Rissoella diaphana, copyright Ennio Squizzatto.

Belongs within: Heterobranchia.
Contains: Acteonidae.

The Acteonimorpha are a clade of marine gastropods supported by molecular data. It unites the Rissoellidae, minute gastropods with transparent, conical shells, with the Acteonoidea, which have fusiform shells with an elongate aperture and a well-developed head-shield.

<==Acteonimorpha BR17
    |--Rissoellidae [Heterophrosynidae, Jeffresiidae, Jeffreysiidae, Rissoelloidea] BR17
    |    |--Cithna C64
    |    |    |--C. albida Carpenter 1864 C64
    |    |    `--C. tumens C64
    |    `--Rissoella Gray 1847 BR05 [=Jeffreysia Alder 1850 BR17]
    |         |--*R. diaphana (Alder 1848) [=Rissoa diaphana, *Jeffreysia diaphana] BR17
    |         |--‘Jeffreysia’ alderi C64
    |         |--‘Jeffreysia’ bifasciata C64
    |         |--R. contrerasi Rolán & Fernández 2004 F05
    |         |--R. flemingi HS01
    |         |--R. galba Robertson 1961 BC01
    |         |--R. globularis C60 [=Jeffreysia globularis N79]
    |         |--R. micra PC08
    |         |--‘Jeffreysia’ opalina N79
    |         |--R. ornata Simone 1995 S11
    |         |--R. rissoaformis HS01
    |         |--‘Jeffreysia’ translucens Carpenter 1864 C64
    |         `--R. verruculosa PP64
    `--Acteonoidea [Acteonacea, Acteonidea, Pleurocoela] BR05
         |--Acteonidae S-PS96
         |--Zardinella Bandel 1994 [Zardinellidae] BR05
         |    `--*Z. cingulata (Zardini 1978) [=Coelostylina cingulata] BR17
         |--Tubifer Piette 1856 BR05 [Tubiferacea, Tubiferidae BR17]
         |    `--*T. nudus Pette 1856 BR17
         |--Cylindrobullinidae [Cylindrobullininae, Cylindrobullinoidea] BR17
         |    |--Consobrinella consobrina B91, CM01
         |    |--Cylindrobullina Ammon 1878 BR05
         |    |    |--*C. fragilis (Dunker 1847) [=Tornatella fragilis] BR17
         |    |    `--C. convexa Batten & Stokes 1986 NE02
         |    `--Ceritellopsis Fischer 1961 B91
         |         |--C. parvula B91
         |         `--+--C. gramanni B91
         |            `--+--*C. complacita (Huckriede 1967) [=Ceritella (*Ceritellopsis) complacita] B91
         |               `--C. plicatula B91
         `--Aplustridae [Bullinadae, Bullinidae] BR17
              |--Palaeohydatina flouesti TTE93
              |--Hydatina Schumacher 1817 BR05
              |    `--H. physis (Linnaeus 1758) F27 (see below for synonymy)
              |--Nonacteonina Stephenson 1941 [Nonacteoninidae] BR05
              |    `--*N. graphoides Stephenson 1941 BR17
              |--Aplustrum Schumacher 1817 [=Amplustrum (l. c.); Amplustrina, Aplustrinae] BR05
              |    `--*A. fasciatum Schumacher 1817 BR17
              |--Sulcoactaeon Cossmann 1895 [Sulcoactaeonidae] BR05
              |    |--*S. striatosulcata (Zittel & Goubert 1861) [=Acteonina striatosulcata] BR17
              |    `--S. pulloides (Hudleston 1896) GK02
              `--Bullina Férussac 1822 BR17, BR05
                   |--B. amplustre (Linnaeus 1758) BR17, L58 [=Bulla amplustre BR17, Aplustrum aplustre H09]
                   |--B. undulata [=Bulla undulata] BR17
                   `--B. velum [=Bulla velum] BR17

Hydatina physis (Linnaeus 1758) F27 [=Bulla physis H09, Bullina physis BR17, *Hydatina filosa Schumacher 1817 BR17]

*Type species of generic name indicated


[B91] Bandel, K. 1991. Gastropods from brackish and fresh water of the Jurassic–Cretaceous transition (a systematic reevaluation). Berliner Geowissenschaftliche Abhandlungen, Reihe A 134: 9–55.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BC01] Boyko, C. B. & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[C64] Carpenter, P. P. 1864. Supplementary report on the present state of our knowledge with regard to the Mollusca of the west coast of North America. Report of the British Association for the Advancement of Science 33: 517–686.

[C60] Cox, L. R. 1960. Gastropoda: general characteristics of Gastropoda. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I84–I169. Geological Society of America, and University of Kansas Press.

[CM01] Cózar, P., & B. L. Mamet. 2001. Planohowchinia (Lasiodiscidae, Foraminiferida), a new Late Viséan genus, southwestern Spain. Journal of Foraminiferal Research 31 (3): 228–232.

[F05] Fernández, J. 2005. Noticia de nuevos táxones para la ciencia en el ámbito Íbero-Balear y Macaronésico. Nuevos táxones animales descritos en la península Ibérica y Macaronesia desde 1994 (IX). Graellsia 61 (2): 261–282.

[F27] Finlay, H. J. 1927. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[GK02] Gründel, J., & T. Kowalke. 2002. Palaeorissoinidae, a new family of marine and brackish water Rissooidea (Gastropoda, Littorinimorpha). Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 226 (1): 43–57.

[HS01] Hayward, B. W., A. B. Stephenson, M. S. Morley, W. M. Blom, H. R. Grenfell, F. J. Brook, J. L. Riley, F. Thompson & J. J. Hayward. 2001. Marine biota of Parengarenga Harbour, Northland, New Zealand. Records of the Auckland Museum 37: 45–80.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[L58] Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii: Holmiae.

[N79] Norman, A. M. 1879. The Mollusca of the fiords near Bergen, Norway. Journal of Conchology 2: 8–77.

[NE02] Nützel, A., & D. H. Erwin. 2002. Battenizyga, a new Early Triassic gastropod genus with a discussion of the caenogastropod evolution at the Permian/Triassic boundary. Paläontologische Zeitschrift 76 (1): 21–27.

[PP64] Peres, J. M., & J. Picard. 1964. Nouveau manuel de bionomie benthique de la mer Mediterranee. Recueil des Travaux de la Station Marine d'Endoume, Bulletin 31 (27): 5–137.

[PC08] Ponder, W. F., D. J. Colgan, J. M. Healy, A. Nützel, L. R. L. Simone & E. E. Strong. 2008. Caenogastropoda. In: Ponder, W. F., & D. L. Lindberg (eds) Phylogeny and Evolution of the Mollusca pp. 331–383. University of California Press: Berkeley.

[S-PS96] Salvini-Plawen, L. v., & G. Steiner. 1996. Synapomorphies and plesiomorphies in higher classification of Mollusca. In: Taylor, J. D. (ed.) Origin and Evolutionary Radiation of the Mollusca pp. 29–51. Oxford University Press: Oxford.

[S11] Simone, L. R. L. 2011. Phylogeny of the Caenogastropoda (Mollusca), based on comparative morphology. Arquivos de Zoologia 42 (4): 161–323.

[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.

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