Neoseiulus

Neoseiulus californicus feeding on two-spotted spider mite Tetranychus urticae, photographed by Lyle Buss.


Belongs within: Neoseiulini.

Neoseiulus is a large genus of about 300 species of predatory mite. Pesticide-resistant strains of Neoseiulus fallacis have been used in Australia for the control of spider mites in apple orchards (James 2001).

Characters (from Chant & McMurtry 2003): Dorsal shield generally significantly longer than wide, but with some species more rotund; ornamentation variable, usually reticulate, with or without waist at level of seta R1; posterior margin rounded; dorsal setal pattern 10A:9B, with 19 pairs of setae, including r3 and R1; dorsal setae short to medium in length, approximately subequal except that Z5, and sometimes Z4, are longer in some species; setiform, occasionally strongly serrated; seta R1 inserted on dorsal shield of females of a few species. Sternal shield with L/W ratio usually approximating 1:1, but in some species sternal shield is much longer than wide; seta St3 usually inserted on sternal shield, sometimes inserted on separate platelet; posterior margin straight or concave. Genital shield narrow or of medium width, truncate posteriorly, with genital setae, sometimes reticulated. Female ventrianal shield without conspicuous waist on most species, never vase-shaped; preanal setae JV1, JV2 and ZV2 arranged in broad triangular pattern, with seta JV2 well-removed from anterior margin of shield; metapodal plates normal. Peritreme usually extending to level of seta j1, shorter in some species; peritremal shield narrow, fused anteriorly with dorsal shield. Leg I always and legs II-III usually without macrosetae, at most with slightly adapted seta on genu III, more rarely on genu II. Leg IV with 0-3 macrosetae, most commonly one.

<==Neoseiulus Hughes 1948 (see below for synonymy) CM03
    |  i. s.: N. aequisetus (Wainstein 1962) CM03
    |         N. caudiglans P91
    |         N. cucumeroides (De Leon 1959) CM03
    |         N. driggeri (Specht 1968) CM03
    |         N. gracilentus (Hirschmann 1962) CM03
    |         N. indicus (Narayanan & Kaur 1960) CM03
    |         N. longisiphonulus Wu & Lan 1989 CM03
    |         N. mazurensis (Kropczynska 1965) CM03
    |         N. monomacroseta (Tseng 1976) CM03
    |         N. mumae (Shehata & Zaher 1969) CM03
    |         N. ornatus (Athias-Henriot 1957) CM03
    |         N. plantagenis (Kolodochka 1981) CM03
    |         N. scoticus (Collyer 1957) CM03
    |         N. shiheziensis Wu & Li 1987 CM03
    |         N. tiensaiensis (Tseng 1983) CM03
    |--+--N. rancidus (Chaudhri, Akbar & Rasool 1979) [=*Amathia rancidus] CM03
    |  `--N. xizangensis (Zhu & Chen 1985) CM03
    `--+--+--+--+--N. cavagnaroi (Schuster 1966) CM03
       |  |  |  |--N. gracilis (Muma 1962) CM03
       |  |  |  |--N. inflatus Kuznetsov 1984 CM03
       |  |  |  |--N. inornatus (Schuster & Pritchard 1963) CM03
       |  |  |  |--N. kennetti (Schuster & Pritchard 1963) [=Amblyseius kennetti] CM03
       |  |  |  `--N. kodryensis (Kolodochka 1980) CM03
       |  |  `--+--N. alustoni (Livschitz & Kuznetsov 1972) CM03
       |  |     |--N. angeliquae (Schicha 1987) CM03
       |  |     |--N. australograminis (Wainstein 1977) CM03
       |  |     |--N. cangaro (Schicha 1987) CM03
       |  |     |--N. dungeri (Karg 1977) CM03
       |  |     |--N. segnis (Wainstein & Arutunjan 1970) CM03
       |  |     |--N. tenuisetae Karg 1993 CM03
       |  |     `--N. vallis (Schuster & Pritchard 1963) CM03
       |  `--+--+--N. alpinus (Schweizer 1922) (see below for synonymy) CM03
       |     |  |--N. arutunjani (Wainstein & Begljarov 1971) CM03
       |     |  |--N. balisungsongus (Schicha & Corpuz-Raros 1992) CM03
       |     |  |--N. bibens (Blommers 1973) CM03
       |     |  |--N. brevicalyx Karg 1993 CM03
       |     |  |--N. cinctutus (Livschitz & Kuznetsov 1972) CM03
       |     |  |--N. clinopodii (Ke & Xin 1982) CM03
       |     |  |--N. dicercellatus (Wu & Ou 1999) CM03
       |     |  |--N. eucolli Karg 1993 CM03
       |     |  |--N. exiguus (Merwe 1968) CM03
       |     |  |--N. foramenis (Karg 1976) CM03
       |     |  |--N. kerri Muma 1965 CM03
       |     |  |--N. liticellus (Athias-Henriot 1966) CM03
       |     |  |--N. longispinosus (Evans 1952) CM03 [=Amblyseius longispinosis M91]
       |     |  |--N. lushanensis Zhu & Chen 1985 CM03
       |     |  |--N. makuwa (Ehara 1972) CM03
       |     |  |--N. marginatus (Wainstein 1961) CM03
       |     |  |--N. mckenziei (Schuster & Pritchard 1963) CM03
       |     |  |--N. montanus Tuttle & Muma 1973 CM03
       |     |  |--N. neoaurescens Moraes & Mesa 1988 CM03
       |     |  |--N. perfectus (Chaudhri 1968) CM03
       |     |  |--N. polyporus (Wainstein 1962) CM03
       |     |  |--N. pseudolongispinosus (Xin & Liang 1981) CM03
       |     |  |--N. scapilatus (Merwe 1965) CM03
       |     |  |--N. subtilisetosus (Begljarov 1962) CM03
       |     |  |--N. sugonjaevi (Wainstein & Abbasova 1974) CM03
       |     |  |--N. swartii Zack 1969 CM03
       |     |  |--N. teke (Pritchard & Baker 1962) CM03
       |     |  |--N. tervus Meshkov 1994 CM03
       |     |  |--N. womersleyi (Schicha 1975) [=Amblyseius womersleyi] CM03
       |     |  `--N. zeitunicus (Wainstein & Arutunjan 1970) CM03
       |     `--+--*N. barkeri Hughes 1948 CM03 [=Amblyseius barkeri CM03; incl. A. mckenziei RH91]
       |        |--N. aleurites Ragusa & Athias-Henriot 1983 CM03
       |        |--N. benicus (El-Badry 1968) CM03
       |        |--N. bindrai (Gupta 1976) CM03
       |        |--N. brevispinus (Kennett 1958) CM03
       |        |--N. calorai (Corpuz & Rimando 1966) CM03
       |        |--N. carunculus Chaudhri et al. 1979 CM03
       |        |--N. cydnodactylon (Shehata & Zaher 1969) CM03
       |        |--N. erugatus Ragusa & Athias-Henriot 1983 CM03
       |        |--N. ghanii (Muma 1967) CM03
       |        |--N. hamus Karg 1993 CM03
       |        |--N. harrowi (Collyer 1965) CM03
       |        |--N. huffakeri (Schuster & Pritchard 1963) CM03
       |        |--N. kermanicus (Daneshvar 1987) CM03
       |        |--N. liangi Chant & McMurtry 2003 [=N. ornatus Liang & Ke 1984 non Athias-Henriot 1957] CM03
       |        |--N. loxus (Schuster & Pritchard 1963) CM03
       |        |--N. marinellus (Muma 1962) CM03
       |        |--N. masiaka (Blommers & Chazeau 1974) CM03
       |        |--N. mycophilus (Karg 1970) CM03
       |        |--N. oahuensis (Prasad 1968) CM03
       |        |--N. pegasus (Schuster 1966) CM03
       |        |--N. picketti (Specht 1968) CM03
       |        |--N. pieteri (Schultz 1972) CM03
       |        |--N. planatus (Muma 1962) CM03
       |        |--N. pristisimilis Karg 1993 CM03
       |        |--N. pseudoherbarius Meshkov 1994 CM03
       |        |--N. rangatensis (Gupta 1977) CM03
       |        |--N. rufus Denmark & Evans in Denmark et al. 1999 CM03
       |        |--N. setulus (Fox 1947) CM03
       |        |--N. sioux (Chant & Hansell 1971) CM03
       |        |--N. sospesitis (Khan & Chaudhri 1969) CM03
       |        |--N. stolidus (Chaudhri 1968) CM03
       |        |--N. tareensis (Schicha 1983) CM03
       |        |--N. transversus Denmark & Muma 1973 CM03
       |        |--N. uliginosus (Karg 1976) CM03
       |        |--N. usitatus (Merwe 1965) CM03
       |        |--N. vagus (Denmark 1965) CM03
       |        |--N. vanderlindei (Merwe 1965) CM03
       |        |--N. vehementis (Khan & Chaudhri 1969) CM03
       |        `--N. venustus (Chaudhri 1968) CM03
       `--+--+--N. leucophaeus (Athias-Henriot 1959) [=Amblyseius leucophaeus, *Phytodromus leucophaeus] CM03
          |  `--+--N. akakia Beard 2001 CM03
          |     |--N. allenrolfius Denmark 1993 CM03
          |     |--N. brigarina Beard 2001 CM03
          |     |--N. cappari Beard 2001 CM03
          |     |--N. carverae Beard 2001 CM03
          |     |--N. dodoneae (Schicha 1980) CM03
          |     |--N. edestes Beard 2001 CM)3
          |     |--N. harveyi McMurtry & Schicha 1987 CM03
          |     |--N. lyra Beard 2001 CM03
          |     |--N. paloratus Beard 2001 CM03
          |     |--N. pannuceus Beard 2001 CM03
          |     |--N. queenslandensis McMurtry & Schicha 1987 CM03
          |     |--N. sparaxis Beard 2001 CM03
          |     |--N. specus Beard 2001 CM03
          |     |--N. steinerae Beard 2001 CM03
          |     |--N. tabis (Schuster & Pritchard 1963) CM03
          |     |--N. wanrooyae Beard 2001 CM03
          |     `--N. warrum Beard 2001 CM03
          `--+--+--+--N. canadensis (Chant & Hansell 1971) [=Amblyseius canadensis] CM03
             |  |  |--N. lateralis (Tuttle & Muma 1973) CM03
             |  |  `--N. subsolidus (Begljarov 1960) CM03
             |  `--+--N. letrauformis (Schicha & Corpuz-Raros 1993) [=Amblyseius letrauformis] CM03
             |     `--+--+--N. aceriae (Gupta 1975) CM03
             |        |  |--N. baraki (Athias-Henriot 1966) CM03
             |        |  |--N. benjamini (Schicha 1981) CM03
             |        |  |--N. bheraensis Chaudhri et al. 1979 CM03
             |        |  |--N. coatesi (Schultz 1972) CM03
             |        |  |--N. dhooriae (Gupta 1976) CM03
             |        |  |--N. echinochlovorus Schicha & Corpuz-Raros 1993 CM03
             |        |  |--N. inabanus (Ehara 1972) CM03
             |        |  |--N. interfolius (De Leon 1962) CM03
             |        |  |--N. lula (Pritchard & Baker 1962) CM03
             |        |  |--N. mumai (Denmark 1965) CM03
             |        |  |--N. paspalivorus (De Leon 1957) [=Typhlodromus paspalivorus] CM03
             |        |  |--N. rambami Swirski & Amitai 1990 CM03
             |        |  |--N. shambati (El-Badry 1968) CM03
             |        |  |--N. sporobolus Tuttle & Muma 1973 CM03
             |        |  |--N. tabularis Chaudhri 1979 CM03
             |        |  `--N. taiwanicus (Ehara 1970) CM03
             |        `--+--N. peruanus (El-Banhawy 1979) [=Amblyseius peruanus] CM03
             |           `--+--N. astutus (Begljarov 1960) CM03
             |              |--N. chikmagaluriensis (Gupta 1985) [=Amblyseius (Typhlodromalus) chikmagaluriensis] CM03
             |              |--N. chitradurgae (Gupta 1985) [=Amblyseius (Typhlodromalus) chitradurgae] CM03
             |              |--N. danilevskyi (Wainstein & Arutunjan 1970) CM03
             |              |--N. desertus (Chant 1957) [=Typhlodromus (Amblyseius) desertus] CM03
             |              |--N. fauveli (Athias-Henriot 1978) CM03
             |              |--N. gansuensis Wu & Lan 1991 CM03
             |              |--N. hanselli (Chant & Yoshida-Shaul 1978) CM03
             |              |--N. huron (Chant & Hansell 1971) (see below for synonymy) CM03
             |              |--N. insularis (Athias-Henriot 1978) CM03
             |              |--N. kolodotshkai Kuznetsov 1984 CM03
             |              |--N. luppovae (Wainstein 1962) CM03
             |              |--N. multiporus Wu & Li 1987 CM03
             |              |--N. parvipilis (Athias-Henriot 1978) CM03
             |              |--N. populi Bozai 1997 CM03
             |              |--N. riparius Kolodochka 1991 CM03
             |              |--N. sinaiticum (Amitai & Swirski 1982) CM03
             |              |--N. turangae (Kolodochka 1982) CM03
             |              `--N. versutus (Begljarov 1981) CM03
             `--+--+--+--N. accessus (Ueckermann & Loots 1988) CM03
                |  |  |--N. agrestis (Karg 1960) CM03
                |  |  |--N. arenillus (De Leon 1967) CM03
                |  |  |--N. ceratoni (Ueckermann & Loots 1988) [=Amblyseius (Amblyseius) ceratoni] CM03
                |  |  |--N. cynodonae (Gupta 1977) CM03
                |  |  |--N. disparis (Chaudhri, Akbar & Rasool 1979) [=*Denmarkia disparis] CM03
                |  |  |--N. salish (Chant & Hansell 1971) CM03
                |  |  |--N. shanksi Congdon 2002 CM03
                |  |  |--N. striatus (Wu 1983) CM03
                |  |  |--N. tauricus (Livschitz & Kuznetsov 1972) CM03
                |  |  |--N. umsteadi (Muma, Metz & Farrier 1967) CM03
                |  |  `--N. vasoides Karg 1989 CM03
                |  `--+--N. bicaudus (Wainstein 1962) CM03
                |     |--N. camarus (El-Badry 1968) CM03
                |     |--N. certus Kolodochka 1990 CM03
                |     |--N. conterminus Kolodochka 1990 CM03
                |     |--N. ellesmerei (Chant & Hansell 1971) CM03
                |     |--N. extricatus Kolodochka 1991 CM03
                |     |--N. hirotae (Ehara 1985) CM03
                |     |--N. houstoni Schicha 1985 CM03
                |     |--N. iroquois (Chant & Hansell 1971) CM03
                |     |--N. marinus (Willmann 1952) [=Lasioseius marinus, *Cydnodromus marinus] CM03
                |     |--N. micmac (Chant & Hansell 1971) CM03
                |     |--N. nodus Denmark & Knisley in Knisley & Denmark 1978 CM03
                |     |--N. paraki (Ehara 1967) [=Amblyseius paraki] CM03
                |     |--N. paramarinus Evans 1987 CM03
                |     |--N. provectus Kolodochka 1991 CM03
                |     |--N. simplexus Denmark & Knisley in Knisley & Denmark 1978 CM03
                |     |--N. subreticulatus (Wu 1987) CM03
                |     |--N. tshernovi (Kuznetsov 1984) CM03
                |     |--N. ulatei Denmark & Evans in Denmark et al. 1999 CM03
                |     |--N. vardgesi (Arutunjan 1968) CM03
                |     `--N. zwoelferi (Dosse 1957) CM03
                `--+--+--N. neotunus (De Leon 1973) CM03
                   |  |--N. plumosus (Denmark & Muma 1975) CM03
                   |  `--N. tunus (De Leon 1967) [=Typhlodromips tunus] CM03
                   `--+--N. aegyptocitri (Kandeel & El-Hallawany 1986) CM03
                      |--N. alidis Kolodochka 1989 CM03
                      |--N. amicus (Chant 1959) CM03
                      |--N. anonymus (Chant & Baker 1965) CM03
                      |--N. apeuthes Beard 2001 CM03
                      |--N. apkutik (Chant & Hansell 1971) CM03
                      |--N. argillaceous (Kolodochka & Bondarenko 1993) CM03
                      |--N. aridus (De Leon 1962) CM03
                      |--N. atrii Karg 1989 CM03
                      |--N. atsak (Chant & Hansell 1971) CM03
                      |--N. baticolus (Athias-Henriot 1977) CM03
                      |--N. bayviewensis (Schicha 1977) CM03
                      |--N. bellinus (Womersley 1954) CM03
                      |--N. byssus Denmark & Knisley in Knisley & Denmark 1978 CM03
                      |--N. californicus (McGregor 1954) CM03 (see below for synonymy)
                      |--N. callunae (Willmann 1952) CM03
                      |--N. campana Beard 2001 CM03
                      |--N. caobae De Leon 1965 CM03
                      |--N. caribbeanus (De Leon 1965) CM03
                      |--N. casimiri (Schicha & Elshafie 1980) CM03
                      |--N. chascomensis (Sheals 1962) CM03
                      |--N. chaudhrii Chant & McMurtry 2003 (see below for synonymy) CM03
                      |--N. chilenensis (Dosse 1958) CM03
                      |--N. chinensis Chant & McMurtry 2003 [=N. crataegi Wang & Xu non (Jorgensen & Chant 1960)] CM03
                      |--N. collegae (De Leon 1962) [incl. N. ohioensis (De Leon 1962)] CM03
                      |--N. comitatus (De Leon 1962) CM03
                      |--N. conconiensis Karg 1965 CM03
                      |--N. coprophilus (Karg 1970) CM03
                      |--N. crataegi (Jorgensen & Chant 1960) CM03
                      |--N. cree (Chant & Hansell 1971) CM03
                      |--N. cryptomeriae Zhu & Chen 1983 CM03
                      |--N. cucumeris (Oudemans 1930) CM03 (see below for synonymy)
                      |--N. culpus Denmark & Evans in Denmark et al. 1999 CM03
                      |--N. curvus Wu & Li 1985 CM03
                      |--N. dieteri (Schicha 1979) CM03
                      |--N. dissipatus Kolodochka 1991 CM03
                      |--N. draconis (Chaudhri et al. 1979) CM03
                      |--N. engaddensis (Amitai & Swirski 1970) CM03
                      |--N. eremios Beard 2001 CM03
                      |--N. esculentus (El-Badry 1968) CM03
                      |--N. fallacis (Garman 1948) CM03 [=Amblyseius fallacis J01]
                      |--N. fallacoides Tuttle & Muma 1973 CM03
                      |--N. ficusi (Gupta 1986) CM03
                      |--N. haimatus (Ehara 1967) CM03
                      |--N. helmi (Schicha 1987) CM03
                      |--N. idaeus Denmark & Muma 1973 CM03
                      |--N. imbricatus (Corpuz & Rimando 1966) CM03
                      |--N. inak (Chant & Hansell 1971) CM03
                      |--N. innuit (Chant & Hansell 1971) CM03
                      |--N. insignitus (Merwe 1968) CM03
                      |--N. kapjik (Chant & Hansell 1971 CM03
                      |--N. kearnae Beard 2001 CM03
                      |--N. koyamanus (Ehara & Yokogawa 1977) CM03
                      |--N. krugeri (Merwe 1968) CM03
                      |--N. lablabi (Ghai & Menon 1967) CM03
                      |--N. lamticus (Athias-Henriot 1977) CM03
                      |--N. latoventris Karg & Edland 1987 CM03
                      |--N. lecki Beard 2001 CM03
                      |--N. leigongshanensis Wu & Lan 1989 CM03
                      |--N. longilaterus (Athias-Henriot 1957) CM03
                      |--N. loxtoni (Schicha 1979) CM03
                      |--N. malaban Beard 2001 CM03
                      |--N. martinicensis Moraes, Kreiter & Lofego 2000 CM03
                      |--N. melaleucae (McMurtry & Schicha 1987) CM03
                      |--N. muganicus (Abbasova 1970) CM03
                      |--N. myrtea Chaudhri 1979 CM03
                      |--N. neoghanii (Gupta 1968) CM03
                      |--N. neoparaki (Ehara 1972) CM03
                      |--N. neoreticuloides Liang & Ke 1988 CM03
                      |--N. nescapi (Chant & Hansell 1971) CM03
                      |--N. nestor Beard 2001 CM03
                      |--N. noosae McMurtry & Schicha 1987 CM03
                      |--N. novaescotiae (Chant 1959) CM03
                      |--N. paraibensis (Moraes & McMurtry 1983) CM03
                      |--N. pepperi (Specht 1968) CM03
                      |--N. perspectus Kolodochka 1992 CM03
                      |--N. phragmitidis Bozai 1997 CM03
                      |--N. placitus (Khan & Chaudhri 1969) CM03
                      |--N. poculi Karg 1976 CM03
                      |--N. pseudoumbraticus (Chant & Yoshida-Shaul 1982) CM03
                      |--N. recifensis Gondin & Moraes 2001 CM03
                      |--N. reductus (Wainstein 1962) [=Amblyseius reductus] CM03
                      |--N. reticulatus (Oudemans 1930) CM03
                      |--N. reticuloides (Wainstein 1975) CM03
                      |--N. scyphus Schluster & Pritchard 1963 CM03
                      |--N. sehlabatei (El-Banhawy 2002) CM03
                      |--N. sharonensis (Rivnay & Swirski 1980) CM03
                      |--N. spicatus Denmark & Evans in Denmark et al. 1999 CM03
                      |--N. spineus (Tseng 1976) CM03
                      |--N. subrotundus Wu & Lan 1991 CM03
                      |--N. thwaitei (Schicha 1977) CM03
                      |--N. tibielingmiut (Chant & Hansell 1971) CM03
                      |--N. tobon (Chant & Hansell 1971) CM03
                      |--N. tornadus Tuttle & Muma 1973 CM03
                      |--N. triangularis Karg 1994 CM03
                      |--N. umbraticus (Chant 1956) CM03
                      |--N. veigai Gondin & Moraes 2001 CM03
                      |--N. wearnei Schicha 1987 CM03
                      `--N. yanoi (Ehara 1972) CM03

Nomina nuda: Neoseiulus arenarius Denmark & Edland in Chant & McMurtry 2003 CM03
             Neoseiulus communis Denmark & Edland in Chant & McMurtry 2003 CM03

Neoseiulus Hughes 1948 [incl. Amathia Chaudhri, Akbar & Rasool 1979 non Lamouroux 1912, Cydnodromus Muma 1961, Denmarkia Chaudhri, Akbar & Rasool 1979, Dictyonotus Athias-Henrot 1978, Dictydionotus Athias-Henrot 1979, Kashmerius Chaudhri, Akbar & Rasool 1979, Phytodromus Muma 1961, Typhlodromopsis De Leon 1959] CM03

Neoseiulus alpinus (Schweizer 1922) [incl. N. aurescens (Athias-Henriot 1961), N. polonicus (Willmann 1949)] CM03

Neoseiulus californicus (McGregor 1954) CM03 [=Amblyseius californicus M91; incl. N. mungeri (McGregor 1954) CM03]

Neoseiulus chaudhrii Chant & McMurtry 2003 [=*Kashmerius reductus Chaudhri, Akbar & Rasool 1979 non Amblyseius reductus Wainstein 1962] CM03

Neoseiulus cucumeris (Oudemans 1930) CM03 [=Typhlodromus cucumeris CM03, Amblyseius (*Typhlodromopsis) cucumeris CM03; incl. A. thripsi McGill 1939 RC91, CM03, Neoseiulus thripsi CM03]

Neoseiulus huron (Chant & Hansell 1971) [=Amblyseius huron, *Dictydionotus huron, *Dictyonotus huron] CM03

*Type species of generic name indicated

REFERENCES

[CM03] Chant, D. A., & J. A. McMurtry. 2003. A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part I. Neoseiulini new tribe. International Journal of Acarology 29 (1): 3-46.

[J01] James, D. G. 2001. History and perspectives of biological mite control in Australian horticulture using exotic and native phytoseiids. In Acarology: Proceedings of the 10th International Congress (R. B. Halliday, D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff, eds) pp. 436-443. CSIRO Publishing: Melbourne.

[M91] McMurtry, J. A. 1991. Augmentative releases to control mites in agriculture. In Dusbábek, F. & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 1 pp. 151-158. SPB Academic Publishing: The Hague.

[P91] Petrushov, A. Z. 1991. Selection and mechanisms of ambush resistance in Metaseiulus occidentalis N. (Acarina: Phytoseiidae). In Dusbábek, F. & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 741-748. SPB Academic Publishing: The Hague.

[RC91] Ragusa Di Chiara, S. 1991. Using native phytoseiids in agricultural cropping systems. In Dusbábek, F. & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 1 pp. 159-166. SPB Academic Publishing: The Hague.

[RH91] Rijn, P. C. J. van, & Y. M. van Houten. 1991. Life history of Amblyseius cucumeris and A. barkeri (Acarina: Phytoseiidae) on a diet of pollen. In Dusbábek, F. & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 647-654. SPB Academic Publishing: The Hague.

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