Acteon punctocoelata, copyright J. Goddard.

Belongs within: Opisthobranchia.

The Acteonidae are a group of marine gastropods of uncertain affinities. Though anatomically similar to opisthobranchs, they possess certain plesiomorphic features such as the presence of an operculum to close the shell aperture. The shell is well calcified with a distinct spire and an elongate aperture.

<==Acteonidae [Pupacea, Pupidae, Tornatelladae, Tornatellidae, Tornatellinae]
    |  i. s.: Dactylus Schumacher 1817 BR05
    |           |--D. maurus F66
    |           `--D. mustelinus F66
    |         Microglyphis Dall 1902 O27
    |           |--*M. curtulus [=Acteon curtulus] O27
    |           |--M. breviculus Dall 1902 O27
    |           |--M. estuarinus Dall 1908 O27
    |           `--M. globularis (Smith 1875) [=Admete globularis] PH90
    |--Liocarenus Harris & Burrows 1891 [Liocareninae] BR05
    `--Acteoninae [Actaeoninae] BR05
         |--Tornatella Lamarck 1816 BR05
         |--Solidula Fischer von Waldheim 1807 [Solidulidae] BR05
         |--Nucleopsis Conrad 1865 [Nucleopsinae] BR05
         |--Tornatellaea Conrad 1860 [Tornatellaeinae] BR05
         |--Pupa Röding 1798 BR05
         |    |--P. admodesta Mighels 1845 J49
         |    |--P. affinis (Adams 1855) (see below for synonymy) F27b
         |    |--P. coccinata (Reeve 1842) [=Tornatella coccinata] H09
         |    |--P. nitidula (Lamarck 1822) [=Tornatella nitidula] H09
         |    |--P. solidula (Linnaeus 1758) [=Bulla solidula] H09
         |    |--P. sulcata (Gmelin 1791) [=Voluta sulcata] H09
         |    `--P. suturalis (Adams 1855) [=Solidula suturalis] H09
         `--Acteon Montfort 1810 [=Actaeon (l. c.) non Rang 1829] BR05
              |  i. s.: A. affinis (Sowerby 1836) [=Tornatella affinis] F27a
              |         A. ambiguus (Hutton 1885) [=Admete ambigua] PH90
              |         A. cratericulatus Hedley 1906 [=A. craticulatus Murdoch & Suter 1906] F27a
              |         A. praestitus Finlay 1924 [=A. sulcatus (Hutton 1885) (preoc.)] F27b
              |         A. roseus Hedley 1906 F27a
              |         A. tornatilis C60
              |         A. traski Stearns 1898 O27
              |--*A. (Acteon) fasciatus [=Tornatella fasciata] O27
              `--A. (Rictaxis Dall 1871) O27
                   |--A. (R.) painei Dall 1903 O27
                   `--A. (R.) punctocoelata (Carpenter 1864) [=Tornatella punctocoelata] O27
                        |--A. p. punctocoelata O27
                        `--A. p. vancouverensis Oldroyd 1927 O27

Pupa affinis (Adams 1855) nec Rossmaessler 1839 nec Aradas & Maggiore 1843 (neither preoc. as dif. homonymous gen.) [=Solidula affinis non Tornatella affinis Sowerby 1836 (not preoc. if in dif. gen.), Actaeon pilsbryi Cossmann 1902] F27c

*Type species of generic name indicated


[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[C60] Cox, L. R. 1960. Gastropoda: general characteristics of Gastropoda. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I84–I169. Geological Society of America, and University of Kansas Press.

[F27a] Finlay, H. J. 1927a. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[F27b] Finlay, H. J. 1927b. New specific names for austral Mollusca. Transactions and Proceedings of the New Zealand Institute 57: 488–533.

[F66] Fraser, L. 1866. Communication of a list of mollusks collected by R. Swinhoe, Esq., in Formosa. Proceedings of the Zoological Society of London 1866: 146.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[J49] Johnson, R. I. 1949. Jesse Wedgwood Mighels with a bibliography and a catalogue of his species. Occasional Papers on Mollusks 1 (14): 213–231.

[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. II pt I. Stanford University Press: Stanford University (California).

[PH90] Petit, R. E. & M. G. Harasewych. 1990. Catalogue of the superfamily Cancellarioidea Forbes and Hanley, 1851 (Gastropoda: Prosobranchia). Nautilus Supplement 1: 1–69.

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