Ptycta polluta, photographed by Graham Montgomery.

Belongs within: Psocinae.

Ptycta is a widespread genus of barklice with a particular radiation of species on the Hawaiian islands. The exact limits of the genus are uncertain, however, with a number of species assigned whose relationships require investigation.

Characters (from Mockford 1993): Adults of both sexes macropterous. Nine distal inner labral sensilla. Forewing markings extremely variable: usually a dark mark in pterostigma and dark marks on diagonal from pterostigma base to nodulus, but some species with extensive wing banding. Hypandrium with or without basal sclerites; with asymmetrical median strap based in middle or right, some species with processes at base of strap. Phallosome slightly tapering and rounded anteriorly; distally with pair of semi-membranous (presumed) external parameres and median aedeagal process of varying width, usually bifid at apex. Mape epiproct chair-shaped, extending anteriorly over clunium. Male paraproct with distal process long, slender, acuminate; no bulge on shoulder lateral to process; sense cushion near base of paraproct; basal articular region short and broad. Subgenital plate with distal piece short and wide, about as long as its basal width; pigmented arms directed anterolaterally, somewhat widened distally. Ovipositor valvulae: v2 with long, slender distal process; v3 ovoid; median lobe variable—prominent to absent.

<==Ptycta Enderlein 1925 SN08
    |--*P. haleakalae [=Psocus haleakalae] SN08
    |    |--P. h. haleakalae CT72
    |    `--P. h. hualalai CT72
    |--P. abouchari Badonnel 1986 B86
    |--P. acraea B86
    |--P. afasciata Endang, Thornton & New 2002 NL07
    |--P. apicantha CT72
    |    |--P. a. apicantha CT72
    |    |--P. a. mauiensis CT72
    |    `--P. a. multicantha CT72
    |--P. apicanthoides CT72
    |    |--P. a. apicanthoides CT72
    |    `--P. a. palligena CT72
    |--P. australis Schmidt & Thornton 1993 SN08
    |--P. breuschi Smithers & Thornton 1990 NL07
    |--P. buettikeri Badonnel 1982 L98
    |--P. caboverdensis Meinander 1966 L98
    |--P. campbelli Schmidt & Thornton 1993 SN08
    |--P. colei Schmidt & New 2008 SN08
    |--P. curvata Li 2002 L02
    |--P. dentata B86
    |--P. diadela CT72
    |--P. diastema CT72
    |--P. dicrosa CT72
    |--P. disclera CT72
    |    |--P. d. disclera CT72
    |    `--P. d. iaoensis CT72
    |--P. distincta Smithers & Thornton 1990 NL07
    |--P. distinguenda CT72
    |--P. drepana CT72
    |--P. drepanoides CT72
    |--P. elegantula Li 1999 L02
    |--P. emarginata New 1974 [incl. P. cornigera New 1974] SN08
    |--P. episcia CT72
    |--P. expansa B86
    |--P. fittkaui Badonnel 1986 B86
    |--P. flavipalpi Li & Yang 1987 L03, L02 [=P. flavipalpae (l. c.) L03]
    |--P. floresensis Endang, Thornton & New 2002 NL07
    |--P. freycineti Schmidt & New 2008 SN08
    |--P. frogneri CT72
    |--P. frontalis Thornton 1984 NL07
    |--P. furcata Li 1993 L02
    |--P. giffardi CT72
    |--P. glossoptera New 1974 SN08
    |--P. gynegonia CT72
    |--P. hardyi CT72
    |--P. heterogamias CT72
    |--P. hollowayae SN08
    |--P. incerta Smithers & Thornton 1990 NL07
    |--P. incurvata Thornton 1960 NL07
    |--P. kaala CT72
    |--P. kauaiensis CT72
    |--P. krakatau Vaughan, Thornton & New 1989 NL07
    |--P. lanaiensis CT72
    |    |--P. l. lanaiensis CT72
    |    |--P. l. fusca CT72
    |    |--P. l. halawa CT72
    |    `--P. l. persclera CT72
    |--P. leurothorax CT72
    |--P. lineata Mockford 1974 M93
    |--P. lobophora CT72
    |--P. lugosensis Smithers & Thornton 1990 NL07
    |--P. maculifrons CT72
    |--P. merapiensis Endang, Thornton & New 2002 NL07
    |--P. microctena CT72
    |--P. microglena CT72
    |--P. molokaiensis CT72
    |--P. monticola CT72
    |--P. nadleri Galil 1981 L98 [=P. apicantha nadleri CT72]
    |--P. novohibernica Smithers & Thornton 1990 NL07
    |--P. oahuensis CT72
    |--P. oligocantha CT72
    |--P. pallawahensis Schmidt & New 2008 SN08
    |--P. paranense B86
    |--P. pardena CT72
    |--P. pedina CT72
    |--P. perkinsi CT72
    |--P. persimilis CT72
    |--P. pikeloi CT72
    |--P. pitallo Endang, Thornton & New 2002 NL07
    |--P. placophora CT72
    |--P. polluta (Walsh 1862) (see below for synonymy) M93
    |--P. precincta Thornton 1984 NL07
    |--P. pupukea CT72
    |--P. revoluta Li 2002 L02
    |--P. rhina CT72
    |--P. sardjani Endang & Thornton 1992 NL07
    |--P. schillei (Enderlein 1906) NL07
    |--P. schisma CT72
    |--P. schneideri L98
    |--P. serrata Endang, Thornton & New 2002 NL07
    |--P. simulator CT72
    |    |--P. s. simulator CT72
    |    |--P. s. kilauea CT72
    |    |--P. s. konae CT72
    |    `--P. s. stena CT72
    |--P. sinuatistigma B86
    |--P. stenomedea CT72
    |--P. swezeyi CT72
    |--P. sylvestris CT72
    |--P. symmetrica CT72
    |--P. tapensis Endang, Thornton & New 2002 NL07
    |--P. telma CT72
    |--P. timorensis Endang, Thornton & New 2002 NL07
    |--P. tsutsumii New & Thornton 1976 NL07
    |--P. umbrata New 1974 SN08
    |--P. unica CT72
    |    |--P. u. unica CT72
    |    `--P. u. hawaiiensis CT72
    |--P. verticalis Vaughan, Thornton & New 1991 NL07
    |--P. vittipennis CT72
    `--P. zimmermani CT72

Ptycta polluta (Walsh 1862) [=Psocus pollutus, Psocidus pollutus; incl. Psocus texanus var. submarginatus Aaron 1886] M93

*Type species of generic name indicated


[B86] Badonnel, A. 1986. Psocoptères de Colombie (Insecta, Psocoptera). Spixiana 9 (2): 179–223.

[CT72] Chui, V. W. D., & I. W. B. Thornton. 1972. A numerical taxonomic study of the endemic Ptycta species of the Hawaiian Islands (Psocoptera: Psocidae). Systematic Zoology 21 (1): 7–22.

[L02] Li F. 2002. Psocoptera of China vol. 2. Science Press: Beijing.

[L98] Lienhard, C. 1998. Faune de France. France et Régions Limitrophes. 83. Psocoptères Euro-Méditerranéens. Fédération Française des Sociétés de Sciences Naturelles: Paris.

[L03] Lienhard, C. 2003. Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie 110 (4): 695–721.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.

[SN08] Schmidt, E. R., & T. R. New. 2008. The Psocoptera (Insecta) of Tasmania, Australia. Memoirs of Museum Victoria 65: 71–152.

Last updated: 15 August 2020.

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