P66
    |--A. biplicata (Melvill 1906) [=Mangilia biplicata, Lienardia (Acrista) biplicata] P66
    |--A. disconica (Hervier 1895) [=Glyphostoma disconicum, Lienardia (Acrista) disconica] P66
    |--A. fatima (Thiele 1925) [=Bellardiella fatima, Lienardia (Acrista) fatima] P66
    |--A. gaidei (Hervier 1895) [=Glyphostoma gaidei, Lienardia (Acrista) gaidei] P66
    |--A. marchei (Jousseaume 1884) [=Lienardia (Acrista) marchei] P66
    |    |--A. m. marchei P66
    |    `--A. m. tuberculifera (Hervier 1896) (see below for synonymy) P66
    `--A. semilineata (Garrett 1873) [=Clathurella semilineata, Lienardia (Acrista) semilineata] P66
  Adanaclava Bartsch 1950 BK11
    `--*A. adana Bartsch 1950 [=Crassispira (*Adanaclava) adana] P66
  Clathromangelia Monterosato 1884 BK11
    |--C. quadrillum (Dujardin 1837) (see below for synonymy) P66
    |--C. clathrata (Marcel de Serres 1829) [=Pleurotoma clathrata] P66
    |--C. crebricostata (Carpenter 1865) [=Mangilia (Clathromangilia) crebricostata] O27
    |--C. duperrayi (Peyrot 1903) [=Mangilia duperrayi] P66
    |--C. interlirata (Stearns 1871) [=Mangilia (Clathromangilia) interlirata] O27
    |--C. levidensis (Carpenter 1865) [=Mangilia (Clathromangilia) levidensis] O27
    |--C. lineolata (Reeve 1846) [=Mangilia (Clathromangilia) lineolata] O27
    `--C. rhyssa (Dall 1919) [=Mangilia (Clathromangilia) rhyssa] O27
  Crockerella Hertlein & Strong 1951 P66
    |--*C. crystallina (Gabb 1865) P66 (see below for synonymy)
    |--C. hilli Hertlein & Strong 1951 P66
    |--C. pederseni Hertlein & Strong 1951 P66
    `--C. subdiaphana (Carpenter 1864) P66 [=Mangelia subdiaphana P66, Cytharella subdiaphana D19]
  Euclathurella Woodring 1928 BK11
    |--*E. vendryesiana (Dall 1896) [=Clathurella vendryesiana] P66
    `--E. gertrudis (Toula 1909) [=Pleurotoma (Genota) gertrudis] P66
  Paraclathurella Boettger 1895 BK11
    |--*P. gracilenta (Reeve 1843) (see below for synonymy) P66
    |--P. aditicola Hedley 1922 P66
    |--P. clothonis Hedley 1922 P66
    |--P. densegranosa (Thiele 1925) [=Mangelia (Paraclathurella) densegranosa] P66
    |--P. padangensis (Thiele 1925) [=Mangelia (Paraclathurella) padangensis] P66
    `--P. thecla (Thiele 1925) [=Mangelia (Paraclathurella) thecla] P66
  Pseudoetrema Oyama 1953 BK11
    |--*P. fortilirata (Smith 1879) [=Drillia fortilirata] P66
    |--P. gerthi (Oostingh 1938) [=Clathurella gerthi] P66
    |--P. hukusimensis (Nomura & Zinbo 1935) [=Mangilia (Etrema) hukusimensis] P66
    |--P. ijzermani (Oostingh 1938) [=Clathurella ijzermani] P66
    |--P. kaloedanensis (Oostingh 1938) [=Clathurella kaloedanensis] P66
    `--P. sintikuensis (Nomura 1935) [=Lienardia (Etrema) sintikuensis] P66
  Turrella Laseron 1954 BK11
    |--*T. tenuilirata (Angas 1871) P66 (see below for synonymy)
    |--T. asperrima Laseron 1954 P66
    |--T. crassa Laseron 1954 P66
    |--T. gracilis Laseron 1954 P66
    |--T. granulosissima (Tenison-Woods 1879) P66 (see below for synonymy)
    |--T. letourneauxiana (Crosse & Fischer 1865) [=Pleurotoma letourneauxiana] P66
    |--T. morologus (Hedley 1922) [=Guraleus morologus] P66
    `--T. subcostata Laseron 1954 P66
  Pleurotomoides P66

Acrista marchei tuberculifera (Hervier 1896) [=Glyphostoma tuberculifera, Lienardia (Acrista) marchei tuberculifera] P66

Clathromangelia quadrillum (Dujardin 1837) [=Pleurotoma quadrillum; incl. P. granum Philippi 1844, *Clathromangelia granum, P. rude Philippi 1836 non P. rudis Sowerby 1834] P66

Comarmondia Monterosato 1844 BK11 [=Bellardia Bucquoy, Dautzenberg & Dollfus 1883 nec Allioni 1785 (ICBN) nec Robineau-Desvoidy 1863 P66, Bellardiella Fischer 1883 P66, Bellatula Strand 1929 BK11]

*Comarmondia gracilis (Montagu 1803) [=Murex gracilis, *Bellardia gracilis, *Bellardiella gracilis, *Bellatula gracilis; incl. Fusus branscombi Clark 1849, Murex emarginatus Donovan 1804, M. oblongus Brocchi 1814] P66

*Crockerella crystallina (Gabb 1865) P66 [=Clathurella crystallina P66, Philbertia crystallina D19; incl. C. lowei Dall 1903 P66]

*Paraclathurella gracilenta (Reeve 1843) [=Pleurotoma gracilenta; incl. Pl. contracta Reeve 1843, Cythara elegantissima Melvill & Standen 1903, Pl. fusoides Reeve 1846, Pl. (Drillia) portia Smith 1884] P66

Turrella granulosissima (Tenison-Woods 1879) P66 [=Clathurella granulosissima P66, Guraleus granulosissimus L54, Mangelia granulosissima L54]

*Turrella tenuilirata (Angas 1871) P66 [=Clathurella tenuilirata P66, Guraleus tenuiliratus L54, Pseudoraphitoma tenuilirata L54, Turrella multilirata (l. c.) L54]

*Type species of generic name indicated


[BD12] Bandel, K., & D. T. Dockery III. 2012. Protoconch characters of Late Cretaceous Latrogastropoda (Neogastropoda and Neomesogastropoda) as an aid in the reconstruction of the phylogeny of the Neogastropoda. Freiberger Forschungshefte C542 (psf 20): 93–128.

[BK11] Bouchet, P., Y. I. Kantor, A. Sysoev & N. Puillandre. 2011. A new operational classification of the Conoidea (Gastropoda). Journal of Molluscan Studies 77: 273–308.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BC01 Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[D19] Dall, W. H. 1919. Descriptions of new species of mollusks of the family Turritidae from the west coast of America and adjacent regions. Proceedings of the United States National Museum 56 (2288): 1–86, pls 1–24.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[L54] Laseron, C. F. 1954. Revision of the New South Wales Turridae (Mollusca). Royal Zoological Society of New South Wales: Sydney.

[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. II pt I. Stanford University Press: Stanford University (California).

[P66] Powell, A. W. B. 1966. The molluscan families Speightiidae and Turridae: an evaluation of the valid taxa, both Recent and fossil, with lists of characteristic species. Bulletin of the Auckland Institute and Museum 5: 1–184, pls 1–23.

[PK11] Puillandre, N., Y. I. Kantor, A. Sysoev, A. Couloux, C. Meyer, T. Rawlings, J. A. Todd & P. Bouchet. 2011. The dragon tamed? A molecular phylogeny of the Conoidea (Gastropoda). Journal of Molluscan Studies 77: 259–272.

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