Pentatominae

Brown shield bug Dictyotus caenosus, copyright MAF.


Belongs within: Pentatomoidea.
Contains: Myrocharia, Diemeniini, Poecilometis, Tarisini, Phyllocephalinae, Cephaloplatus, Euschistus, Antestiaria, Pentatoma, Cuspicona.

The Pentatominae is a highly diverse group of stink bugs, the exact definition of which remains debated.

<==Pentatominae [Amyotinae, Graphosomatinae] B27
    |--Mycoolona atricornis G75, G76
    |--+--Mecidea [Mecideinae, Mecideini] G75
    |  `--Diemeniini G75
    |--+--Eysarcoris aeneus G75, G76
    |  |--Neostollia G76
    |  `--Spermatodes G76
    |--Poecilotoma Dallas 1851 [incl. Allportia Schoutenden 1906] G75
    |    |--*P. grandicornis (Erichson 1842) [=Atelocerus grandicornis] G75
    |    |--P. callosa Gross 1975 G75
    |    |--P. perconfusa Gross 1975 G75
    |    `--P. scutellata (Van Duzee 1905) [=Platycoris scutellatus] G75
    |--+--Adelaidena regina G76
    |  `--Kapunda Distant 1911 G75, G76
    |       |--K. troughtoni (Distant 1910) (see below for synonymy) G76
    |       `--K. tepperi Gross 1976 G76
    |--+--Bachesua Gross 1975 G75
    |  |    `--*B. enigmatica Gross 1975 G75
    |  `--Ippatha Distant 1910 G75
    |       |--*I. australiensis Distant 1910 G75
    |       `--I. angustilinea Musgrave 1930 G75
    |--+--Anaximenes G76
    |  `--Piezodorus Fieber 1861 G75, G76
    |       |--P. lituratus (Fabricius 1794) L-SSY04 (see below for synonymy)
    |       |--P. guildinii H91
    |       |--P. hybneri (Gmelin 1790) (see below for synonymy) G76
    |       `--P. pallescens K08
    |--Podopini [Podopinae] G75
    |    |--Coracanthella geophila G75
    |    |--Weda G75
    |    |--Amaurochrous G75
    |    `--Scotinophara G75
    |         |--S. allanae G75
    |         `--S. geophila (Montr. 1858) [=Scutellera geophila] D14
    |--+--Sciomenida Gross 1976 G76
    |  |    `--*S. mcfarlandi Gross 1976 G76
    |  |--Menida Motschulsky 1861 G75, G76
    |  |    |--*M. violacea Motschulsky 1861 G76
    |  |    `--M. personata Distant 1881 G76
    |  `--Lathraedoeus G76
    |       |--*L. speculator G76
    |       |--L. inconstans [=Menida inconstans] G76
    |       `--L. leucophaeus [=Antestia leucophaea, Menida leucophaea] G76
    |--Halyini [Halyinae] G76
    |    |--Poecilometis G76
    |    |--Atelias variegatus G76
    |    |--Coctoteris Stål 1858 G76
    |    |--Tinganina Bergroth 1909 G76
    |    |--Neosurenus Distant 1920 G76
    |    |--Elemana Distant 1912 G76
    |    |--Baracellus Distant 1903 G76
    |    |--Austromalaya Kirkaldy 1908 G76
    |    |--Bathrus Dallas 1851 G76
    |    |--Dalpada Amyot & Serville 1843 G76
    |    |--Acanthidiellum Kirkaldy 1904 [incl. Bromocoris Horvarth 1915] G76
    |    |--Ectenus Dallas 1851 G76
    |    |--Accarana Distant 1888 G76
    |    |--Auxentius Horvarth 1915 G76
    |    |--Neoptolemus Horvarth 1915 G76
    |    |--Babylas Horvarth 1915 G76
    |    |--Brizica Walker 1867 G76
    |    |--Dippilana Distant 1910 G76
    |    |    `--*D. membranacea Distant 1910 [incl. Poecilometis gibbiceps Bergroth 1916] G76
    |    |--Dictyometis Gross 1976 G76
    |    |    `--*D. spriggi Gross 1976 G76
    |    |--Parapoecilometis Gross 1976 G76
    |    |    `--*P. campbelli Gross 1976 G76
    |    |--Polycarmes Stål 1867 G76
    |    |    `--P. punctatissimus (Montr. 1858) [=Acanthidium punctatissimum] D14
    |    |--Anchises Stål 1867 G76
    |    |    `--A. parvulus (Westwood 1837) [=Halys parvulus, Theseus parvulus] G76
    |    |--Theseus Stål 1867 G76
    |    |    |--*T. modestus (Stål 1865) (see below for synonymy) G76
    |    |    `--T. distanti Gross 1972 [=T. scutellatus Distant 1910 non Poecilometis scutellatus Distant 1899] G76
    |    |--Aglaophon Stål 1876 G76
    |    |    |--*A. varius Stål 1876 G76
    |    |    `--A. variegatus Gross 1976 G76
    |    |--Paragenor Gross 1976 G76
    |    |    |--*P. halei Gross 1976 G76
    |    |    |--P. atomicus Gross 1976 G76
    |    |    `--P. robustus Gross 1976 G76
    |    |--Alcaeus Dallas 1851 [incl. Muritha] G76
    |    |    |--*A. varicornis (Westwood 1837) [=Atelocerus varicornis; incl. Al. subspinosus] G76
    |    |    |--A. hermannsburgi (Distant 1910) [=*Muritha hermannsburgi] G76
    |    |    |--A. lignicolor Walker 1867 G76
    |    |    `--A. unifornis Gross 1976 G76
    |    |--Neagenor Bergroth 1891 [=Agenor Dallas 1851 (preoc.)] G76
    |    |    |--*N. spinosus (Dallas 1851) [=*Agenor spinosus] G76
    |    |    |--N. anteaureus Gross 1976 G76
    |    |    |--N. minor Gross 1976 G76
    |    |    `--N. transcontinentalis Gross 1976 G76
    |    `--Omyta Spinola 1850 G76
    |         |--O. centrolineata (Westwood 1837) G76 (see below for synonymy)
    |         |    |--O. c. centrolineata G76
    |         |    `--O. c. siccior Gross 1976 G76
    |         |--O. delineata G76
    |         `--O. spinosa (Dallas 1851) G75 [=Poecilotoma spinosa G75; incl. Platycoris rufescens G76]
    |--+--+--Menestheus cuneatus G75
    |  |  |--Neomenestheus G75
    |  |  |--Hillieria Distant 1910 G75
    |  |  |    `--*H. acuminata Distant 1910 G75
    |  |  |--Paramenestheus Breddin 1900 G75
    |  |  |    |--*P. terricolor Breddin 1900 (see below for synonymy) G75
    |  |  |    `--P. nercivus (Dallas 1851) [=Sciocoris nercivus] G75
    |  |  |--Pseudaelia Distant 1910 G75
    |  |  |    |--*P. clementi Distant 1910 G75
    |  |  |    |--P. gnielae Gross 1975 G75
    |  |  |    `--P. rectimargo Gross 1975 G75
    |  |  `--Eribotes Stål 1867 G75
    |  |       |--*E. australis (Dallas 1852) [=Sciocoris australis] G75
    |  |       |--E. bicuspis G75
    |  |       |--E. challengeri G75
    |  |       |--E. delitescens G75
    |  |       |--E. hobartensis Distant 1910 G75
    |  |       |--E. indicator (Walker 1868) [=Sciocoris indicator] G75
    |  |       `--E. reconditus Bergroth 1920 G75
    |  `--+--Tarisini G75
    |     |--Kitsonia Gross 1975 G75
    |     |    `--*K. rubrocoriosa Gross 1975 G75
    |     |--+--Macrocarenus Stål 1873 G75
    |     |  |    `--*M. acuminatus (Dallas 1851) [=Eurygaster acuminatus] G75
    |     |  `--Macrocarenoides Gross 1975 G75
    |     |       `--*M. scutellatus (Distant 1899) [=Macrocarenus scutellatus] G75
    |     |--+--Ochisme Kirkaldy 1904 G75
    |     |  |    `--O. australis (Dallas 1851) [=Trachyops australis; incl. Hypolcus apricus Bergroth 1916] G75
    |     |  |--Dysnoetus Bergroth 1918 G75
    |     |  |    `--D. rugulosus (Horvarth 1902) [=Dictyotus rugulosus; incl. *Dy. anceps Bergroth 1918] G75
    |     |  |--Turrubulana Distant 1910 G75
    |     |  |    `--*T. plana Distant 1910 G75
    |     |  `--Amphidexius Bergroth 1918 G75
    |     |       `--*A. suspensus Bergroth 1918 G75
    |     `--+--Kumbutha quadrinotata G75
    |        |--Cooperocoris Gross 1975 G75
    |        |    `--*C. australis Gross 1975 G75
    |        |--Ooldeon Gross 1975 G75
    |        |    `--*O. maculipes Gross 1975 G75
    |        |--Brizocoris Gross 1975 G75
    |        |    `--*B. dimorphicus Gross 1975 G75
    |        |--Thryptomenecoris Gross 1975 G75
    |        |    `--*T. officeri Gross 1975 G75
    |        |--Eremophilacoris Gross 1975 G75
    |        |    `--*E. aridicolus Gross 1975 G75
    |        |--Pirricoris Gross 1975 G75
    |        |    `--*P. annectans Gross 1975 G75
    |        |--Tepperocoris Gross 1975 G75
    |        |    |--*T. lofteyensis Gross 1975 G75
    |        |    `--T. vandijki Gross 1975 G75
    |        `--Buthumka Gross 1975 G75
    |             |--*B. orthoscutellata Gross 1975 G75
    |             |--B. reducta Gross 1975 G75
    |             `--B. transitionalis Gross 1975 G75
    `--Pentatomini [Strachiaria, Strachiini] R74
         |  i. s.: Sibaria R74
         |         Agroecus R74
         |         Tibraca R74
         |         Menecles R74
         |         Hymenarcys R74
         |         Berecynthus R74
         |         Galedanta R74
         |         Dichelops R74
         |         Padaeus Stål 1862 R74
         |           |--*P. viduus (Vollenhoven 1868) [=Euschistus viduus] R74
         |           |--P. teapensis (Distant 1890) [=Euschistus teapensis] R74
         |           |--P. trivittatus R74
         |           `--P. verrucifer Stål 1862 [=Euschistus verrucifer] R74
         |         Eurydema [incl. Strachia] SS02
         |           |--E. dominulus (see below for synonymy) H01
         |           |--E. festivum [incl. E. festivum var. decoratum] H01
         |           |--E. fieberi [incl. E. fieberi var. armeniacum] H01
         |           |--E. oleraceum (Linnaeus 1758) L-SSY04 (see below for synonymy)
         |           |--E. ornatum H01
         |           `--E. sexpunctatum H01
         |         Biprorulus bibax WEE70
         |         Vitellus insularis Stal 1876 WEE70, K08
         |--+--Phyllocephalinae G75
         |  `--+--Cephaloplatus WEE70
         |     `--Caystrus G76 [incl. Neodius G75]
         |--+--+--Arniscus Distant 1899 G75
         |  |  |    `--A. humeralis G75
         |  |  `--Tholosanus Distant 1899 G75
         |  |       |--*T. proximus (Dallas 1851) (see below for synonymy) G75
         |  |       `--T. colligatus (Walker 1867) [=Pentatoma colligata] G75
         |  `--+--Utheria Gross 1975 G75
         |     |    `--*U. dixoni Gross 1975 G75
         |     |--Paradictyotus Gross 1975 G75
         |     |    `--*P. fuscus Gross 1975 G75
         |     `--Dictyotus Dallas 1851 WEE70, G75
         |          |--*D. tasmanicus Dallas 1851 G75
         |          |--D. caenosus (Westwood 1837) (see below for synonymy) G75
         |          |--D. conspicuus Gross 1975 G75
         |          |--D. inconspicuus Dallas 1851 [incl. Pentatoma aequalis Walker 1867] G75
         |          |--D. incultus (Erichson 1842) [=Cimex incultus] G75
         |          |--D. lincolnensis Gross 1975 G75
         |          |--D. pallipes (Westwood 1837) [=Pentatoma pallipes] G75
         |          |--D. roei (Westwood 1837) [=Pentatoma roei; incl. D. affinis Dallas 1851] G75
         |          `--D. scissicollis (Herrich-Schaeffer 1844) [=Pentatoma scissicolle] G75
         |--+--Codophila G76
         |  |--Proxys G75
         |  |--Euschistus G75
         |  |--Hyparete vitiensis Distant 1901 G76, K08
         |  |--Macromolus G76
         |  |--Lubentius G76
         |  |--Hypsethocus G76
         |  |--Dolycoris Mulsant & Rey 1866 G76, L-SSY04
         |  |    `--D. baccarum H01
         |  |--Monteithiella Gross 1976 G76
         |  |    `--*M. humeralis (Walker 1868) (see below for synonymy) G76
         |  |--Pseudapines Bergroth 1912 G76
         |  |    `--*P. geminata (Van Duzee 1905) [=Apines geminata] G76
         |  |--Notius Dallas 1851 G76
         |  |    |--*N. depressus Dallas 1851 G76
         |  |    `--N. consputus G76
         |  |--Agonoscelis Spinola 1837 G76
         |  |    |--*A. nubila (Fabricius 1775) [=Cimex nubilus] G76
         |  |    `--A. rutila (Fabricius 1775) [=Cimex rutilus] G76
         |  |--Anaxarchus G76
         |  |    |--A. pardalinus D14
         |  |    `--A. reyi (Montr. 1858) [=Pentatoma reyi] D14
         |  |--Eurinome G76
         |  |    |--E. inconspicua (Montr. 1858) [=Pentatoma inconspicua; incl. P. basiventris Sign. 1861] D14
         |  |    `--E. submarginalis G75
         |  |--Carpocoris Kolenati 1846 G76, L-SSY04
         |  |    |--C. fuscispinus H01
         |  |    |--C. lunulatus H01
         |  |    `--C. purpureipennis [incl. C. nigricornis] H01
         |  `--Hypogomphus Spinola 1850 G76
         |       |--*H. rugosus Spinola 1850 G76
         |       |--H. bipunctatus (Guérin-Méneville 1838) [=Platycoris bipunctata] G76
         |       |--H. bufo WEE70
         |       `--H. rotundatus (Dallas 1851) [=Mazium rotundatum] G76
         `--+--+--Murgantia histrionica G76
            |  `--Stenozygum Fieber 1861 G76
            |       |--S. coloratum [incl. *S. variegatum Fieber 1861] G76
            |       |--S. australe G76
            |       |--S. flavifrons Distant 1914 D14
            |       |--S. meridionale Gross 1976 G76
            |       |--S. miniatulum G76
            |       |--S. persignatum G76
            |       `--S. speciosum G76
            `--+--Antestiaria G75
               `--+--+--Alciphron glaucus G76
                  |  |--Chlorocoris subrugosus G76
                  |  |--Acrosternum hilare G76, H91
                  |  |--Pentatoma G75
                  |  |--Plautia G76
                  |  |    |--P. affinis G76
                  |  |    `--P. brunnipennis (Montr. 1861) [=Pentatoma brunnipennis] D14
                  |  |--Glaucias G76
                  |  |    |--G. amyoti G76
                  |  |    |--G. impar G76
                  |  |    `--G. sulcata G76
                  |  `--Nezara Amyot & Serville 1843 G76
                  |       |--N. viridula (Linnaeus 1758) [=Cimex viridulus] G76
                  |       |    |--N. v. viridula G76
                  |       |    `--*N. v. smaragdula (Fabricius 1775) [=Cimex smaragdulus] G76
                  |       `--N. heegeri G89
                  `--+--Cuspicona G76
                     |--Morna Stål 1867 G76
                     |    `--M. florens (Walker 1867) [=Cuspicona florens; incl. *M. cornuta Haglund 1868] G76
                     |--Petalaspis Bergroth 1916 G76
                     |    `--*P. tescorum Bergroth 1916 G76
                     |--Everardia Gross 1975 G76
                     |    `--*E. picta Gross 1975 G76
                     |--Diaphyta Bergroth 1891 (see below for synonymy) G76
                     |    |--D. fulvescens (Dallas 1851) (see below for synonymy) G76
                     |    |--D. pulchra (Dallas 1851) [=Cuspicona pulchra] G76
                     |    |--D. rosea Bergroth 1912 G76
                     |    `--D. tasmani (Distant 1910) [=Vandiemenia tasmani] G76
                     `--Ocirrhoe Stål 1867 G76
                          |--O. australis (Westwood 1837) (see below for synonymy) G76
                          |--O. coronata Gross 1975 G76
                          |--O. dallasi Gross 1975 G76
                          |--O. lutescens Distant 1900 G76
                          |--O. prasinata (Stål 1859) [=Cuspicona prasinata] G76
                          |--O. unimaculata (Westwood 1837) (see below for synonymy) G76
                          `--O. wilsoni Gross 1975 G76

Pentatominae incertae sedis:
  Halyomorpha D14
    |--H. canalana Distant 1914 D14
    `--H. picus H01
  Ambybelus elegans Montr. 1864 D14
  Catacanthus G76
    |--C. carrenoi (Le Guillon 1841) D14 [=Raphigaster carrenoi D14; incl. Cermatulus pulcher G75]
    |--C. incarnatus G76
    |--C. nigripes C91
    |--C. punctus G76
    `--C. viridicatus Distant 1881 K08
  Hypsithocus Bergroth 1927 B27
    `--*H. hudsonae Bergroth 1927 B27
  Bathycoelia Amyot & Serville 1843 [incl. Furtina Stal 1867, Gastraulax Herr. 1844] D14
    `--B. longirostris Montr. 1861 [=Furtina longirostris, Gastraulax longirostris] D14
  Cosmopepla ingressa [=Eysarcoris ingressus] G76
  Sepontia G76
  Stagonomus G76
  Amblycara gladiatora G76
  Hyrmine Stål 1876 G76
    |--*H. sexpunctata (Linnaeus 1758) [=Cimex sexpunctatus] G76
    `--H. dispar (Westwood 1837) G76
  Parocirrhoe G76
  Graphosoma G75
    |--G. italicum I92
    `--G. rubrolineatum G75
  Aeschrus G75
  Aeliini [Aeptini, Dymantaria] G75
    |--Dymantis G75
    `--Aelia G75
         |--A. acuminata H01
         |--A. americana G75
         `--A. sibirica H01
  Galgacus labidus (Erichson 1842) [incl. Dictyotus macer Walker 1867] G75
  Myrocharia G75
  Andrallus CGW91
  Oechalia Stål 1862 CGW91, G75
    `--*O. schellenbergii (Guérin-Méneville 1831) (see below for synonymy) G75
  Platynopus melacanthus (Boisduval 1835) CGW91, D14 [=Pentatoma melacanthum D14]
  Amyotea CGW91
  Cermatulus Dallas 1851 CGW91, G75
    `--*C. nasalis (Westwood 1837) [=Aelia nasalis] G75
  Jalloides Schouteden 1907 G75
    `--*J. rubricosus (Stål 1870) [=Jalla rubricosa] G75
  Sciocoris H01 [Sciocorinae, Sciocorini WEE70]
    |--S. microphthalmus H01
    |--S. obscurus Dallas 1851 G76
    `--S. umbrinus H01

Diaphyta Bergroth 1891 [=Periboea Stål 1867 (preoc.) non Kunth 1843 (ICBN); incl. Vandiemenia Distant 1910] G76

Diaphyta fulvescens (Dallas 1851) [=Cuspicona fulvescens; incl. C. pulchra Stål 1867 non Dallas 1851, *Diaphyta pulchra, *Periboea pulchra] G76

Dictyotus caenosus (Westwood 1837) [=Pentatoma caenosus; incl. D. bipunctatus Dallas 1851, P. latifrons Walker 1867, D. plebejus Stål 1859, D. polysticticus White in Dallas 1857, Pentatoma tibialis Walker 1867, P. vilis Walker 1867] G75

Eurydema dominulus [incl. Strachia festiva var. albiventris, E. dominulus var. dauricum, E. dominulus var. lhesgica] H01

Eurydema oleraceum (Linnaeus 1758) L-SSY04 [incl. E. oleraceum var. angulare H01, E. oleraceum var. annulatum H01, E. oleraceum var. consimile H01]

Kapunda troughtoni (Distant 1910) [=Sciocoris troughtoni; incl. S. herrmansi Distant 1910, S. rubens Distant 1910, *Kapunda typica Distant 1911] G76

*Monteithiella humeralis (Walker 1868) [=Strachia humeralis; incl. Antestia orbona, Pentatoma pallipes Dallas 1851 non Westwood 1837] G76

Ocirrhoe australis (Westwood 1837) [=Rhynchocoris australis; incl. Cuspicona inconspicua Stål 1867 non Dallas 1851, *Ocirrhoe inconspicua] G76

Ocirrhoe unimaculata (Westwood 1837) [=Rhynchocoris unimaculata; incl. Acanthosoma chlorophila Walker 1867] G76

*Oechalia schellenbergii (Guérin-Méneville 1831) [=Pentatoma schellenbergii; incl. O. consocialis] G75

Omyta centrolineata (Westwood 1837) G76 [=Atelocerus centrolineatus G76, Poecilotoma centrolineata G75; incl. *O. deyrollei Spinola 1850 G76]

*Paramenestheus terricolor Breddin 1900 [incl. P. abditus Bergroth 1916, P. brevis (Van Duzee 1905), P. semoni (Horvarth 1900)] G75

Piezodorus hybneri (Gmelin 1790) [=Cimex hybneri; incl. C. rubrofasciatus Fabricius 1787 non De Geer 1783, Piezodorus rubrofasciatus] G76

Piezodorus lituratus (Fabricius 1794) L-SSY04 [=Cimex lituratus G76; incl. P. lituratus var. alliaceus H01, *P. degeeri Fieber 1861 G76]

*Theseus modestus (Stål 1865) [=Poecilometis modestus; incl. Spudeus lyricus Distant 1899, Theseus lyricus, T. nigrescens Distant 1910, Poecilometis plenus Walker 1869, P. scutellatus Distant 1899, Theseus turneri Distant 1910] G76

*Tholosanus proximus (Dallas 1851) [=Sciocoris proximus; incl. Pentatoma laticeps Walker 1867, Dictyotus laticeps, T. piceatus Distant 1910] G75

*Type species of generic name indicated

REFERENCES

[B27] Bergroth, E. 1927. Hemiptera Heteroptera from New Zealand. Transactions and Proceedings of the New Zealand Institute 57: 671–684.

[CGW91] Carver, M., G. F. Gross & T. E. Woodward. 1991. Hemiptera (bugs, leafhoppers, cicadas, aphids, scale insects etc.) In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1 pp. 429–509. Melbourne University Press: Carlton (Victoria).

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