Epiclerus sp., photographed by Simon Van Noort.

Belongs within: Chalcidoidea.

The Tetracampinae is a group of small parasitic wasps that are mostly known to develop as parasites of leaf-miners or in eggs of leaf-feeding insects (Bouček 1988). Male tetracampines have five tarsomeres in the fore- and hindlegs, but only four in the midlegs (Heraty et al. 2013).

Synapomorphies (from Heraty et al. 2013): Male antenna with six flagellomeres between F1 and clava; head with upper ocular and transfacial sulci present anteriorly; clypeus inflected into oral cavity, not on same plane as face; pronotum with isolated large bristle on posterolateral corner; forewing with speculum absent (specular area setose both dorsally and ventrally).

<==Tetracampinae B88
    |  i. s.: Niticampe Bouček 1988 B88
    |           `--*N. metallica Bouček 1988 B88
    |         Cassidocida Crawford 1913 B88
    |           `--*C. aspidomorphae B88
    |--Diplesiostigma Girault 1920 HB13, B88 [incl. Xanthopirene Girault 1929 B88]
    |    |--*D. particolor Girault 1920 [incl. *Xanthopirene shakespearei Girault 1929] B88
    |    `--D. bisetosum Bouček 1988 B88
    `--+--+--Foersterella Dalla Torre 1897 HB13, B88 [=Hyperbius Förster 1878 non Stal 1867 B88]
       |  |    |--*F. flavipes [=Tetracampe flavipes] B88
       |  |    |--F. reptans MH11
       |  |    `--F. scaposa Bouček 1988 B88
       |  `--Tetracampe Förster 1841 HB13, B88
       |       |--*T. impressa B88
       |       |--T. africana [=Cassidocida africana] B88
       |       `--T. avia Bouček 1988 B88
       `--Epiclerus Haliday 1844 HB13, B88 [incl. Diparellomyia Girault 1913 B88]
            |--*E. panyas [=Entedon panyas] B88
            |--E. aligherini (Girault 1926) [=Tetracampe aligherini] B88
            |--E. australiensis (Girault & Dodd in Girault 1915) (see below for synonymy) B88
            |--E. haeckeli (Girault 1913) [=*Diparellomyia haeckeli] B88
            |--E. longicornis Bouček 1988 B88
            `--E. temenus MH11

Epiclerus australiensis (Girault & Dodd in Girault 1915) [=Tetracampe australiensis; incl. Toxeuma pax Girault 1915] B88

*Type species of generic name indicated


[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[HB13] Heraty, J. M., R. A. Burks, A. Cruaud, G. A. P. Gibson, J. Liljeblad, J. Munro, J.-Y. Rasplus, G. Delvare, P. Janšta, A. Gumovsky, J. Huber, J. B. Woolley, L. Krogmann, S. Heydon, A. Polaszek, S. Schmidt, D. C. Darling, M. W. Gates, J. Mottern, E. Murray, A. D. Molin, S. Triapitsyn, H. Baur, J. D. Pinto, S. van Noort, J. George & M. Yoder. 2013. A phylogenetic analysis of the megadiverse Chalcidoidea (Hymenoptera). Cladistics 29: 466-542.

[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PLoS One 6 (11): e27023.

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