Hemileiidae

Dorsal (left) and ventral views of Hemileius paraguayensis, from Balogh & Balogh (1992b).

Belongs within: Oripodoidea.
Contains: Hemileius (Hemileius), Hemileius (Tuberemaeus).

The Hemileiidae are a group of oripodoid mites with a sacculate octotaxic system and lacking pteromorphs (Balogh & Balogh 1992).

Characters (from Balogh & Balogh 1992a, as Hemileius group of Scheloribatidae): Octotaxic system with sacculi; pteromorphae absent. Usually four pairs of genital setae, exceptionally five; one pair of aggenital setae; two pairs of anal setae; three pairs of adanal setae.

<==Hemileiidae [Hemileinae] S04
    |--Constrictobates Balogh & Mahunka 1966 [Constrictobatinae] S04
    |    `--*C. lineolatus Balogh & Mahunka 1966 S04
    |--Mucrobates Balogh & Mahunka 1979 S04
    |    `--*M. fissuratus Balogh & Mahunka 1979 S04
    |--Plumobates Balogh & Mahunka 1966 S04
    |    `--*P. decoratus Balogh & Mahunka 1966 S04
    |--Zeascheloribates Luxton 1982 S04
    |    `--*Z. palustris Luxton 1982 S04
    |--Baobabula Mahunka 1975 S04
    |    |--*B. mussardi Mahunka 1975 S04
    |    `--B. impolita (Warburton 1912) [=Notaspis impolita] S04
    |--Monoschelobates Balogh & Mahunka 1969 S04
    |    |--*M. parvus Balogh & Mahunka 1969 S04
    |    `--M. translamellatus Pérez-Íñigo & Baggio 1991 S04
    |--Balazsella Mahunka 1983 S04
    |    |--*B. pilosetosa Mahunka 1983 S04
    |    |--B. ilhabellae Balogh & Palacios-Vargas 1996 S04
    |    `--B. mexicana Balogh & Palacios-Vargas 1996 S04
    |--Heteroleius Balogh & Mahunka 1966 S04
    |    |--*H. longissimus Balogh & Mahunka 1966 S04
    |    |--H. navicula (Berlese 1913) [=Oribatula navicula] S04
    |    `--H. planus (Aoki 1984) [=Symphauloppia plana] S04
    |--Nasozetes Sellnick 1930 [incl. Kinabaluella Mahunka 1966] S04
    |    |--*N. sumatrensis Sellnick 1930 S04
    |    |--N. choerognathus Willmann 1931 S04
    |    |--N. porcella (Mahunka 1996) [=Kinabaluella porcella] S04
    |    `--N. stunkardi Sengbusch 1957 S04
    |--Simkinia Krivolutsky 1966 S04
    |    |--*S. turanica Krivolutsky 1966 [incl. Hemileius ovalis Kulijev 1968] S04
    |    |--S. elongata Krivolutsky 1969 S04
    |    |--S. montana Krivolutsky & Grishina 1970 S04
    |    |--S. schachtachtinskoi (Kulijev 1961) [=Oribatula schachtachtinskoi] S04
    |    `--S. tianshanica Krivolutsky 1971 S04
    |--Siculobata Grandjean 1953 S04
    |    |--S. (Siculobata) [incl. Floribates Norton & Kethley 1989] S04
    |    |    |--*S. (S.) sicula (Berlese 1892) [=Oppia tibialis sicula] S04
    |    |    |    |-S. s. sicula S04
    |    |    |    `--S. s. platensis (Berlese 1916) [=Oribatula (Hemileius) platensis] S04
    |    |    |--S. (S.) florens (Berlese 1908) [=Oribatula florens] S04
    |    |    `--S. (S.) malabarica Ramani & Haq 1998 S04
    |    `--S. (Vesiculobates Hammer 1979) S04
    |         `--S. (*V.) silvatica (Hammer 1979) [=*Vesiculobates silvaticus] S04
    |--Exoribatula Jacot 1936 S04
    |    |--E. (Exoribatula) [incl. Nesoribatula Aoki 1964] S04
    |    |    |--*E. (E.) biundata Jacot 1936 S04
    |    |    |--E. (E.) ensifer (Mahunka 1983) [=Nesoribatula ensifer] S04
    |    |    |--E. (E.) pacifica (Aoki 1964) [=Nesoribatula pacifica] S04
    |    |    `--E. (E.) rotunda (Mahunka 1988) [=Nesoribatula rotunda] S04
    |    `--E. (Multoribates Hammer 1961) S04
    |         |--E. (*M.) chavinensis (Hammer 1961) [=*Multoribates chavinensis] S04
    |         |--E. (M.) longior (Hammer 1958) [=Scheloribates longior] S04
    |         |--‘Multoribates’ minor M83
    |         |--E. (M.) multisetosus (Balogh 1970) [=Tuberemaeus multisetosus] S04
    |         `--E. (M.) parvialatus (Hammer 1958) [=Scheloribates parvialatus] S04
    `--Hemileius Berlese 1916 S04
         |--H. (Hemileius) S04
         |--H. (Tuberemaeus) S04
         |--H. (Turcibates Ayyildiz & Luxton 1989) [incl. Tenuileius Lee 1989] S04
         |    |--H. (*T.) parvus (Ayyildiz & Luxton 1989) (see below for synonymy) S04
         |    |--H. (T.) lanceolatus Bayartogtokh & Aoki 1997 S04
         |    |--H. (T.) minimus Lee 1989 S04 [=H. (*Tenuileius) minimus BB92a]
         |    |--H. (T.) paratenuis Lee 1989 [=H. (Tenuileius) paratenuis] S04
         |    `--‘Turcibates’ porosus Ayyildiz & Luxton 1989 BB92b
         `--H. (Urubambates Hammer 1961) S04
              |--H. (*U.) punctatus (Hammer 1961) [=*Urubambates punctatus] S04
              |--H. (U.) calcaratus (Mahunka 1984) [=Urubambates calcaratus] S04
              |--H. (U.) flagellatus (Mahunka 1985) [=Urubambates flagellatus] S04
              |--H. (U.) paraguayensis (Balogh & Mahunka 1981) [=Urubambates paraguayensis] S04
              |--H. (U.) perlongus (Vasiliu & Calugar 1976) [=Urubambates perlongus] S04
              `--H. (U.) romanicus (Vasiliu & Calugar 1981) [=Urubambates romanicus] S04

Hemileius (*Turcibates) parvus (Ayyildiz & Luxton 1989) [=*Turcibates parvus; incl. Dometorina rossica Bashkirova 1958 (n. d.)] S04

*Type species of generic name indicated

REFERENCES

[BB92a] Balogh, J., & P. Balogh. 1992a. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[BB92b] Balogh, J., & P. Balogh. 1992b. The Oribatid Mites Genera of the World vol. 2. Plates. Hungarian Natural History Museum: Budapest.

[M83] Martin, N. A. 1983. Miscellaneous observations on a pasture fauna: an annotated species list. DSIR Entomology Division Report 3: 1-98.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.