Xanthocaecilius quillayute, photographed by Dann Thombs.

Belongs within: Caeciliusetae.
Contains: Enderleinella, Paracaecilius, Fuelleborniella, Coryphacini, Austrocaeciliini, Caecilius, Dypsocus.

The Caeciliusidae are a group of barklice with forewing veins bearing setae on the dorsal surface only. They are found living on vegetation, most commonly on living trees (New & Lienhard 2007).

Characters (from Mockford 1993): Inner clypeal shelf variable. Distal inner labral sensilla five to seven. Lacinial tip variable: broad and flat, broad and denticulate, slender and bicuspid, or slender and bilobed. Labral stylets present or absent. Maxillary palpus with mx4 shorter than mx2. Setae of forewings slanting distad, relatively short, usually in one rank except on costa. Setae on veins in distal half of forewing only on dorsal surface. Hindwing margin mostly ciliated except basal two-thirds of front margin. Spine of free margin of paraproct small or absent. Endophallus with median bulb, usually not divided on midline, ovipositor valvulae with v3 a sclerotic bulge from base of v2.

<==Caeciliusidae [Caeciliini]
    |--Aphyopsocus Smithers 1982 [Aphyopsocinae] SN08
    |    `--*A. prolixus Smithers 1982 SN08
    |--Paracaeciliinae SN08
    |    |--Enderleinella NL07
    |    `--Paracaecilius SN08
    |--Amphicaecilius Li 2002 [Amphicaeciliinae] L02
    |    |--*A. pulcherrimus Li 2002 L02
    |    |--A. daqingshanicus Li 2002 L02
    |    |--A. floribundus Li 2002 L02
    |    `--A. pulchellus Li 2002 L02
    |--Fuelleborniellinae [Fuellebornelliini] NL07
    |    |--Fuelleborniella S95
    |    `--Ypsiloneura B55
    |         |--Y. kirkpatricki B55
    |         `--Y. monostyla Badonnel 1955 B55
    `--Caeciliusinae [Dypsocinae] SN08
         |  i. s.: Disialacaecilia Li 2002 L02 [=Disiacaecilia L03, Disialacaecilius L03]
         |           `--*D. ningxiaensis Li 2002 L02
         |         Parvialacaecilia Li 2002 L02
         |           `--*P. hebeiensis Li 2002 L02 [=P. hedeiensis L03]
         |         Anoculaticaeca Li 1997 L03, L02 [=Anoculaticaecilius (l. c.) L03]
         |           `--*A. chuanshaanica Li 1997 L02, L03, L02 [=Anoculaticaecilius chuanshaanicus (l. c.) L03]
         |         Licaecilius Lienhard 2003 L03 [=Neocaecilius Li 2002 non Li 2000 L02]
         |           |--*L. triradiatus (Li 2002) [=*Neocaecilius triradiatus] L03
         |           `--L. mangshiensis (Li 2002) [=Neocaecilius mangshiensis] L03
         |         Bivalvicaecilia Li 2002 L02 [=Bivalvicaecilius L03]
         |           |--*B. longiansa Li 2002 L02
         |           `--B. abbreviata (Li 1993) [=Caecilius abbreviatus, Asiocaecilius abbreviatus] L02
         |         Phymocaecilius Li 2002 L02 [=Phtmocaecilius L03, Phymocaecilus L03, Phymocaeilius L03]
         |           |--*P. subulosus Li 2002 L02
         |           |--P. fortis (Li 1995) [=Caecilius fortis, Velenzuala forti] L02
         |           |--P. fuscifascus Li 2002 L02
         |           |--P. guizhouensis Li 2002 L02
         |           `--P. puniceifascus Li 2002 L02
         |--Coryphacini SN08
         |--Austrocaeciliini NL07
         |--Caeciliusini SN08
         |    |--Caecilius NL07
         |    `--Dypsocus NL07
         |--Bassocaeciliini SN08
         |    |--Bassocaecilius Schmidt & New 2008 SN08
         |    |    `--*B. rawlinsoni Schmidt & New 2008 SN08
         |    `--Clinocaecilius Schmidt & New 2008 SN08
         |         `--*C. edwardsi Schmidt & New 2008 SN08
         |--Epicaeciliini SN08
         |    |--Nothocaecilius Schmidt & New 2008 SN08
         |    |    `--*N. thomasi Schmidt & New 2008 SN08
         |    `--Tasmanocaecilius Schmidt & New 2008 SN08
         |         `--*T. truchanasi Schmidt & New 2008 SN08
         `--Maoripsocini SN08
              |--*Maoripsocopsis sumatrensis Mockford 2000 NL07
              `--Maoripsocus Tillyard 1923 SN08
                   |--*M. semifuscatus Tillyard 1923 SN08
                   |--M. concavistigma (Schmidt & Thornton 1993) [=Caecilius concavistigma] SN08
                   |--M. hobartensis Schmidt & New 2008 SN08
                   |--M. juneae (Schmidt & Thornton 1993) [=Caecilius juneae] SN08
                   |--M. pedderi Schmidt & New 2008 SN08
                   |--M. spiralosus Schmidt & New 2008 SN08
                   |--M. tahunensis Schmidt & New 2008 SN08
                   |--M. wedgei Schmidt & New 2008 SN08
                   `--M. weindorferi Schmidt & New 2008 SN08

Caeciliusidae incertae sedis:
  Xanthocaecilius Mockford 1989 M93
    |--X. quillayute (Chapman 1930) [=Caecilius quillayute] M93
    `--X. sommermanae (Mockford 1955) [=Caecilius sommermanae] M93
  Orocaecilius Mockford 1999 L02
  Lacroixiella Badonnel 1943 L98
    `--*L. martini (Lacroix 1919) [=Caecilius martini] L98
  Eopsocites Hong 2002 H02
    `--*E. fushunensis Hong 2002 H02
  Stenopterites Hong 2002 H02
    |--*S. guchengziensis Hong 2002 H02
    `--S. longus Hong 2002 H02
  Fushunopsocus Hong 2002 H02
    `--*F. latus Hong 2002 H02
  Palaeopsocus Kolbe 1883 H02
  Ptenolasia Enderlei 1991 H02
  Liaoxipsocus Hong 1983 H02
    `--L. beipiaoensis (Hong 1983) L02
  Beipiaopsocus Hong 1983 H02
    `--B. divaricatus (Hong 1983) L02
  Huaxiapsocus Hong 1983 H02
    `--H. parvus (Hong 1983) L02
  Psocites Hong 1983 [incl. Parapsocus Hong 1983] H02
    `--P. pectinatus (Hong 1992) L02

*Type species of generic name indicated


[B55] Badonnel, A. 1955. Psocoptères de l'Angola. Diamang Publicações Culturais 26: 11–267.

[H02] Hong Y. 2002. Amber Insect of China. Beijing Scientific and Technological Publishing House.

[L02] Li F. 2002. Psocoptera of China vol. 1. Science Press: Beijing.

[L98] Lienhard, C. 1998. Faune de France. France et Régions Limitrophes. 83. Psocoptères Euro-Méditerranéens. Fédération Française des Sociétés de Sciences Naturelles: Paris.

[L03] Lienhard, C. 2003. Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie 110 (4): 695–721.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.

[SN08] Schmidt, E. R., & T. R. New. 2008. The Psocoptera (Insecta) of Tasmania, Australia. Memoirs of Museum Victoria 65: 71–152.

[S95] Smithers, C. N. 1995. Psocoptera (Insecta) of Christmas Island. Invertebrate Taxonomy 9: 529–561.

Last updated: 9 August 2020.

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