Specimen of Substilbula pallidiclava (missing gaster), photographed by John M. Heraty.

Belongs within: Eucharitidae.
Contains: Tricoryna, Austeucharis, Chalcura, Schizaspidia.

The Eucharitinae are a group of wasps in which many species of which have the scutellum strongly produced apically.

Characters (from Bouček 1988): Male antennae often with branches; pronotum dorsally strongly shortened and mostly concealed by the head; prepectus evidently fused with pronotum, although fusion sometimes indicated as a groove or carina; axillae well developed; scutellum often apically produced.

<==Eucharitinae [Eucharissinae] B88
    |--Psilocharitini HB13
    |    |--‘Psilocharis’ afra MH11
    |    `--+--Neolosbanus HB13
    |       |    |--N. palgravei MH11
    |       |    `--N. purpureoventris MH11
    |       `--Psilocharis MH11
    |            |--P. pacifica MH11
    |            `--P. theocles MH11
    `--Eucharitini HB13
         |--+--Neostilbula ranomafanae MH11
         |  `--Pseudometagea HB13
         |       |--P. montana MH11
         |       `--+--P. bakeri MH11
         |          `--P. schwarzii MH11
         `--+--+--Galearia MH11
            |  |    |--G. bruchi MH11
            |  |    `--G. latreillei HB13
            |  `--+--Kapala Cameron 1884 HB13, GM79
            |     |    `--K. floridana MH11
            |     `--Colocharis napoana MH11
            `--+--+--Ancylotropus HB13
               |  |    |  i. s.: A. manipurensis [=Schizaspidia manipurensis, Stilbula manipurensis] B88
               |  |    |--A. cariniscutis MH11
               |  |    `--+--A. montanus MH11
               |  |       `--Eucharissa natalica MH11
               |  `--+--Schizaspidia MH11
               |     `--Saccharissa Kirby 1886 MH11, B88
               |          |--*S. contigens (Walker 1862) [=Eucharis contigens, S. contingens Kirby 1886] B88
               |          |--S. alcocki MH11
               |          |--S. latifurca Bouček 1988 B88
               |          `--S. vicina [=Schizaspidia vicina] B88
               `--+--Eucharis HB13
                  |    |--E. adscendens B88
                  |    `--E. albipennis MH11
                  `--+--+--Obeza MH11
                     |  `--Lophyrocera HB13
                     |       |--L. pretendens MH11
                     |       `--L. variabilis MH11
                     `--+--Pseudochalcura HB13
                        |    |--P. gibbosa HB13
                        |    `--P. prolata MH11
                        `--Substilbula Bouček 1988 MH11, B88
                             |--*S. bidentata (Girault 1913) [=Stilbula bidentata] B88
                             |--S. albipennis (Girault 1929) [=Stilbula albipennis] B88
                             |--S. australiana (Girault 1913) [=Stilbula australiana] B88
                             `--S. pallidiclava (Girault 1934) [=Stilbula pallidiclava] B88

Eucharitinae incertae sedis:
  Psilogastrellus Ghesquière 1946 [=Psilogaster Blanchard 1840 non R.L. 1817; incl. Pachyeucharis] B88
  Striostilbula Bouček 1988 B88
    `--*S. quadridigitata (Girault 1929) [=Stilbula quadridigitata] B88
  Thoracanthoides Girault 1928 B88
    `--*T. albispina Girault 1928 B88
  Parapsilogastrus Ghesquière 1946 (see below for synonymy) B88
    |--*P. fausta (Walker 1839) (see below for synonymy) B88
    `--P. ulyanovi (Girault 1940) [=Epimetagea ulyanovi] B88
  Propsilogaster Girault 1940 B88
    |--*P. biclavata Girault 1940 B88
    `--P. alcicornis Bouček 1988 B88
  Rhipipalloidea Girault 1934 B88
    |--*R. mira Girault 1934 [=Rhipipallus mirus] B88
    `--R. gruberi Girault 1940 [=Rhipipallus gruberi] B88
  Stilbuloida Bouček 1988 B88
    |--*S. doddi (Bingham 1906) [=Schizaspidia doddi] B88
    `--S. calomyrmecis (Brues 1934) [=Schizaspidia calomyrmecis] B88
  Losbanus Ishii 1932 B88
    |--*L. uichancoi B88
    |--L. antennatus (Gahan 1940) [=Psilogaster antennata] B88
    `--L. minutus Bouček 1988 B88
  Stilbula Spinola 1811 [incl. Eltolada Cameron 1909] B88
    |--*S. cyniformis [=Ichneumon cyniformis] B88
    |--S. albipetiole Girault 1929 B88
    |--S. arenae Girault 1934 B88
    |--S. brunneipetiole (Girault 1934) [=Epimetagea brunneipetiole] B88
    |--S. octodigitata Girault 1929 B88
    |--S. peduncularis Westwood 1874 B88
    |--S. polyrhachicida [=Schizaspidia polyrhachicida] B88
    |--S. quinqueguttata (Girault 1915) [=Schizaspidia quinqueguttata] B88
    |--S. toga Girault 1937 B88
    `--*Eltolada’ trimaculata B88
  Tricoryna B88
  Austeucharis B88
  Chalcura B88

Parapsilogastrus Ghesquière 1946 [=Parapsilogaster Girault 1915 non Bigot 1882, Psilogasteroides Girault 1913 non Brethes 1910] B88

*Parapsilogastrus fausta (Walker 1839) [=Eucharis fausta, Epimetagea faustus (l. c.), *Parapsilogaster fausta, *Psilogasteroides fausta; incl. Epimetagea phidiasae Girault 1940] B88

*Type species of generic name indicated


[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[GM79] Gordh, G., A. S. Menke, E. C. Dahms & J. C. Hall. 1979. The privately printed papers of A. A. Girault. Memoirs of the American Entomological Institute 28: 1-400.

[HB13] Heraty, J. M., R. A. Burks, A. Cruaud, G. A. P. Gibson, J. Liljeblad, J. Munro, J.-Y. Rasplus, G. Delvare, P. Janšta, A. Gumovsky, J. Huber, J. B. Woolley, L. Krogmann, S. Heydon, A. Polaszek, S. Schmidt, D. C. Darling, M. W. Gates, J. Mottern, E. Murray, A. D. Molin, S. Triapitsyn, H. Baur, J. D. Pinto, S. van Noort, J. George & M. Yoder. 2013. A phylogenetic analysis of the megadiverse Chalcidoidea (Hymenoptera). Cladistics 29: 466-542.

[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PLoS One 6 (11): e27023.

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