Cerobasis guestfalica, photographed by Brian Valentine.

Belongs within: Psocoptera.

The Trogioidea is a group of small barklice with wings reduced or absent.

Characters (from Li 2002; translated using Mandarin Tools, all errors my own): Small size, wings (if present) not more than buds. Antennae with more than twenty segments, not annulated. Epicranial sutures complete or lateral arms absent; eyes large and not hairy, ocelli absent or three present, closely placed; lacinia tip not toothed or with only one large tooth, labial palp with four slender segments, terminal segment axe-shaped. Tarsus three-segmented, claw without preapical tooth.

<==Trogioidea [Psoquilloidea]
    |--Archaeatropidae R02
    |--Empheriidae R02
    |--Anomocopeus Badonnel 1967 [Anomocopeidae] L02
    |--Psoquillidae M93
    |    |--Balliella Badonnel 1949 M93
    |    |    `--*B. ealensis Badonnel 1949 M93
    |    |--Psoquilla Hagen 1865 [incl. Heteropsocus Verrill 1902] M93
    |    |    |--*P. marginepunctata Hagen 1865 M93 [incl. Heteropsocus dispar Verrill 1902 M93, P. dispar B49]
    |    |    `--P. infuscata Badonnel 1949 B49
    |    `--Rhyopsocus Hagen 1876 [incl. Deipnopsocus Enderlein 1903, Rhyopsocopsis Pearman 1929] M93
    |         |--*R. eclipticus Hagen 1876 [incl. R. phillipsae Sommerman 1956] M93
    |         |--R. afer (Badonnel 1948) [=Deipnopsocus afer] B49
    |         |--R. bentonae Sommerman 1956 M93
    |         |--R. bicornis Badonnel 1986 B86
    |         |--R. disparilis (Pearman 1931) M93 (see below for synonymy)
    |         |--R. maculosus M93
    |         |--R. micropterus Mockford 1971 M93
    |         |--R. orthatus B86
    |         |--R. pandanicola Thornton, Lee & Chui 1972 S95
    |         |--R. speciophilus MG56
    |         `--R. texanus (Banks 1930) (see below for synonymy) M93
    `--Trogiidae [Atropidae] R02
         |--Myrmicodipnella Enderlein 1909 M93
         |    `--*M. aptera Enderlein 1909 M93
         |--Trogium Iliger 1798 [incl. Atropos Leach 1815, Clothilla Westwood 1841] M93
         |    `--*T. pulsatorium (Linnaeus 1758) (see below for synonymy) M93
         |--Helminotrogia Li 2002 L02
         |    `--*H. bipunctatia Li 2002 L02
         |--Phlebotrogia Li 2002 L02
         |    `--*P. chinensis Li 2002 L02
         |--Lepinotus Heyden 1850 (see below for synonymy) M93
         |    |--*L. inquilinus Heyden 1850 (see below for synonymy) M93
         |    |--L. angolensis Badonnel 1955 B55
         |    |--L. patruelis Pearman 1931 M93
         |    |--L. quadrispinosus [=L. (Heterolepinotus) quadrispinosus] B55
         |    |--L. reticulatus Enderlein 1905 M93
         |    `--L. tasmaniensis Hickman 1934 M93
         `--Cerobasis Kolbe 1882 (see below for synonymy) M93
              |--+--C. caboverdensis M93
              |  |--C. harteni M93
              |  |--C. intermedia M93
              |  |--C. lambda M93
              |  |--C. papillata M93
              |  `--C. treptica M93
              `--+--*C. guestfalica (Kolbe 1880) (see below for synonymy) M93
                 |--C. alfredi M93
                 |--C. annulata (Hagen 1865) [=Clothilla annulata, Atropos annulata, Myopsocnema annulata] M93
                 |--C. bundyi M93
                 |--C. canariensis M93
                 |--C. lapidaria Badonnel 1955 M93, B55 [=C. lapidarius B55]
                 |--C. maculiceps M93
                 |--C. maderensis M93
                 |--C. multispinosa M93
                 `--C. recta M93

Cerobasis Kolbe 1882 [incl. Albardia Jacobson & Bianchi 1904, Hyperetes Kolbe 1880 (preoc.), Myopsocnema Enderlein 1905, Tichobia Kolbe 1882, Zlinia Obr 1948] M93

*Cerobasis guestfalica (Kolbe 1880) [=Hyperetes guestfalicus; incl. Tichobia alternans Kolbe 1882, Albardia alternans, Cerobasis muraria Kolve 1882, Hyperetes pinicola Kolbe 1881, H. tessulatus Hagen 1883] M93

Lepinotus Heyden 1850 [incl. Cuixa Navás 1927, Heterolepinotus Obr 1948, Paradoxenus Molshulsky 1852, Paradoxides Motschulsky 1851] M93

*Lepinotus inquilinus Heyden 1850 [=Clothilla inquilina, Atropos inquilina; incl. Cuixa canaria Navás 1927, A. distincta Kolbe 1888, Clothilla distincta, Cl. picea Hagen 1861, Atropos picea, Lepinotus piceus, Paradoxides psocoides Motschulsky 1851, Paradoxenus psocoides, Termes pulsatorium Scopoli 1763 non Linnaeus 1758, Atropos sericea Kolbe 1883, Lepinotus sericeus] M93

Rhyopsocus disparilis (Pearman 1931) M93 [=Deipnopsocus spheciophilus var. disparilis M93, R. spheciophilus disparilis MG56]

Rhyopsocus texanus (Banks 1930) [=Deipnopsocus texanus; incl. R. pescadori García Aldrete 1984, R. squamosus Mockford & Gurney 1956] M93

*Trogium pulsatorium (Linnaeus 1758) [=Termes pulsatorium, Atropos pulsatoria, Clothilla pulsatoria, Hemerobius pulsatorius Fabricius 1775, Psocus pulsatorius, Psylla pulsatoria Billburg 1820, Troctes pulsatorius; incl. Termes lignarium de Geer 1778, Atropos lignaria, Clothilla ocelloria Weber 1906, C. studiosa Westwood 1841] M93

*Type species of generic name indicated


[B49] Badonnel, A. 1949. Psocoptères de la Cote d'Ivoire. Mission Paulian-Delamare (1945). Revue Française d'Entomologie 16: 20-46.

[B55] Badonnel, A. 1955. Psocoptères de l'Angola. Diamang Publicações Culturais 26: 11-267.

[B86] Badonnel, A. 1986. Psocoptères de Colombie (Insecta, Psocoptera). Spixiana 9 (2): 179-223.

[L02] Li F. 2002. Psocoptera of China vol. 1. Science Press: Beijing.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[MG56] Mockford, E. L. & A. B. Gurney. 1956. A review of the psocids, or book-lice and bark-lice, of Texas (Psocoptera). Journal of the Washington Academy of Sciences 46 (11): 353-368.

[R02] Rasnitsyn, A. P. 2002. Superorder Psocidea Leach, 1815. In History of Insects (A. P. Rasnitsyn & D. L. J. Quicke, eds) pp. 125-133. Kluwer Academic Publishers: Dordrecht.

[S95] Smithers, C. N. 1995. Psocoptera (Insecta) of Christmas Island. Invertebrate Taxonomy 9: 529-561.

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