Buresium naso, photographed by Mike Gates.

Belongs within: Chalcidoidea.
Contains: Eurytominae.

The Eurytomidae are a family of chalcidoid wasps including both parasitic and phytophagous species. The pronotum in eurytomids is large and subrectangular, and the majority of species are non-metallic in coloration and bear strong piliferous puncturation (Bouček 1988).

Characters (from Bouček 1988): Body usually non-metallic, black or marked with yellow or reddish; thorax mostly with coarse piliferous puncturation; pronotum subrectangular, large; notauli complete though posteriorly sometimes shallow; lower mesepimeron not separated; forewing without linea calva; tarsi five-segmented; gaster convex, non-collapsing, ovipositor hardly exserted (if apex narrowed in a cauda then covered by extended epipygium).

<==Eurytomidae HB13
    |--Buresiinae HB13
    |    |--Buresium HB13
    |    |    |--B. naso HB13
    |    |    `--B. rufum HB13
    |    `--Macrorileya HB13
    `--+--Rileyinae [Rileyini] B88
       |    |  i. s.: Archirileya B88
       |    |         Heimbrella Subba Rao 1978 B88
       |    |         Calorileya Crawford 1910 B88
       |    |         Dougiola Bouček 1988 B88
       |    |           `--*D. koebelei (Ashmead 1900) [=Systole koebelei] B88
       |    |         Matna Bouček 1988 B88
       |    |           `--*M. fallax Bouček 1988 B88
       |    |--+--Heimbra [Heimbrinae, Heimbrini] HB13
       |    |  |    `--H. opaca HB13
       |    |  `--Neorileya flavipes HB13
       |    `--Rileya Ashmead 1888 (see below for synonymy) B88
       |         |  i. s.: *R. cecidomyiae B88
       |         |         R. asiatica B88
       |         |         R. spadix (Girault 1915) [=*Pararileya spadix] B88
       |         |         R. vardyi B88
       |         |--+--R. atopogaster MH11
       |         |  `--R. grisselli MH11
       |         `--+--R. longitergum HB13
       |            `--+--R. pallidipes HB13
       |               `--Austrophotismus Girault 1938 HB13, B88
       |                    `--A. daicles (Walker 1839) [=Miscogaster daicles; incl. *A. ater Girault 1938] B88
       `--Eurytominae HB13

Rileya Ashmead 1888 [incl. Ashmeadia Howard 1889, Pararileya Girault 1915, Tragiicola Brethes 1918, Xenopelte Cameron 1913] B88

*Type species of generic name indicated


[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[HB13] Heraty, J. M., R. A. Burks, A. Cruaud, G. A. P. Gibson, J. Liljeblad, J. Munro, J.-Y. Rasplus, G. Delvare, P. Janšta, A. Gumovsky, J. Huber, J. B. Woolley, L. Krogmann, S. Heydon, A. Polaszek, S. Schmidt, D. C. Darling, M. W. Gates, J. Mottern, E. Murray, A. D. Molin, S. Triapitsyn, H. Baur, J. D. Pinto, S. van Noort, J. George & M. Yoder. 2013. A phylogenetic analysis of the megadiverse Chalcidoidea (Hymenoptera). Cladistics 29: 466-542.

[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PLoS One 6 (11): e27023.

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