Lachesilla pedicularia, photographed by Alby Oakshott.

Belongs within: Lachesilloidea.

Lachesilla is a speciose genus of barklice with very sparse and short ciliation on the forewing and a distally open phallosome in the males. The type species L. pedicularia is mostly associated with fallen vegetation, including dead grasses, and has a global distribution as a result of being transported with harvested grain crops (Mockford 1993).

Characters (from Mockford 1993): Ocelli present. In forewing, Rs fork stem only slightly curved; veins Cu1a free from M. Phallosome with base an apodeme or two separate apodemes; aedeagal arch absent, external parameres absent as such. Hypandrium usually bearing pair of lateral processes ('claspers'). Ovipositor valvulae: v3 usually rounded at apex, occasionally pointed but point never spinose.

<==Lachesilla Westwood 1840 SN08 (see below for synonymy)
    |--L. corona group M93
    |    |--L. corona Chapman 1930 M93
    |    `--L. dividiforceps M93
    |--L. texcocana group M93
    |    |--L. delta M93
    |    `--L. texcocana García Aldrete 1972 M93
    |--L. patzunensis group M93
    |    |--L. sulcata García Aldrete 1986 M93
    |    `--L. tapanatepeca M93
    |--L. centralis group M93
    |    |--L. centralis García Aldrete 1983 M93
    |    |--L. cintalapa García Aldrete 1983 M93
    |    `--L. perezi García Aldrete 1983 M93
    |--L. riegeli group M93
    |    |--L. riegeli Sommerman 1946 M93
    |    |--L. tropica García Aldrete 1982 M93
    |    `--L. ultima García Aldrete 1982 M93
    |--L. andra group M93
    |    |--L. andra Sommerman 1946 M93
    |    |--L. arnae Sommerman 1946 M93
    |    |--L. dona Sommerman 1946 M93
    |    |--L. kerzhneri M93
    |    |--L. kola Sommerman 1946 M93
    |    |--L. nubilis (Aaron 1886) [=Caecilius nubilis] M93
    |    |--L. nubiloides García Aldrete 1975 M93
    |    `--L. punctata (Banks 1905) [=Elipsocus punctatus] M93
    |--L. forcepeta group M93
    |    |--L. alpejia García Aldrete 1988 M93
    |    |--L. anna Sommerman 1946 M93
    |    |--L. bottimeri Mockford & Gurney 1956 M93
    |    |--L. chapmani Sommerman 1946 M93
    |    |--L. contraforcepeta Chapman 1930 M93
    |    |--L. denticulata García Aldrete 1988 M93
    |    |--L. forcepeta Chapman 1930 M93
    |    |--L. gracilis García Aldrete 1988 M93
    |    |--L. kathrynae Mockford & Gurney 1956 M93
    |    |--L. major Chapman 1930 [=L. forcepeta var. major] M93
    |    |--L. penta Sommerman 1946 M93
    |    `--L. sandersoni Mockford 1974 M74
    |--+--L. fuscipalpis Badonnel 1971 M93
    |  `--L. rufa group M93
    |       |--L. abiesicola M93
    |       |--L. arida Chapman 1930 M93
    |       |--L. broadheadi M93
    |       |--L. chiricahua García Aldrete 1990 M93
    |       |--L. cupressicola M93
    |       |--L. jeanae Sommerman 1946 M93
    |       |--L. nita Sommerman 1946 M93
    |       |--L. pinicola M93
    |       |--L. rufa (Walsh 1863) [=Psocus rufus, Caecilius rufus, Pterodela rufa; incl. C. impacatus Aaron 1886] M93
    |       |--L. turneri M93
    |       |--L. yakima Mockford & García Aldrete 1974 M93
    |       `--L. yuccalnicola M93
    `--L. pedicularia group L98
         |--*L. pedicularia (Linnaeus 1758) SN08 (see below for synonymy)
         |--L. aldretei Badonnel 1986 B86
         |--L. bernardi Badonnel 1938 M93, G74
         |--L. dimorpha Lienhard 1981 L98
         |--L. greeni (Pearman 1933) M93 [=Terracaecilius greeni M93; incl. L. greeni f. longipennis Roesler 1939 G74]
         |--L. meinanderi Lienhard 1998 L98
         |--L. merzi Lienhard 1989 L98
         |--L. pacifica Chapman 1930 [incl. L. silvicola Chapman 1930, L. telsa Sommerman 1946] M93
         |--L. pallida (Chapman 1930) M93 [=*Terracaecilius pallidus NL07]
         |--L. quercus (Kolbe 1880) L98 (see below for synonymy)
         |--L. rena Sommerman 1946 M93
         |--L. rossica Roesler 1953 L98
         |--L. sauteri Lienhard 1977 L98
         |--L. tanaidana Roesler 1953 L98
         |--L. tectorum Badonnel 1931 [incl. L. pilosa Badonnel 1966] L98
         `--+--L. aethiopica (Enderlein 1902) [=Pterodela pedicularia var. aethiopica] M93
            `--L. nuptialis M93

Lachesilla incertae sedis:
  L. anura B55
  L. brinchangensis New & Lee 1991 NL07
  L. buettikeri New 1979 L98
  L. cameruna B55
  L. chiapensis Garcia Aldrete 1974 L02
  L. cornisterna B86
  L. cornuta Badonnel 1948 B55
  L. crutifurca Li 2002 [=L. crutifurcus] L03
  L. dichodolichna (Li 2002) [=Dicrolachesillus dichodolichnus] L03
  L. fusca Badonnel 1949 B55
  L. gigantea B55
  L. graminicola Badonnel 1986 B86
  L. gridellii (Navás 1927) (n. d.) [=Caecilius gridellii] L98
  L. hirsuta Lienhard 2002 NL07
  L. intrans Li 1993 L03, L02 [=Dicrolachesillus intrans L03]
  L. juniperana Garcia Aldrete 1991 L02
  L. latinerva Badonnel 1955 B55
  L. micrura Badonnel 1955 B55
  L. monocera Li 2002 L02
  L. mucronata Badonnel 1946 B55
  L. muncunilli (Navás 1913) L98
  L. novemimaculata Li 1993 L03, L02 [=Dicrolachesillus novemimaculatus L03]
  L. pachyura Badonnel 1955 B55
  L. qianshanensis (Li 2002) [=*Dicrolachesillus qianshanensis] L03
  L. rhizophila Li 2002 L02
  L. sabinae Li 1993 L03, L02 [=Dicrolachesillus sabinae L03]
  L. septenaria Li 2002 L02
  L. wutaishanensis (Li 2002) [=Dicrolachesillus wutaishanensis] L03
  L. ximaensis Li 2002 L02

Nomen nudum: Lachesilla albertina García Aldrete in Mockford 1993 M93

Lachesilla Westwood 1840 SN08 [=Pterodela Kolbe 1880 L02; incl. Dicrolachesillus Li 2002 L03, Leptopsocus Reuter 1899 L02, Terracaecilius Chapman 1930 L02]

*Lachesilla pedicularia (Linnaeus 1758) SN08 [=Hemerobius pedicularius SN08, Caecilius pedicularius M93, C. (*Pterodela) pedicularia M93, NL07, Psocus pedicularius L02; incl. Hemerobius abdominalis Fabricius 1775 L02, Ps. abdominalis M93, Ps. binotatus Rambur 1842 M93, Pterodela pedicularia var. brevipennis Enderlein 1903 M93, Lachesilla pedicularia var. brevipennis G74, Ps. domesticus Burmeister 1839 M93, Ps. dubius Stephens 1836 M93, *Leptopsocus exiguus Reuter 1899 NL07, M93, Termes fatidicum Linnaeus 1758 L02, Atropos fatidicus L02, Hemerobius fatidicus M93, Hyperetes fatidicus L02, Lachesilla fatidica NL07, Lachesis fatidica L02, Psocus fatidicus M93, Hemerobius flavicans Linnaeus 1758 L98, Elipsocus flavicans L02, Psocus flavicans L95, Ps. geologus Walsh 1862 M93, Pterodela pedicularia ab. giardi Enderlein 1901 M93, P. gobiernoi Navás 1913 L98, Lachesilla limbata Enderlein 1924 M93, Ps. nigricans Stephens 1836 M93, Caecilius nigrotuberculatus Curran 1925 M93, Hemerobius pusillus Müller 1776 M93, Caecilius pusillus M93, Psocus pusillus M93, Ps. salicis Hagen 1861 M93, Caecilius salicis L02, Lachesilla stigmalis Navás 1932 M93]

Lachesilla quercus (Kolbe 1880) L98 [=Caecilius (Pterodela) quercus L02, Dicrolachesillus quercus L03; incl. Pterodela livida Enderlein 1903 L98, Lachesilla lavidia G74, L. (Pterodela) machi Navas 1936 L02, L. wittmeri New 1979 L02]

*Type species of generic name indicated


[B55] Badonnel, A. 1955. Psocoptères de l'Angola. Diamang Publicações Culturais 26: 11–267.

[B86] Badonnel, A. 1986. Psocoptères de Colombie (Insecta, Psocoptera). Spixiana 9 (2): 179–223.

[G74] Günther, K. K. 1974. Staubläuse, Psocoptera. Die Tierwelt Deutschlands 61: 1–315.

[L95] Latreille, P. A. 1795. Extrait d'un mémoire pour servir de suite à l'histoire des termès, ou fourmis blanches. Bulletin de la Société Philomatique, a ses Correspondans 41–42: 1–3.

[L02] Li F. 2002. Psocoptera of China vol. 2. Science Press: Beijing.

[L98] Lienhard, C. 1998. Faune de France. France et Régions Limitrophes. 83. Psocoptères Euro-Méditerranéens. Fédération Française des Sociétés de Sciences Naturelles: Paris.

[L03] Lienhard, C. 2003. Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie 110 (4): 695–721.

[M74] Mockford, E. L. 1974. Records and descriptions of Cuban Psocoptera. Entomologica Americana 48 (2): 103–215.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.

[SN08] Schmidt, E. R., & T. R. New. 2008. The Psocoptera (Insecta) of Tasmania, Australia. Memoirs of Museum Victoria 65: 71–152.

Last updated: 10 August 2020.

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