Matsumuraiella radiopicta, photographed by Kazunori Yoshizawa.

Belongs within: Psocomorpha.
Contains: Stenopsocidae, Caeciliusidae, Amphipsocidae.

The Caeciliusetae are a clade of barklice characterised by the absence of a preapical tooth on the pretarsal claw, and reduction of the external valve of the gonapophyses.

Characters (from Yoshizawa 2002): Small to large in size, about 2–10 mm in length. Sexual dimorphism present. Head with rounded vertex; eyes variable; ocelli complete, clustered on small tubercle; mandible more or less elongate, outer margin angled; galea ball-shaped; labial palpus somewhat triangular. Prothorax less bulged dorsally; preepisternum of prothorax elongate. Pterothorax greatly bulged dorsally in winged form; mesothorax with broad precoxal bridge and narrow trochantin; metathorax with broad membranous region. Forewing variable in venation, size, shape, and colour; posteroproximal corner smoothly rounded. Hindwing with smoothly rounded posteroproximal corner; ciliation variable; Sc clear; Rs and M + Cu fused for long distance basally; Rs and M fused. Tarsi 2-segmented; claws without preapical tooth. Male genitalia: Hypandrium simple, posterior margin smoothly rounded. Phallosome: aedeagus pointed apically, strongly arched in lateral aspect; paramere extended along aedeagus, strongly arched in lateral aspect; phallobase simple and narrow, anterior margin not divided; endophallus usually with weakly sclerotized region but never with rod-like sclerites. Female genitalia: Subgenital plate simply rounded or slightly hollowed posteriorly. Gonapophyses variable; external valve more or less reduced.

<==Caeciliusetae [Caecilietae]
    |--Asiopsocidae [Asiopsocoidea] Y02
    |    |--Pronotiopsocus Mockford 1983 M93
    |    |    `--*P. amazonicus M93
    |    |--Notiopsocus Banks 1913 [incl. Lenkoella Machado-Allison & Papavero 1964] M93
    |    |    |--*N. simplex M93
    |    |    |--N. aldretei Y02
    |    |    |--N. machadoi Badonnel 1955 B55
    |    |    `--N. vilhenai Badonnel 1955 B55
    |    `--Asiopsocus Günther 1968 Y02
    |         |--*A. mongolicus Günther 1968 L02
    |         |--A. meridionalis M93
    |         |--A. sonorensis Mockford & García Aldrete 1976 M93
    |         `--A. wulingshanensis Li 2002 L02
    `--Caeciliusoidea [Caecilioidea] Y02
         |--Stenopsocidae Y02
         `--+--Caeciliusidae Y02
            `--+--Amphipsocidae Y02
               `--Dasydemellidae [Dasydemellinae] Y02
                    |--Ptenopsila M93
                    |--Teliapsocus Chapman 1930 M93
                    |    |--*T. conterminus (Walsh 1863) (see below for synonymy) M93
                    |    `--T. distinctus Badonnel 1986 B86
                    |--Matsumuraiella Enderlein 1906 L02
                    |    |--*M. radiopicta Enderlein 1906 L02
                    |    |--M. auriformis Li 1989 L02
                    |    |--M. enderleini Banks 1937 L02
                    |    |--M. guichengana Li 1997 L02
                    |    |--M. maculosa Li 1997 L02
                    |    |--M. perducta Li 2002 L02
                    |    |--M. quadripunctata Li 1993 L02
                    |    `--M. wangae Li 1997 L02
                    `--Dasydemella Enderlein 1909 SN04
                         |--*D. sylvestrii Enderlein 1909 SN04
                         |--D. aragua Mockford 1996 SN04
                         |--D. calocephala (Li 1992) [=Matsumuraiella calocephala] L02
                         |--D. cirrata Li 2002 L02
                         |--D. colossa Li 1999 L02
                         |--D. compressa (Li 1989) [=Matsumuraiella compressa] L02
                         |--D. crassistigma Mockford 1996 SN04
                         |--D. dezayasi Mockford 1974 SN04
                         |--D. diffusa (Li 1992) [=Matsumuraiella diffusa] L02
                         |--D. estriata Li 1995 L02
                         |--D. formosa Li 2002 L02
                         |--D. gynopeza Roesler 1940 SN04
                         |--D. modesta (Banks 1938) [=Elipsocus modestus] SN04
                         |--D. montana Badonnel 1986 SN04
                         |--D. obrienorum Garcia Aldrete 1978 SN04
                         |--D. planistigma Mockford 1996 SN04
                         |--D. septenpunctata (Li & Yang 1992) [=Matsumuraiella septenpunctata] L02
                         |--D. setosa Roesler 1940 SN04
                         |--D. sichuanana Li 1997 L02
                         |--D. stigmata Badonnel 1987 SN04
                         |--D. stipitiformis Li 1997 L02
                         |--D. xiaochuni Li 2002 L02
                         `--D. yalinana Li 2002 L02

*Teliapsocus conterminus (Walsh 1863) [=Psocus conterminus, Elipsocus conterminus, Matsumuraiella contermina; incl. Psocus canadensis Provancher 1876, Caecilius definitus Aaron 1883] M93

*Type species of generic name indicated


[B55] Badonnel, A. 1955. Psocoptères de l'Angola. Diamang Publicações Culturais 26: 11-267.

[B86] Badonnel, A. 1986. Psocoptères de Colombie (Insecta, Psocoptera). Spixiana 9 (2): 179-223.

[L02] Li F. 2002. Psocoptera of China vol. 1. Science Press: Beijing.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[SN04] Schmidt, E. R., & T. R. New. 2004. A systematic and phylogenetic revision of the family Elipsocidae (Insecta: Psocoptera), with the erection of two new families: Lesneiidae and Sabulopsocidae. Invertebrate Systematics 18: 157-213.

[Y02] Yoshizawa, K. 2002. Phylogeny and higher classification of suborder Psocomorpha (Insecta: Psocodea: ‘Psocoptera’). Zoological Journal of the Linnean Society 136: 371-400.

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