Belongs within: Pentatomoidea.
Contains: Myrocharia, Edessa, Diemeniini, Eysarcoris, Menida, Poecilometis, Tarisini, Phyllocephalinae, Cephaloplatus, Dictyotus, Euschistus, Strachiaria, Antestiaria, Pentatoma, Nezara, Cuspicona.
The Pentatominae is a highly diverse group of stink bugs, the exact definition of which remains debated. Members of the group combine a head extending well in front of the eyes with spiracles on the second abdominal segment concealed by the metapleura. The scutellum is usually triangular and smaller than the pronotum but in some members, such as the Ippatha group and the Tarisini and related taxa, it is greatly enlarged to cover most of the abdomen and/or semielliptical in shape (Gross 1975). Most pentatomines are plant feeders but one group of genera (including Oechalia, Cermatulus and relatives) are predatory.
Characters (from Gross 1975): Head always extended well in front of eyes; scutellum usually triangular, if enlarged then corium anteriorly concealing some anterior laterotergites; posterolateral margin of pronotum straight or shallowly excavated, meso- and metanota not visible.
<==Pentatominae [Amyotinae, Graphosomatinae]
|--Mycoolona atricornis G75, G76
|--+--Mecidea [Mecideinae, Mecideini] G75
| `--Diemeniini G75
|--Eysarcoris group G75
| |--Eysarcoris G76
| |--Neostollia G76
| `--Spermatodes G76
|--Poecilotoma Dallas 1851 [incl. Allportia Schoutenden 1906] G75
| |--*P. grandicornis (Erichson 1842) [=Atelocerus grandicornis] G75
| |--P. callosa Gross 1975 G75
| |--P. perconfusa Gross 1975 G75
| `--P. scutellata (Van Duzee 1905) [=Platycoris scutellatus] G75
|--Kapunda group G75
| |--Adelaidena regina G76
| `--Kapunda Distant 1911 G76
| |--K. troughtoni (Distant 1910) (see below for synonymy) G76
| `--K. tepperi Gross 1976 G76
|--Ippatha group G75
| |--Bachesua Gross 1975 G75
| | `--*B. enigmatica Gross 1975 G75
| `--Ippatha Distant 1910 G75
| |--*I. australiensis Distant 1910 G75
| `--I. angustilinea Musgrave 1930 G75
|--Podopini [Podopinae] G75
| |--Coracanthella geophila G75, CGW91
| |--Weda G75
| |--Amaurochrous G75
| `--Scotinophara G75
| |--S. allanae G75
| `--S. geophila (Montr. 1858) [=Scutellera geophila] D14
|--Menida group G75
| |--Menida G76
| |--Sciomenida Gross 1976 G76
| | `--*S. mcfarlandi Gross 1976 G76
| `--Lathraedoeus G76
| |--*L. speculator G76
| |--L. inconstans [=Menida inconstans] G76
| `--L. leucophaeus (Walk. 1867) G76, D00a [=Antestia leucophaea G76, Menida leucophaea G76]
|--Piezodorus group G75
| |--Anaximenes G76
| `--Piezodorus Fieber 1861 G76
| |--P. lituratus (Fabricius 1794) G76 (see below for synonymy)
| |--P. guildinii H91
| |--P. hybneri (Gmelin 1790) G76 (see below for synonymy)
| |--P. incarnatus [=Cimex incarnatus; incl. Rhaphigaster parilis Walker 1867] D00b
| |--P. inexpertus (Walker 1867) [=Rhaphigaster inexpertus] D00b
| |--P. pallescens (Walker 1868) K08, D00b [=Rhaphigaster pallescens D00b]
| `--P. punctiventris (Dall. 1851) [=Rhaphigaster punctiventris, Antestia punctiventris] D00b
|--Halyini [Halyinae] G76
| |--Poecilometis G76
| |--Atelias variegatus G76
| |--Coctoteris Stål 1858 G76
| |--Tinganina Bergroth 1909 G76
| |--Neosurenus Distant 1920 G76
| |--Elemana Distant 1912 G76
| |--Baracellus Distant 1903 G76
| |--Austromalaya Kirkaldy 1908 G76
| |--Bathrus Dallas 1851 G76
| |--Dalpada Amyot & Serville 1843 G76
| |--Acanthidiellum Kirkaldy 1904 [incl. Bromocoris Horvarth 1915] G76
| |--Ectenus Dallas 1851 G76
| |--Accarana Distant 1888 G76
| |--Auxentius Horvarth 1915 G76
| |--Neoptolemus Horvarth 1915 G76
| |--Babylas Horvarth 1915 G76
| |--Brizica Walker 1867 G76
| |--Dippilana Distant 1910 G76
| | `--*D. membranacea Distant 1910 [incl. Poecilometis gibbiceps Bergroth 1916] G76
| |--Dictyometis Gross 1976 G76
| | `--*D. spriggi Gross 1976 G76
| |--Parapoecilometis Gross 1976 G76
| | `--*P. campbelli Gross 1976 G76
| |--Polycarmes Stål 1867 G76
| | `--P. punctatissimus (Montr. 1858) [=Acanthidium punctatissimum] D14
| |--Anchises Stål 1867 G76
| | `--A. parvulus (Westwood 1837) [=Halys parvulus, Theseus parvulus] G76
| |--Theseus Stål 1867 G76
| | |--*T. modestus (Stål 1865) (see below for synonymy) G76
| | `--T. distanti Gross 1972 [=T. scutellatus Distant 1910 non Poecilometis scutellatus Distant 1899] G76
| |--Aglaophon Stål 1876 G76
| | |--*A. varius Stål 1876 G76
| | `--A. variegatus Gross 1976 G76
| |--Paragenor Gross 1976 G76
| | |--*P. halei Gross 1976 G76
| | |--P. atomicus Gross 1976 G76
| | `--P. robustus Gross 1976 G76
| |--Alcaeus Dallas 1851 [incl. Muritha] G76
| | |--*A. varicornis (Westwood 1837) [=Atelocerus varicornis; incl. Al. subspinosus] G76
| | |--A. hermannsburgi (Distant 1910) [=*Muritha hermannsburgi] G76
| | |--A. lignicolor Walker 1867 G76
| | `--A. unifornis Gross 1976 G76
| |--Neagenor Bergroth 1891 [=Agenor Dallas 1851 (preoc.)] G76
| | |--*N. spinosus (Dallas 1851) [=*Agenor spinosus] G76
| | |--N. anteaureus Gross 1976 G76
| | |--N. minor Gross 1976 G76
| | `--N. transcontinentalis Gross 1976 G76
| `--Omyta Spinola 1850 G76
| |--O. centrolineata (Westwood 1837) G76 (see below for synonymy)
| | |--O. c. centrolineata G76
| | `--O. c. siccior Gross 1976 G76
| |--O. delineata G76
| `--O. spinosa (Dallas 1851) G75 [=Poecilotoma spinosa G75; incl. Platycoris rufescens G76]
|--+--Menestheus group G75
| | |--Menestheus cuneatus G75
| | |--Neomenestheus G75
| | |--Hillieria Distant 1910 G75
| | | `--*H. acuminata Distant 1910 G75
| | |--Paramenestheus Breddin 1900 G75
| | | |--*P. terricolor Breddin 1900 (see below for synonymy) G75
| | | `--P. nercivus (Dallas 1851) [=Sciocoris nercivus] G75
| | |--Pseudaelia Distant 1910 G75
| | | |--*P. clementi Distant 1910 G75
| | | |--P. gnielae Gross 1975 G75
| | | `--P. rectimargo Gross 1975 G75
| | `--Eribotes Stål 1867 G75
| | |--*E. australis (Dallas 1852) [=Sciocoris australis] G75
| | |--E. bicuspis G75
| | |--E. challengeri G75
| | |--E. delitescens G75
| | |--E. hobartensis Distant 1910 G75
| | |--E. indicator (Walker 1868) [=Sciocoris indicator] G75
| | `--E. reconditus Bergroth 1920 G75
| `--+--Tarisini G75
| |--Kitsonia Gross 1975 G75
| | `--*K. rubrocoriosa Gross 1975 G75
| |--Macrocarenus group G75
| | |--Macrocarenus Stål 1873 G75
| | | `--*M. acuminatus (Dallas 1851) [=Eurygaster acuminatus] G75
| | `--Macrocarenoides Gross 1975 G75
| | `--*M. scutellatus (Distant 1899) [=Macrocarenus scutellatus] G75
| |--Ochisme group G75
| | |--Ochisme Kirkaldy 1904 G75
| | | `--*O. australis (Dallas 1851) [=*Trachyops australis; incl. Hypolcus apricus Bergroth 1916] G75
| | |--Dysnoetus Bergroth 1918 G75
| | | `--D. rugulosus (Horvarth 1902) [=Dictyotus rugulosus; incl. *Dy. anceps Bergroth 1918] G75
| | |--Turrubulana Distant 1910 G75
| | | `--*T. plana Distant 1910 G75
| | `--Amphidexius Bergroth 1918 G75
| | `--*A. suspensus Bergroth 1918 G75
| `--Kumbutha group G75
| |--Kumbutha quadrinotata G75
| |--Cooperocoris Gross 1975 G75
| | `--*C. australis Gross 1975 G75
| |--Ooldeon Gross 1975 G75
| | `--*O. maculipes Gross 1975 G75
| |--Brizocoris Gross 1975 G75
| | `--*B. dimorphicus Gross 1975 G75
| |--Thryptomenecoris Gross 1975 G75
| | `--*T. officeri Gross 1975 G75
| |--Eremophilacoris Gross 1975 G75
| | `--*E. aridicolus Gross 1975 G75
| |--Pirricoris Gross 1975 G75
| | `--*P. annectans Gross 1975 G75
| |--Tepperocoris Gross 1975 G75
| | |--*T. lofteyensis Gross 1975 G75
| | `--T. vandijki Gross 1975 G75
| `--Buthumka Gross 1975 G75
| |--*B. orthoscutellata Gross 1975 G75
| |--B. reducta Gross 1975 G75
| `--B. transitionalis Gross 1975 G75
`--Pentatomini [Strachiini] R74
| i. s.: Sibaria R74
| Agroecus R74
| Tibraca limbativentris Stål 1858 R74, D00a [incl. Mormidea walkeri Leth. & Sev. 1893 D00a]
| Menecles incertus R74, E66
| Berecynthus R74
| Galedanta R74
| Dichelops R74
| |--D. divisus (Walker 1867) [=Diceraeus divisus] D00b
| |--D. mutabilis (Walker 1867) [=Diceraeus mutabilis] D00b
| `--D. pulchricornis (Walker 1867) [=Diceraeus pulchricornis] D00b
| Padaeus Stål 1862 R74
| |--*P. viduus (Vollenhoven 1868) [=Mormidea viduus] R74
| |--P. teapensis (Distant 1890) [=Euschistus teapensis] R74
| |--P. trivittatus R74
| `--P. verrucifer Stål 1862 [=Euschistus verrucifer] R74
|--+--Phyllocephalinae G75
| `--Cephaloplatus group G75
| |--Cephaloplatus WEE70
| `--Caystrus G76 [incl. Neodius G75]
|--+--Tholosanus group G75
| | |--Arniscus Distant 1899 G75
| | | `--A. humeralis G75
| | `--Tholosanus Distant 1899 G75
| | |--*T. proximus (Dallas 1851) (see below for synonymy) G75
| | `--T. colligatus (Walker 1867) [=Pentatoma colligata] G75
| `--Dictyotus group G75
| |--Dictyotus WEE70
| |--Utheria Gross 1975 G75
| | `--*U. dixoni Gross 1975 G75
| `--Paradictyotus Gross 1975 G75
| `--*P. fuscus Gross 1975 G75
|--Carpocoris group G75
| |--Codophila G76
| |--Proxys G76
| |--Euschistus R74
| |--Hyparete vitiensis Distant 1901 G76, K08
| |--Macromolus G76
| |--Lubentius G76
| |--Hypsithocus Bergroth 1927 G76, B27
| | `--*H. hudsonae Bergroth 1927 B27
| |--Dolycoris Mulsant & Rey 1866 G76, L-SSY04
| | `--D. baccarum [incl. Pentatoma inconcisa Walker 1867] D00b
| |--Monteithiella Gross 1976 G76
| | `--*M. humeralis (Walker 1868) G76 (see below for synonymy)
| |--Pseudapines Bergroth 1912 G76
| | `--*P. geminata (Van Duzee 1905) [=Apines geminata] G76
| |--Notius Dallas 1851 G76
| | |--*N. depressus Dallas 1851 G76
| | `--N. consputus G76
| |--Agonoscelis Spinola 1837 G76
| | |--*A. nubila (Fabricius 1775) [=Cimex nubilus] G76
| | `--A. rutila (Fabricius 1775) G76 (see below for synonymy)
| |--Anaxarchus G76
| | |--A. pardalinus D14
| | `--A. reyi (Montr. 1858) [=Pentatoma reyi] D14
| |--Eurinome G76
| | |--E. inconspicua (Montrouz. 1858) D14 (see below for synonymy)
| | `--E. submarginalis (Tryon 1892) G75, D00a (see below for synonymy)
| |--Carpocoris Kolenati 1846 L-SSY04
| | |--C. fuscispinus H01
| | |--C. lunulatus H01
| | `--C. purpureipennis [incl. C. nigricornis] H01
| `--Hypogomphus Spinola 1850 G76
| |--*H. rugosus Spinola 1850 G76
| |--H. bipunctatus (Guérin-Méneville 1838) [=Platycoris bipunctata] G76
| |--H. bufo WEE70
| `--H. rotundatus (Dallas 1851) [=Mazium rotundatum] G76
`--+--Strachiaria G76
`--+--Antestiaria G75
`--+--Pentatoma group G75
| |--Alciphron glaucus G76
| |--Chlorocoris subrugosus G76
| |--Acrosternum hilare G76, H91
| |--Pentatoma G76
| |--Nezara G76
| |--Glaucias G76
| | |--G. amyoti CGW91
| | |--G. impar G76
| | `--G. sulcata G76
| `--Plautia G76
| |--P. affinis G76
| |--P. brunnipennis (Montr. 1861) [=Pentatoma brunnipennis] D14
| |--P. discolor (Walker 1867) [=Pentatoma discolor] D00a
| |--P. fimbriata (Fabr. 1794) (see below for synonymy) D00a
| |--P. grossepunctata (Kirby in Andrews 1900) [=Pentatoma grossepunctata] D00a
| `--P. prolata (Walker 1867) [=Pentatoma prolata] D00a
`--Rhynchocoris group G75
|--Biprorulus bibax WEE70
|--Cuspicona G76
|--Parocirrhoe G76
|--Petalaspis Bergroth 1916 G76
| `--*P. tescorum Bergroth 1916 G76
|--Everardia Gross 1975 G76
| `--*E. picta Gross 1975 G76
|--Rhynchocoris C96
| |--R. humeralis C96
| `--R. plagiatus (Walker 1867) [=Cuspicona plagiata; incl. R. taprobanensis Bergr. 1891] D00b
|--Morna Stål 1867 G76
| |--M. florens (Walker 1867) [=Cuspicona florens; incl. *M. cornuta Haglund 1868] G76
| |--M. aggressor (Fabr. 1775) [=Cimex aggressor, Hoffmanseggiella aggressor] D00b
| `--M. leucospila (Walker 1867) [=Cuspicona leucospila] D00b
|--Diaphyta Bergroth 1891 (see below for synonymy) G76
| |--D. fulvescens (Dallas 1851) G76 (see below for synonymy)
| |--D. pulchra (Dallas 1851) [=Cuspicona pulchra] G76
| |--D. rosea Bergroth 1912 G76
| `--D. tasmani (Distant 1910) [=Vandiemenia tasmani] G76
|--Vitellus G76
| |--V. forficuloides (Walker 1867) [=Cuspicona forficuloides] D00b
| |--V. insularis Stal 1876 K08
| |--V. propinquus (Walker 1867) [=Cuspicona propinqua] D00b
| |--V. ‘proximus’ (Walker 1868 non Walker 1867) [=Cuspicona proxima] D00b
| |--V. strenuus (Walker 1868) [=Cuspicona strenua] D00b
| `--V. transversus (Walker 1867) [=Cuspicona transversa] D00b
`--Ocirrhoe Stål 1867 G76
|--O. australis (Westwood 1837) (see below for synonymy) G76
|--O. coronata Gross 1975 G76
|--O. dallasi Gross 1975 G76
|--O. inconspicua (Dallas 1851) [=Cuspicona inconspicua] D00b
|--O. lutescens Distant 1900 G76
|--O. prasinata (Stål 1859) [=Cuspicona prasinata] G76
|--O. unimaculata (Westwood 1837) G76 (see below for synonymy)
`--O. wilsoni Gross 1975 G76
Pentatominae incertae sedis:
Halyomorpha D14
|--H. canalana Distant 1914 D14
`--H. picus H01
Ambybelus elegans Montr. 1864 D14
Catacanthus G76
|--C. carrenoi (Le Guillon 1841) D14 [=Raphigaster carrenoi D14; incl. Cermatulus pulcher G75]
|--C. incarnatus G76
|--C. nigripes C91
|--C. punctus G76
`--C. viridicatus Distant 1881 K08
Bathycoelia Amyot & Serville 1843 [incl. Furtina Stal 1867, Gastraulax Herr. 1844] D14
`--B. longirostris Montr. 1861 [=Furtina longirostris, Gastraulax longirostris] D14
Cosmopepla G76
|--C. bimaculata E66
`--C. ingressa [=Eysarcoris ingressus] G76
Sepontia G76
Stagonomus G76
Amblycara gladiatora G76
Hyrmine Stål 1876 G76
|--*H. sexpunctata (Linnaeus 1758) G76 (see below for synonymy)
`--H. dispar (Westwood 1837) G76
Aeliini [Aeptini, Dymantaria] G75
|--Dymantis G75
`--Aelia G75
|--A. acuminata H01
|--A. americana G75
`--A. sibirica H01
Galgacus labidus (Erichson 1842) [incl. Dictyotus macer Walker 1867] G75
Myrocharia G75
Andrallus CGW91
Oechalia Stål 1862 CGW91, G75
`--*O. schellenbergii (Guérin-Méneville 1831) (see below for synonymy) G75
Platynopus melacanthus (Boisduval 1835) CGW91, D14 [=Pentatoma melacanthum D14]
Amyotea CGW91
Cermatulus Dallas 1851 CGW91, G75
`--*C. nasalis (Westwood 1837) [=Aelia nasalis] G75
Jalloides Schouteden 1907 CGW91, G75
`--*J. rubricosus (Stål 1870) [=Jalla rubricosa] G75
Sciocoris G76 [Sciocorinae, Sciocorini WEE70]
|--S. microphthalmus H01
|--S. obscurus Dallas 1851 G76
`--S. umbrinus H01
Crollius Distant 1901 D01
`--*C. conspersus (Walker 1867) [=Podops conspersus] D01
Microdeuterus aequalis Walker 1867 D01
Brachystethus D01
|--‘Eysarcoris’ alienus Walker 1867 (n. d.) D00b
|--B. discolor (Walk. 1867) [=Ochlerus discolor; incl. B. marginifer Walker 1868] D00b
|--B. fulvicornis (Walker 1867) (see below for synonymy) D00b
`--B. piceolus Walker 1867 D01
Ochrophara corinna (Kirby 1891) [=Pentatoma corinna] D00a
Brachymna Stål 1861 [incl. Balsa Walker 1867 non Walker 1860] D00a
`--B. tenuis Stål 1861 [incl. Balsa extenuata Walker 1867] D00a
Plexippus D00a
|--P. affinis Distant 1900 D00a
`--P. fulvescens (Dallas 1851) D00a [=Rhaphigaster fulvescens D00a, R. flavescens (l. c.) D00b]
Peribalus D00a
|--P. abbreviatus (Uhler 1872) [=Holcostethus abbreviatus] D00a
|--P. dubius (Dall. 1851) [=Pentatoma dubia] D00a
`--P. vernalis H01
Tolumnia D00b
|--T. aeola (Dall. 1851) [=Pentatoma aeola] D00a
|--T. basalis [incl. Pentatoma inobtrusa Walker 1867] D00b
`--T. latipes [incl. Pentatoma contingens Walker 1867, P. trispila Walker 1867] D00b
Brachynema biplaga (Walker 1867) [=Rhaphigaster biplaga] D00a
Thyanta D00a
|--T. custator E66
|--T. parvula (Dall. 1851) [=Rhaphigaster parvulus] D00a
|--T. pallidovirens WS04
| |--T. p. pallidovirens E66
| `--T. p. accerra E66
`--T. perditor [incl. Euschistus adjunctor Walker 1867, E. fasciatus Walker 1867] D00b
Afrania Stål 1864 [incl. Sala Walk. 1867] D00a
|--A. exigua (Walker 1868) [=Sala exigua] D00a
`--A. wahlbergi (Stål 1853) [=Strachia wahlbergi; incl. Sala colorata Walker 1867] D00a
Arocera D00b
|--A. apta (Walker 1967) [=Strachia apta] D00b
|--A. nigropicta (Walker 1867) [=Strachia nigropicta] D00b
`--A. placens (Walker 1867) [=Strachia placens] D00a
Vulsirea violacea (Fabr. 1803) D00a (see below for synonymy)
Banasa D00a
|--B. imbuta (Walker 1867) [=Rhaphigaster imbutus] D00b
|--B. inopinata (Walker 1867) [=Rhaphigaster inopinatus] D00a
`--B. varians Stål 1872 D00a (see below for synonymy)
Pallantia macula (Dall. 1851) [=Rhaphigaster macula; incl. Pentatoma diffusa Walker 1867] D00a
Sabaeus D00a
|--S. humeralis (Dall. 1851) [=Rhaphigaster humeralis; incl. Cuspicona smaragdina Walker 1867] D00a
`--S. spinosus (Dall. 1851) [=Rhaphigaster spinosus; incl. Mormidea ductor Walker 1867] D00a
Hyllus D00a
|--H. fasciatus Distant 1900 D00a
`--H. florens (Walker 1867) [=Mormidea florens] D00b
Zangis D00a
|--Z. impar (Walker 1867) [=Rhaphigaster impar; incl. R. ludekingii Voll. 1868] D00a
|--Z. subpunctata (Walker 1867) [=Pentatoma subpunctata; incl. Rhaphigaster melanostictus Voll. 1868] D00a
`--Z. varicornis (Dall. 1851) [=Rhaphigaster varicornis] D00a
Astyanax subactus (Walk. 1867) [=Hoplistodera subacta] D00a
Brachycoris insignis Distant 1900 D00b
Thoreyella concolor (Walker 1867) [=Mormidea concolor] D00b
Utana Distant 1900 D00b
`--*U. suprema (Walker 1867) [=Cuspicona suprema] D00b
Pugione D00b
Hoffmanseggiella D00b
|--H. defensor (Fabr. 1775) [=Cimex defensor; incl. Cuspicona vigens Walker 1867] D00b
`--H. hamata (Walker 1867) [=Cuspicona hamata; incl. C. contraria Walker 1867] D00b
Priassus exemptus (Walker 1868) [=Prionaca exempta; incl. Priassus carinatus Horv. 1889] D00b
Amyntor Stål 1867 [incl. Bolaca Walker 1867] D00b
`--A. unicolor (Walker 1867) [=Bolaca unicolor; incl. Oestopis terra Dist. 1879, Amyntor terra] D00b
Compastes obtusus (Walker 1868) (see below for synonymy) D00b
Prionochilus [incl. Lelia Walker 1867] D00b
`--P. decempunctatus (Motsch. 1859) [=Tropicoris decempunctatus; incl. Lelia porrigens Walker 1867] D00b
Leovitius Distant 1900 D00b
`--*L. macranthus (Dall. 1851) [=Rhaphigaster macranthus] D00b
Prionaca tortuosa Walker 1867 [incl. P. tauricornis Walker 1867] D00b
Araducta Walker 1867 D00b
|--A. bella Distant 1900 D00b
|--A. glabrata Walker 1867 D00b
|--A. malayana Distant 1900 D00b
|--A. repellens (Kirby 1892) [=Rhaphigaster repellens] D00b
`--A. sordida (Kirby 1892) [=Rhaphigaster sordida] D00b
Edessa D00b
Asyla Walker 1867 D00b
`--A. indicatrix Walker 1867 D00b
Boea Walker 1867 D00b
|--B. auriflura Walker 1867 D00b
|--B. postica Walker 1867 D00b
`--B. purpurascens Walker 1867 D00b
Taurocerus [incl. Canaca Walker 1867] D00b
|--T. abruptus (Walker 1867) [=Canaca abrupta] D00b
`--T. achilles [incl. T. divergens Walker 1867] D00b
Mormidea D00b
|--M. croceipes [incl. M. compta Walker 1867] D00b
|--M. integella [=Euschistus (Paraschistus) integellus] R74
|--M. lugens E66
|--M. nigriceps Walker 1868 (n. d.) D00b
|--M. pedestris Walker 1867 (n. d.) D00b
|--M. pulchella Walker 1868 D00b
|--M. punctifera (Walker 1867) [=Eysarcoris punctifer] D00b
`--M. tetra Walker 1868 D00b
Alcimus lineosus Walker 1867 D00b
Fecelia nigridens (Walker 1867) [=Loxa nigridens] D00b
Aspavia D00b
|--A. hastator D00b
`--A. infivita (Walker 1867) [=Mormidea infivita] D00b
Carbula D00b
|--C. contigua (Walker 1867) [=Mormidea contigua] D00b
|--C. insocia (Walker 1868) [=Eysarcoris insocius] D00b
|--C. lateralis (Walker 1867) [=Mormidea lateralis] D00b
`--C. socia (Walker 1867) [=Mormidea socia] D00b
Aeschrocoris testudinarius (Walker 1867) [=Aeschrus testudinarius] D00b
Cataulax D00b
|--C. punctipes Walker 1868 (n. d.) D00b
|--C. subvittatus Walker 1868 (n. d.) D00b
`--C. varicornis (Walker 1867) [=Pentatoma varicornis] D00b
Tropicorypha placida (Walker 1867) [=Pentatoma placida; incl. P. immunis Walker 1868] D00b
Holcostethus D00b
|--H. apicalis [incl. Pentatoma confinis Walker 1867] D00b
`--H. heterocerus (Walker 1867) [=Pentatoma heterocera] D00b
Runibia D00b
|--R. alligata (Walker 1867) [=Strachia alligata] D00b
|--R. euopta (Walker 1867) [=Strachia euopta] D00b
`--R. perspicua [incl. Strachia eucosma Walker 1867] D00b
Compsoprepes D00b
|--C. callideoides (Walker 1867) [=Strachia callideoides] D00b
`--C. calliphoroides (Walker 1867) [=Strachia calliphoroides] D00b
Jurtina chlorospila (Walker 1867) [=Bathycoelia chlorospila] D00b
Loxa D00b
|--L. affinis [incl. L. invaria Walker 1867] D00b
`--L. flavicollis [incl. L. deducta Walker 1867] D00b
Herda punctata [incl. Mormidea unisignata Walker 1867] D00b
Niphe subferruginea [incl. Pentatoma lateralis Walker 1867] D00b
Adria parvula [incl. Pentatoma brevittata Walker 1867] D00b
Paryphia pulchella [incl. Strachia notabilis Walker 1867] D00b
Dismegistus fimbriatus [incl. Strachia aenescens Walker 1867] D00b
Dinocoris tripterus [incl. Cataulax centralis Walker 1868] D00b
Pharypia nitidiventris [incl. Vulsirea transducta Walker 1867] D00b
Rhaphigaster D00b
|--R. nebulosa (Poda 1761) L-SSY04 [incl. R. grisea H01]
|--R. obliquus Walker 1867 D00b
|--R. patulus Walker 1867 (n. d.) D00b
|--R. quinquemaculatus Walker 1868 D00b
|--R. venustulus Walker 1867 D00b
`--R. viridipes Walker 1867 D00b
Arvelius mormideoides Walker 1868 (n. d.) D00b
Diceraeus piceus Walker 1868 D00b
Agonoscelis rutila (Fabricius 1775) G76 [=Cimex rutilus G76; incl. Strachia aspersa Walker 1867 D00b, tetragona Walker 1867 D00b]
Banasa varians Stål 1872 D00a [incl. Pentatoma basalis Walker 1867 D00b, Rhaphigaster derivatus Walker 1867 D00b, R. dimidiatus Stål 1858 D00a, Banasa dimidiata E66, P. subrufescens Walker 1867 D00a]
Brachystethus fulvicornis (Walker 1867) [=Rhaphigaster fulvicornis; incl. B. biguttatus Walker 1868, B. parvus Dist. 1889] D00b
Compastes obtusus (Walker 1868) [=Pentatoma obtusa; incl. Homalogonia maculata Jakowl. 1876, Compastes minor Atkins 1889] D00b
Diaphyta Bergroth 1891 [=Periboea Stål 1867 (preoc.) non Kunth 1843 (ICBN); incl. Vandiemenia Distant 1910] G76
Diaphyta fulvescens (Dallas 1851) G76 [=Cuspicona fulvescens G76, Periboea fulvescens D00b; incl. C. longiceps Walker 1867 D00b, C. pulchra Stål 1867 non Dallas 1851 G76, *Diaphyta pulchra G76, *Periboea pulchra G76]
Eurinome inconspicua (Montrouz. 1858) D14 [=Pentatoma inconspicua D14; incl. P. basiventris Sign. 1861 D14, P. circumdata Walker 1867 D00a]
Eurinome submarginalis (Tryon 1892) G75, D00a [=Pentatoma submarginalis D00a; incl. P. marginalis Walker 1867 non P. marginale Herr.-Schäff. 1836 D00a, Eurinome marginalis D00a]
*Hyrmine sexpunctata (Linnaeus 1758) G76 [=Cimex sexpunctatus G76; incl. Rhaphigaster nigrifascia Walker 1867 D00b]
Kapunda troughtoni (Distant 1910) [=Sciocoris troughtoni; incl. S. herrmansi Distant 1910, S. rubens Distant 1910, *Kapunda typica Distant 1911] G76
*Monteithiella humeralis (Walker 1868) G76 [=Strachia humeralis G76; incl. Antestia orbona G76, Pentatoma pallipes Dallas 1851 non Westwood 1837 G76, Antestia pallipes D00a]
Ocirrhoe australis (Westwood 1837) [=Rhynchocoris australis; incl. Cuspicona inconspicua Stål 1867 non Dallas 1851, *Ocirrhoe inconspicua] G76
Ocirrhoe unimaculata (Westwood 1837) G76 [=Rhynchocoris unimaculata G76; incl. Acanthosoma chlorophila Walker 1867 G76, Cuspicona fasciata Dallas 1851 D00b]
*Oechalia schellenbergii (Guérin-Méneville 1831) [=Pentatoma schellenbergii; incl. O. consocialis] G75
Omyta centrolineata (Westwood 1837) G76 [=Atelocerus centrolineatus G76, Poecilotoma centrolineata G75; incl. *O. deyrollei Spinola 1850 G76]
*Paramenestheus terricolor Breddin 1900 [incl. P. abditus Bergroth 1916, P. brevis (Van Duzee 1905), P. semoni (Horvarth 1900)] G75
Piezodorus hybneri (Gmelin 1790) G76 [=Cimex hybneri G76; incl. Rhaphigaster extenuatus Walker 1867 D00b, C. rubrofasciatus Fabricius 1787 non De Geer 1783 G76, Piezodorus rubrofasciatus G76]
Piezodorus lituratus (Fabricius 1794) G76 [=Cimex lituratus G76; incl. P. lituratus var. alliaceus H01, *P. degeeri Fieber 1861 G76]
Plautia fimbriata (Fabr. 1794) [=Cimex fimbriatus; incl. Rhaphigaster rufoviridis Walker 1867] D00a
*Theseus modestus (Stål 1865) [=Poecilometis modestus; incl. Spudeus lyricus Distant 1899, Theseus lyricus, T. nigrescens Distant 1910, Poecilometis plenus Walker 1869, P. scutellatus Distant 1899, Theseus turneri Distant 1910] G76
*Tholosanus proximus (Dallas 1851) [=Sciocoris proximus; incl. Pentatoma laticeps Walker 1867, Dictyotus laticeps, T. piceatus Distant 1910] G75
Vulsirea violacea (Fabr. 1803) D00a [=Cimex violaceus D00a; incl. V. lativentris Walker 1867 D00b, V. superba Walker 1867 D00a]
*Type species of generic name indicated
REFERENCES
[B27] Bergroth, E. 1927. Hemiptera Heteroptera from New Zealand. Transactions and Proceedings of the New Zealand Institute 57: 671–684.
[C96] Calder, A. A. 1996. Click beetles: genera of the Australian Elateridae (Coleoptera). CSIRO Australia: Collingwood.
[CGW91] Carver, M., G. F. Gross & T. E. Woodward. 1991. Hemiptera (bugs, leafhoppers, cicadas, aphids, scale insects etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1 pp. 429–509. Melbourne University Press: Carlton (Victoria).
[C91] CSIRO. 1991. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1. Melbourne University Press: Carlton (Victoria).
[D00a] Distant, W. L. 1900a. Rhynchotal notes.—IV. Heteroptera: Pentatominae (part). Annals and Magazine of Natural History, series 7, 5: 386–397.
[D00b] Distant, W. L. 1900b. Rhynchotal notes.—IV. Heteroptera: Pentatominae (part) (concluded). Annals and Magazine of Natural History, series 7, 5: 420–435.
[D01] Distant, W. L. 1901. Rhynchotal notes.—VIII. Heteroptera: fam. Coreidae. Annals and Magazine of Natural History, series 7, 7: 6–22.
[D14] Distant, W. L. 1914. Rhynchota from New Caledonia and the surrounding islands. In: Sarasin, F., & J. Roux (eds) Nova Caledonia: Forschungen in Neu-Caledonian und auf den Loyalty-Inseln. A. Zoologie vol. 1 pt 4 pp. 369–390, pls 11–12. C. W. Kreidels Verlag: Wiesbaden.
[E66] Evans, H. E. 1966. The Comparative Ethology and Evolution of the Sand Wasps. Harvard University Press: Cambridge (Massachusetts).
[G75] Gross, G. F. 1975. Plant-feeding and Other Bugs (Hemiptera) of South Australia. Heteroptera—Part I. Handbook of the Flora and Fauna of South Australia.
[G76] Gross, G. F. 1976. Plant-feeding and Other Bugs (Hemiptera) of South Australia. Heteroptera—Part II. Handbook of the Flora and Fauna of South Australia.
[H91] Hokkanen, H. M. T. 1991. Trap cropping in pest management. Annual Review of Entomology 36: 119–138.
[H01] Horváth, G. 1901. Hemipterák [Hemiptera]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 245–274. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.
[K08] Kirkaldy, G. W. 1908. A catalogue of the Hemiptera of Fiji. Proceedings of the Linnean Society of New South Wales 33: 345–391, pl. 4.
[L-SSY04] López-Sebastián, E., J. Selfa & J. Ylla. 2004. Primeros datos del parasitismo de Ooencyrtus pityocampae (Mercet, 1921) sobre Graellsia isabelae (Graells, 1849) en condiciones de laboratorio. Graellsia 60 (1): 121–123.
[R74] Rolston, L. H. 1974. Revision of the genus Euschistus in Middle America (Hemiptera, Pentatomidae, Pentatomini). Entomologica Americana 48 (1): 1–102.
[WS04] Wang, Q., & G. L. Shi. 2004. Mating frequency, duration, and circadian mating rhythm of New Zealand wheat bug Nysius huttoni White (Heteroptera: Lygaeidae). New Zealand Entomologist 27: 113–117.
[WEE70] Woodward, T. E., J. W. Evans & V. F. Eastop. 1970. Hemiptera (bugs, leafhoppers, etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 387–457. Melbourne University Press.
Last updated: 9 May 2022.
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