Acteon punctocoelata, copyright J. Goddard.

Belongs within: Acteonimorpha.

The Acteonidae are a group of marine gastropods of uncertain affinities. Though anatomically similar to opisthobranchs, they possess certain plesiomorphic features such as the presence of an operculum to close the shell aperture. The shell is well calcified with a distinct spire and an elongate aperture. The radula usually lacks a rachidian tooth and possesses from five to over two hundred laterals each side.

<==Acteonidae [Pupacea, Pupidae, Tornatelladae, Tornatellidae, Tornatellinae]
    |--Liocarenus Harris & Burrows 1891 [Liocareninae] BR05
    |    `--*L. conovuliformis (Deshayes 1824) [=Auricula conovuliformis] BR17
    `--Acteoninae [Actaeoninae] BR05
         |--Nucleopsis Conrad 1865 [Nucleopsinae] BR05
         |    `--*N. subvaricata (Conrad 1860) [=Acteonina subvaricata] BR17
         |--Tornatellaea Conrad 1860 [Tornatellaeinae] BR05
         |    |--*T. bella Conrad 1860 BR17
         |    |--T. fontis (Duméril 1806) TTE93
         |    `--T. heberti TTE93
         |--Tornatella Lamarck 1816 BR05
         |    |--*T. flammea (Bruguière 1789) [=Buccinum flammeum] BR17
         |    |--T. bifasciata G20
         |    |--T. minuta Petterd 1879 P79
         |    `--T. ovalis H86
         |--Acteon Montfort 1810 [=Actaeon (l. c.) non Rang 1829] BR05
         |    |  i. s.: A. affinis (Sowerby 1836) [=Tornatella affinis] F27a
         |    |         A. ambiguus (Hutton 1885) [=Admete ambigua] PH90
         |    |         A. cratericulatus Hedley 1906 P61 [=A. craticulatus Murdoch & Suter 1906 F27a]
         |    |         A. panamensis Dall 1908 PP78
         |    |         A. praestitus Finlay 1924 [=A. sulcatus (Hutton 1885) (preoc.)] F27b
         |    |         A. roseus Hedley 1906 F27a
         |    |         A. tenellus N79
         |    |         A. traski Stearns 1898 O27
         |    |--A. (Acteon) O27
         |    |    |--*A. (A.) tornatilis (Linnaeus 1758) [=Bulla tornatilis] BR17
         |    |    `--A. (A.) fasciatus [=Tornatella fasciata] O27
         |    `--A. (Rictaxis Dall 1871) O27
         |         |--A. (R.) painei Dall 1903 O27
         |         `--A. (R.) punctocoelata (Carpenter 1864) [=Tornatella punctocoelata] O27
         |              |--A. p. punctocoelata O27
         |              `--A. p. vancouverensis Oldroyd 1927 O27
         `--Pupa Röding 1798 [=Solidula Fischer von Waldheim 1807; Solidulidae] BR05
              |--*P. solidula (Linnaeus 1758) P61, H09 (see below for synonymy)
              |--P. admodesta Mighels 1845 J49
              |--P. affinis (Adams 1855) (see below for synonymy) F27b
              |--P. alba (Hutton 1873) P61
              |--P. armifera W79
              |--P. (Leucochila) chordata C64
              |--P. cinerea [incl. Turbo quinquedentatus, Bulimus similis] G40
              |--P. claustralis S79
              |--P. coccinata (Reeve 1842) H09 (see below for synonymy)
              |--P. contracta W79
              |--P. cylindracea S79
              |--P. doliolum S79
              |--P. fallax [incl. P. parraiana] G79
              |--P. granum S79
              |--P. huttoni (Kirk 1882) P61
              |--P. kirki (Hutton 1873) P61
              |--P. nitidula (Lamarck 1822) [=Tornatella nitidula] H09
              |--P. pellucida G79
              |--P. pentodon W79
              |--P. polydon S79
              |--P. sterrii Forster 1840 KM20
              |--P. strobeli [incl. P. strobeli var. scharffi] S79
              |--P. sulcata (Gmelin 1791) [=Voluta sulcata] H09
              |--P. suturalis (Adams 1855) [=Solidula suturalis] H09
              |--P. triplicata S79
              |--P. uva G20
              `--P. variabilis S79

Acteonidae incertae sedis:
  Dactylus Schumacher 1817 BR05
    |--D. maurus F66
    `--D. mustelinus F66
  Microglyphis Dall 1902 O27
    |--*M. curtulus [=Acteon curtulus] O27
    |--M. breviculus Dall 1902 O27
    |--M. estuarinus Dall 1908 O27
    `--M. globularis (Smith 1875) [=Admete globularis] PH90
  Bullinula Swainson 1840 P61
    `--*B. lineata (Gray 1825) P61 (see below for synonymy)
  Neactaeonina Thiele 1912 P61
    |--*N. cingulata [=Acteonina cingulata] P61
    `--N. inexpectata Dell 1956 P61

*Bullinula lineata (Gray 1825) P61 [=Bulla lineata P61, Bullina lineata BR17; incl. Bulla scabra Gmelin 1791 (preoc.) BR17, F27a, Bullina scabra F27a, Voluta ziczac Muhlfeldt 1878 non Schroeter 1804 F27a]

Pupa affinis (Adams 1855) nec Rossmaessler 1839 nec Aradas & Maggiore 1843 (neither preoc. as dif. homonymous gen.) [=Solidula affinis non Tornatella affinis Sowerby 1836 (not preoc. if in dif. gen.), Actaeon pilsbryi Cossmann 1902] F27b

Pupa coccinata (Reeve 1842) H09 [=Tornatella coccinata H09, Buccinulus coccinatus B79, Solidula coccinata B79]

*Pupa solidula (Linnaeus 1758) P61, H09 [=Bulla solidula H09, *Solidula solidula BR17, Voluta solidula P61]

*Type species of generic name indicated


[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[B79] Brazier, J. 1879. List of marine shells collected on Fitzroy Island, north coast of Australia; with notes on their geographical range. Journal of Conchology 2: 186–199.

[C64] Carpenter, P. P. 1864. Supplementary report on the present state of our knowledge with regard to the Mollusca of the west coast of North America. Report of the British Association for the Advancement of Science 33: 517–686.

[F27a] Finlay, H. J. 1927a. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[F27b] Finlay, H. J. 1927b. New specific names for austral Mollusca. Transactions and Proceedings of the New Zealand Institute 57: 488–533.

[F66] Fraser, L. 1866. Communication of a list of mollusks collected by R. Swinhoe, Esq., in Formosa. Proceedings of the Zoological Society of London 1866: 146.

[G79] Gibbons, J. S. 1879. Notes on the habits and distribution, &c., of certain W. Indian Pulmonifera. Journal of Conchology 2: 129–134.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[G40] Gray, J. E. 1840. A Manual of the Land and Fresh-water Shells of the British Islands, with figures of each of the kinds. By William Turton, M.D. A new edition, thoroughly revised and much enlarged. Longman, Orme, Brown, Green, and Longmans: London.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[H86] Hutton, F. W. 1886. The Mollusca of the Pareora and Oamaru systems of New Zealand. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 205–237.

[J49] Johnson, R. I. 1949. Jesse Wedgwood Mighels with a bibliography and a catalogue of his species. Occasional Papers on Mollusks 1 (14): 213–231.

[KM20] Kury, A. B., A. C. Mendes, L. Cardoso, M. S. Kury & A. A. Granado. 2020. WCO-Lite: Online world catalogue of harvestmen (Arachnida, Opiliones). Version 1.0—Checklist of all valid nomina in Opiliones with authors and dates of publication up to 2018. Published by the author: Rio de Janeiro.

[N79] Norman, A. M. 1879. The Mollusca of the fiords near Bergen, Norway. Journal of Conchology 2: 8–77.

[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. 2 pt 1. Stanford University Press: Stanford University (California).

[PH90] Petit, R. E. & M. G. Harasewych. 1990. Catalogue of the superfamily Cancellarioidea Forbes and Hanley, 1851 (Gastropoda: Prosobranchia). Nautilus 103 (Suppl. 1): 1–69.

[P79] Petterd, W. F. 1879. New species of Tasmanian marine shells. Journal of Conchology 2: 102–105.

[PP78] Poorman, F. L., & L. H. Poorman. 1978. Additional molluscan records from Bahía de Los Angeles, Baja California Norte. Veliger 20 (4): 369–374.

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4th ed. Whitcombe and Tombs Limited: Christchurch.

[S79] Scharff, R. 1879. Land and freshwater shells collected in the alluvium of the Garonne, France. Journal of Conchology 2: 315–316.

[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.

[W79] Walker, B. 1879. Catalogue of the shell-bearing Mollusca of Michigan. Journal of Conchology 2: 325–337.

Last updated: 5 September 2021.

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